Rubiaceae
Tocoyena

Tocoyena formosa (Cham. & Schltdl.) K.Schum.

First published in C.F.P.von Martius & auct. suc. (eds.), Fl. Bras. 6(6): 347 (1889)
This species is accepted
The native range of this species is Suriname to Paraguay. It is a subshrub or shrub and grows primarily in the wet tropical biome.

Distribution

Native to:

Bolivia, Brazil North, Brazil Northeast, Brazil South, Brazil Southeast, Brazil West-Central, Paraguay, Suriname

Synonyms

Homotypic Synonyms

Heterotypic Synonyms

Classification

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Publications

POWO follows these authorities in accepting this name:

  • Delprete, P.G. & Cortés-B., R. (2006 publ. 2007). A synopsis of the Rubiaceae of the states of Mato Grosso and Mato Grosso do Sul, Brazil, with a key to genera, and a preliminary species list. Revista de Biologia Neotropical 3: 13-96.
  • Delprete, P.G. (2008). Revision of Tocoyena (Rubiaceae: Gardenieae) from the states of Goiás and Tocantins and a new species endemic to white-sand areas in the Brazilian cerrado. Journal of the Botanical Research Institute of Texas 2: 983-993.
  • Delprete, P.G. (2010). Rubiaceae. Flora dos estados de Goiás e Tocantins 40: 1-1610. Universidade Federal de Goiás.
  • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.
  • Jørgensen, P.M., Nee, M.H. & Beck., S.G. (eds.) (2013). Catálogo de las plantas vasculares de Bolivia. Monographs in Systematic Botany from the Missouri Botanical Garden 127: 1-1741. Missouri Botanical Garden.
  • Oliveira-Filho, A.T. (2006). Catálogo das Árvores nativas de Minas Gerais: 1-423. Editora UFLA, Lavas, Brasil.
  • Ramos Varjão, R., Gomes Jardim, J. & de Souza Canceição, A. (2013). Rubiaceae Juss. de caatinga na APA Serra Branco/Raso da Catarina, Bahia, Brasil. Biota Neotropica 13: 105-123.
  • Sant'Ana Melhem, T., das Graças Lapa Wanderley, M., Ehlin Martins, S., Jung-Mendaçolli, S.L., Shepherd, G.J. & Kirizawa, M. (eds.) (2007). Flora Fanerogâmica do Estado de São Paulo 5: 1-476. Instituto de Botânica, São Paulo.
  • Zappi, D.C. & al. (2003). Lista das plantas vasculares de Catoles. Boletim de Botânica da Universidade de São Paulo 21(2): 345-398.

Kew Backbone Distributions

  • Oliveira-Filho, A.T. (2006). Catálogo das Árvores nativas de Minas Gerais: 1-423. Editora UFLA, Lavas, Brasil.

Other Data

Other Kew resources that provide information on this taxon:

Date Reference Identified As Barcode Type Status Has image?
Jun 1, 2017 Silva, S.P.C. [565] Brazil K001192566 Yes
Jun 1, 2017 Nunes, T.S.; Carneiro, A.; Rocha, A.; Andrade, B.S.; Silva, B.M. [648] Brazil K001192562 Yes
Jun 1, 2017 Cavalcante, T.B. [35068] Brazil K001192569 Yes
Sep 15, 2005 Marilene; Krieger, P.L., et al. [100079] Brazil K000036485 Yes
Feb 15, 2005 Prado, J., et al. Brazil K000251343 Yes
Feb 1, 2005 Giulietti, A.M.; Harley, R.M. [1851] Brazil K000173308 Yes
Jan 1, 2001 André, M., et al. [4132] Brazil K000265121 Yes
Feb 1, 1999 Nic Lughadha, E.; França, F.; Gasson, P.; Harley, R. [PCD6018] Brazil K000013358 Yes
Feb 1, 1999 Stannard, B.; Guedes, M.L.; Saar, E.; Passos, L; Funck, L. [4621] Brazil K000173306 Yes
Sep 1, 1998 Atkins, S.; Sakuragui, C.M.; Harley, R.M.; Sano, P.T.; Souza, V.C. [CFCR14769] Brazil K000015240 Yes
Sep 1, 1998 Atkins, S.; Sakuragui, C.M.; Harley, R.M.; Sano, P.T.; Souza, V.C. [CFCR14769A] Brazil K000015241 Yes
Sep 1, 1998 Felix, L.P.; Dorneles, G.V. [1206] Brazil K000015236 Yes
Jan 1, 1998 Heringer, E.P. [5529] Brazil K001135940 Yes
Jan 1, 1998 Arbo, M.M.; Jardim, J.G.; Ferrucci, M.S.; de Sant'Ana, S.C. [7412] Brazil K000265201 Yes
Jan 1, 1997 Ganev, W. [1748] Brazil K000001904 Yes
Jan 1, 1997 Hind, D.J.N.; Nic Lughadha, E.; Queiroz, R.F. [H50580] Brazil K000001903 Yes
Oct 1, 1996 Laurenio, A.; Gomes, A.P.S.; Tscha, M. [345] Brazil K000015242 Yes
Oct 1, 1996 Andrade, K.; Freire, E.; Campelo, M.J.; Hora, M.J. [292] Brazil K000015243 Yes
Jun 1, 1996 Rodal, M.J.N.; Laurenio, A. [512] Brazil K000015239 Yes
Jun 1, 1996 Gomes, A.P.S.; Laurenio, A. [16] Brazil K000015238 Yes
Jan 1, 1994 Krapovickas, A.; Harley, R.M.; Cristobal, C.L.; Schinini, A. [45790] Paraguay K000994102 Yes
Oct 1, 1993 Agra, M.F. [1375] Brazil K000015248 Yes
Jan 1, 1993 Harley, R.M.; Hind, D.J.N.; Taylor, N. [27085] Brazil K000015244 Yes
Jan 1, 1993 Stannard, B.; Longhi-Wagner, H.M.; Lewis, G.P.; Giulietti, A.M.; Harley, R.M.; Arraes, M.G. [CFCR6846] Brazil K000015249 Yes
Jan 1, 1976 Irwin, H.S.; Harley, R.M.; Smith, G.L. [31313] Brazil K000015237 Yes
Burchell [7990/2] Brazil K001135919 Yes
Forzza, R.C. [2124] Brazil K001192510 Yes
Lemos, M.J.S. [115] Brazil K001192507 Yes
Ratter, J.A.; Ramos, J. [338] Brazil K001192520 Yes
Bridgewater, S. [S604] Brazil K001192500 Yes
Bernacci [1689] Brazil K001192515 Yes
Harley, R.M.; Steyermark, J.A. [16182] Brazil K000015247 Yes
Philcox, D. [3814] Brazil K001135948 Yes
Nunes, T.S. [463] Brazil K001192502 Yes
Salino, A. [5692] Brazil K001192512 Yes
Philcox, D.; Fereira, A.; Bertoldo, J. [3679] Brazil K001135947 Yes
Hassler, E. [4916] Paraguay K000994101 Type Yes
Weir, J. [III] Brazil K001135943 Yes
Irwin, H.S. [19376] Brazil K001192519 Yes
Melo, E. [2587] Brazil K001192516 Yes
Heringer, E.P. [2726] Brazil K001135933 Yes
Silva, M.M.; Santos, C.C.; Rocha, A.C.S.; Silva, B.M. [444] Brazil K001192509 Yes
Heringer, E.P.; Filgueiras, T.S.; Mendonça, R.C.; Pereira, B.A.S. [7371] Brazil K001135942 Yes
Pirani, J.R.; Harley, R.M.; Stannard, B.L.; Cordeiro, I.; Kameyama, C. [1953] Brazil K001135938 Yes
Ratter, J.A.; Bertolda, J.; Castro, R.A.; Santos, R.R.; Souza, R. [1097] Brazil K001192521 Yes
Eiten, G. [10544] Brazil K001192518 Yes
Assis, M.C. [149] Brazil K001192496 Yes
Harley, R.M.; Souza, R.; Ferreira, A. [10462] Brazil K001135941 Yes
Souza, V.C.; Souza, J.P.; Fanches, I.D.; Mazine, F.F.; Isaac, M.S.; Zandoval, J.A. [9367] Brazil K001135918 Yes
Carvalho-Sobrinho, J.G.; Miranda, E.B.; Castro, R.M.; Araújo, A. [460] Brazil K001192505 Yes
Silva, G.P.; Way, M.J. [SvW3626] Brazil K001192503 Yes
Guedes, M.L. [2950] Brazil K001192517 Yes
Guedes, M.L.; Stannard, B.; Woodgyer, E. [2636] Brazil K001192506 Yes
Argent, G.; Ramos, J.; Richards, P.W.; Souza, R. [6386] Brazil K001135930 Yes
Dubs, B. [1950] Brazil K001192527 Yes
Irwin, H.S.; Soderstrom, T.R. [7425] Brazil K001135921 Yes
Philcox, D.; Fereira, A.; Bertoldo, J. [3454] Brazil K001135932 Yes
Tameirão, E. [3211] Brazil K001192511 Yes
Harley, R.M.; Souza, R. [11027] Brazil K001135937 Yes
Heringer, E.P. [17947] Brazil K001135946 Yes
Souza, V.C.; Ferraro, L. [2572] Brazil K001192513 Yes
Farinaccio, M.A.; Magenta, M.A.; Magenta, E. [479] Brazil K001192495 Yes
Conceição, S.F.; Santos, A.K.A.; Nunes, T.S. [401] Brazil K001192508 Yes
Capellari, L. [5208] Brazil K001192522 Yes
Cobra, L.Q. [266] Brazil K001135929 Yes
Glaziou [14916] Brazil K001135925 Yes
Weir, J. [111] Brazil K001135924 Yes
Furlan, A.; Assos, M.A.; Garcia, F.C.P.; Ruffino, P.H.P.; Carneiro, C.E. [1573] Brazil K001135935 Yes
Rodal, M.J.N.; Silva, T.M.C. [776] Brazil K001192501 Yes
Dubs, B. [2015] Brazil K001192524 Yes
Ramos, J.; Sousa, R. [8] Brazil K001135931 Yes
Glaziou [17642] Brazil K001135926 Yes
Eiten, G. [3376] Brazil K001135927 Yes
Burchell [5852] Brazil K001135939 Yes
Robert, A. [655] Brazil K001135923 Yes
Ratter, J.A.; Filho, J.F.; Santos, J.B.; Oliveira, N.R.; Pais, N.O. [7063] Brazil K001192525 Yes
Ratter, J.A. [7006] Brazil K001192526 Yes
Krapovickas, A. [42801] Brazil K001192523 Yes
Pirani, J.R.; Harley, R.M.; Stannard, B.L.; Furlan, A.; Kameyama, C. [1821] Brazil K001192499 Yes
Martinelli, G. [3224] Brazil K001135936 Yes
Ratter, J.A.; Fonseca, S.G.; de Castro, R.A. [2669] Brazil K001135944 Yes
Ratter, J.A. [2425] Brazil K001135922 Yes
Irwin, H.S.; Soderstrom, T.R. [6365] Brazil K001135920 Yes
Glaziou [21484] Brazil K001135917 Yes
Prance, G.T.; Silva, N.T. [59318] Brazil K001135945 Yes
Philcox, D. [3566] Brazil K001135928 Yes
Tannus, J.L.S. [800] Brazil K001192514 Yes
Pirani, J.R. [1821] Brazil K001192498 Yes
Walter, B.M.T. [2927] Brazil K001192504 Yes
Gardner [(1297) 2197] Brazil Tocoyena hirsuta K001135934 Yes
Jan 1, 1998 Blanchet [3601] Brazil Tocoyena formosa var. microdon K000424435 Unknown Type Material Yes
Ratter, J. [3736] Brazil Tocoyena formosa var. microdon K001192528 Yes
Schultz, J.; Tawjoeran, J. [LBB 12441 A] Suriname Tocoyena surinamensis K000994099 Yes
Jan 1, 1998 Rombouts, H.E. [323] Suriname Tocoyena formosa var. pseudobrasiliensis K000424434 Unknown Type Material Yes
Jan 1, 1998 Gottsberger, I.; Gottsberger, G. [11-11171] Brazil Tocoyena formosa var. pseudobrasiliensis K000172525 Type Yes
Jan 1, 1998 Sellow, F. [s.n.] Brazil Tocoyena formosa var. pseudobrasiliensis K000424433 Unknown Type Material Yes
Gardner [1963] Brazil Tocoyena formosa var. pseudobrasiliensis K001192529 Yes
Teixeira, L.O.A. [261] Brazil Tocoyena formosa var. pseudobrasiliensis K001192537 Yes
Claussen [s.n.] Brazil Tocoyena formosa var. pseudobrasiliensis K001192540 Yes
Eiten, G.; Eiten, L.T. [10485] Brazil Tocoyena formosa var. pseudobrasiliensis K001192536 Yes
Giulietti, A.M.; Harley, R.M.; Stannard, B.; Mamede, M.C.H. [6404] Brazil Tocoyena formosa var. pseudobrasiliensis K001192535 Yes
Riedel [1028] Brazil Tocoyena formosa var. pseudobrasiliensis K001192530 Yes
Philcox, D. [3039] Brazil Tocoyena formosa var. pseudobrasiliensis K001192539 Yes
Gottsberger, I. [2222-1] Brazil Tocoyena formosa var. pseudobrasiliensis K001192534 Yes
Glaziou [13965] Brazil Tocoyena formosa var. pseudobrasiliensis K001192538 Yes
Riedel [s.n.] Brazil Tocoyena formosa var. pseudobrasiliensis K001192531 Yes
Lima, A. [53-1574-a] Brazil Tocoyena formosa var. pseudobrasiliensis K001192532 Yes
Gottsberger, I. [11-41171] Brazil Tocoyena formosa var. pseudobrasiliensis K001192533 Yes
Hassler, E. [7752] Paraguay Tocoyena bullata f. puberula K000994106 Type Yes
[21485] Brazil Tocoyena stipulosa K000424426 Unknown Type Material Yes

Sources

  • Angiosperm Extinction Risk Predictions v1

    • Angiosperm Threat Predictions
    • http://creativecommons.org/licenses/by/4.0

  • Herbarium Catalogue Specimens

    • Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

  • Kew Backbone Distributions

    • The International Plant Names Index and World Checklist of Vascular Plants 2024. Published on the Internet at http://www.ipni.org and https://powo.science.kew.org/
    • © Copyright 2023 World Checklist of Vascular Plants. http://creativecommons.org/licenses/by/3.0

  • Kew Names and Taxonomic Backbone

    • The International Plant Names Index and World Checklist of Vascular Plants 2024. Published on the Internet at http://www.ipni.org and https://powo.science.kew.org/
    • © Copyright 2023 International Plant Names Index and World Checklist of Vascular Plants. http://creativecommons.org/licenses/by/3.0

  • Kew Science Photographs

    • Copyright applied to individual images