Hawaiian Native and Naturalized
Vascular Plants Checklist
(December 2012 update)
Editor
Clyde T. Imada
Hawaiian Native and Naturalized Vascular Plants Checklist
(December 2012 update)
Editor:
Clyde Imada
Hawaii Biological Survey
Bishop Museum
Honolulu, HI 96817
Bishop Musem Technical Report 60
Honolulu, Hawai‘i
December 2012
Published by:
BISHOP MUSEUM
The State Museum of Natural and Cultural History
1525 Bernice Street
Honolulu, Hawai’i 96817–2704, USA
Copyright © 2012 Bishop Museum
All Rights Reserved
Printed in the United States of America
ISSN 1085-455x
Cover illustration: Brighamia rockii, scanned from original illustration by Yevonne Wilson-Ramsey used in
Manual of the Flowering Plants of Hawai‘i (Wagner et al. 1990, Plate 48)
Contribution No. 2012-021 to the Hawaii Biological Survey
Hawaiian Native & Naturalized Vascular Plant Checklist
TABLE OF CONTENTS
INTRODUCTION .......................................................................................................................................................................................... 1
BACKGROUND ............................................................................................................................................................................................ 1
FORMAT ........................................................................................................................................................................................................ 2
Citations ................................................................................................................................................................................................... 3
Explanation of categories included in the Citation field........................................................................................................................... 3
Navigating the Citation field .................................................................................................................................................................... 4
Additional abbreviations .......................................................................................................................................................................... 6
TABLE 1. Numbers of families, genera, and taxa by major group ................................................................................................................ 6
TABLE 2. Origin status of included taxa by major group ............................................................................................................................. 6
TABLE 3. Numbers of native vs. non-native taxa by major group ................................................................................................................ 6
ACKNOWLEDGMENTS .............................................................................................................................................................................. 7
BIBLIOGRAPHIC CITATIONS .................................................................................................................................................................... 8
INDEX OF ACCEPTED FAMILIES AND GENERA ................................................................................................................................. 18
APPENDIX A. Native and naturalized vascular plants of the main Hawaiian Islands ..................................................................... A-1–249
APPENDIX B. Native and naturalized vascular plants of the Northwestern Hawaiian Islands........................................................... B-1–17
APPENDIX C. Synonymy list for Hawaiian native and naturalized vascular plant checklist, arranged alphabetically
by major group and family.............................................................................................................................................................. C-1–31
APPENDIX D. Synonymy list for Hawaiian native and naturalized vascular plant checklist, arranged alphabetically
by old scientific name ..................................................................................................................................................................... D-1–26
APPENDIX E. Newly described taxa since 1990 in the Hawaiian Islands.............................................................................................. E-1–3
APPENDIX F. Taxa resurrected since 1990 in the Hawaiian Islands...................................................................................................... F-1–3
APPENDIX G. New naturalized taxa since 1990 in the Hawaiian Islands........................................................................................... G-1–17
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Hawaiian Native & Naturalized Vascular Plant Checklist
Hawaiian Native and Naturalized Vascular Plants Checklist (December 2012 update)
subsequent to publication of the Manual in 1990. This update is through
December 2012.
INTRODUCTION
A product of the Bishop Museum’s Hawaii Biological Survey, the
Hawaiian Native and Naturalized Vascular Plants Checklist is an authority
file of names of accepted native and naturalized angiosperm, gymnosperm,
and pteridophyte/lycophyte taxa in the Hawaiian Islands. The file is
maintained as a Microsoft Access database in the Department of Natural
Sciences/Botany, Bernice P. Bishop Museum. The foundation for the dicot
and monocot sections of the checklist is the Manual of the Flowering Plants
of Hawai‘i (Wagner et al. 1990, 1999, hereafter referred to as the Manual),
while that of the the fern/lycophyte section is Hawai‘i’s Ferns & Fern
Allies (Palmer 2003). Naturalized records of gymnosperms have been
included since being first reported in 2002 (Oppenheimer 2002).
BACKGROUND
The initial version of this checklist was published as “Checklist of
Native and Naturalized Flowering Plants of Hawai‘i” (Imada et al. 1989).
That checklist presented, in telegraphic format, a concise accounting of all
1,963 taxa of native and naturalized flowering plants to be treated the
following year in the Manual of the Flowering Plants of Hawai‘i (Wagner
et al. 1990). Arranged alphabetically by family, each taxon was listed
alphabetically by scientific name with authorship, distributional status,
endangerment status (if any), and island distribution. In some cases, more
specific locality data was provided, primarily for taxa with localized
distribution or known from only a few collections. The intent of the
checklist was to provide the user a quick-access reference for use in the
field. It was hoped that the checklist would stimulate further studies of
species distributions in the state, especially for undercollected weedy nonnative species.
This checklist is periodically updated to incorporate newly published
native and naturalized island and state records, nomenclatural and
taxonomic changes, and newly described native taxa. Many of the additions
are garnered from the annual Bishop Museum publication series, Records of
the Hawaii Biological Survey (1994 and continuing). Additions of new
island or state records are based on published sources supported by voucher
specimens; not included are citations based on anecdotal reports or casual
observations, or on unpublished checklists or manuscripts. This checklist
tracks and attempts to maintain conformity with the taxonomic concepts of
the Smithsonian Institution’s National Museum of Natural History Flora of
the Hawaiian Islands website
(http://botany.si.edu/pacificislandbiodiversity/hawaiianflora/index.htm),
which periodically publishes online supplements that update names and
distributions since publication of the Manual in 1990. At this writing, the
latest available supplement is dated 12 April 2012 (version 1.3,
http://botany.si.edu/pacificislandbiodiversity/hawaiianflora/Hawaiian_vascu
lar_plant_updates_1.3.pdf).
In 1992, the Hawai‘i State Legislature designated the Hawaii Biological
Survey (HBS) as a program of the Bishop Museum. Building upon a
century of scientific studies documenting and developing reference
collections of the plants and animals of Hawai‘i, Bishop Museum
developed a reputation as the go-to source of information on Hawaiian
organisms. Among its responsibilities, HBS maintains and updates
inventories of the flora and fauna of the Hawaiian Islands. The Records of
the Hawaii Biological Survey series has been published annually since 1994
to disseminate information on new island and state records of plants and
animals. As part of that mandate, the Botany unit has maintained an
inventory of the native and naturalized flowering plants of Hawai‘i,
originally as a searchable database. However, more recent versions have
been published on the Bishop Museum website as portable document
format (pdf) files. The previous pdf online version is dated October 2008.
The current version can be found online at
http://www.bishopmuseum.org/research/natsci/botany/botdbhome.html. A
searchable database format is also planned for the current version.
Pending nomenclatural or taxonomic changes or in-press taxa are
generally excluded until they are published, although exceptions are made.
Proposed additions or changes to the checklist are reviewed and evaluated
by staff botanists, often with outside consultation, before acceptance. While
the formal checklists in Appendix A and B list only currently accepted
names, Appendices C and D provide cross-referencing for all name changes
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Hawaiian Native & Naturalized Vascular Plant Checklist
The Manual proved to be a tremendous reference source for stabilizing
taxonomic concepts for Hawaiian plants and for catalyzing additional work
in Hawaiian taxonomy, as amply attested to by the impressive list of postManual bibliographic citations included in this checklist. This checklist
includes 2,983 taxa, over 1,000 more than were included in the original
checklist in 1989. Partly this is due to the added inclusion of 270 native and
naturalized gymnosperms and ferns/lycophytes, but also includes 60+
newly recognized native taxa (see Appendix E), 40+ resurrected native taxa
(Appendix F), and 570+ newly documented naturalized taxa (Appendix G).
The latter represents an annual average of 26 new naturalizations since the
Manual was published in 1990. Grand totals for families, genera, and taxa
included in this checklist can be found in Tables 1, 2, and 3 (page 5).
includes a listing of all the relevant family and generic changes affecting
Hawaiian plants. Abbreviations of author names follows Brummitt and
Powell (1992). If the author name is not included there, the author citation
provided by The Plant List (http://www.theplantlist.org) is followed.
Checklists such as this one become outdated as soon as they are
published, and error corrections, queries, or new information are much
appreciated; they can be relayed to the author at
cimada@bishopmuseum.org.
Distributional origin status is provided following the scientific name.
The categories are as follows:
end
end?
FORMAT
For ease of formatting, this checklist is split up into two geographic
units. Appendix A covers the eight main Hawaiian Islands, listed in order
geographically from northwest to southeast (Ni‘ihau, Kaua‘i, O‘ahu,
Moloka‘i, Läna‘i, Maui, Kaho‘olawe, Hawai‘i); also included as entities
separate from nearby Ni‘ihau are Ka‘ula Rock and Lehua Island. Appendix
B covers the Northwestern Hawaiian Islands (Papahänaumokuäkea), also
geographically from northwest to southeast (Kure, Midway, Pearl &
Hermes, Lisianski, Laysan, Gardner Pinnacles, French Frigate Shoals,
Necker, Nihoa). In each checklist, dicots are listed first, followed by
gymnosperms, monocots, and pteridophytes. A nested alphabetic
arrangement is followed within each group—family, genus, species, and
infraspecies.
ind
ind?
nat
nat?
pol
pol?
?
Family classifications for the dicots and monocots continue to follow
the Cronquist system as adopted in the Manual (Wagner et al. 1990). While
for now it will make navigation between the Manual and this checklist
easier for the user, it is inevitable that the overwhelming molecular
evidence for revised family concepts compiled by the Angiosperm
Phylogeny Group (http://www.mobot.org/MOBOT/ research/APweb/) will
eventually be adopted by Bishop Museum’s Herbarium Pacificum and will
be reflected in this checklist.
endemic, occurring only in the Hawaiian Islands
possibly endemic, or possibly conspecific with a species occurring
elsewhere
indigenous, occurring naturally in the Hawaiian Islands but also
elsewhere in the world
probably indigenous, but possibly introduced in the Hawaiian
Islands by man
naturalized, introduced by man and now established and
maintaining or spreading their populations by vegetative or sexual
means
probably naturalized, but possibly indigenous or a Polynesian
introduction
Polynesian introduction (intentional or unintentional)
possibly a Polynesian introduction, or perhaps introduced in
historical times (nat)
species not confirmed to be truly naturalized, including present-day
adventive and escaped plants, or historical collections not recently
vouchered
Island abbreviation codes are compiled here and are also located at the
foot of each checklist page. East Maui and West Maui were separated out if
they were so listed in a publication. However, if a distribution is listed as
“Maui,” it cannot be assumed that the taxon has been documented by
vouchers on both East and West Maui, and no effort was made to clarify
that status for this edition of the checklist. No effort was made to similarly
designate distribution by region on other islands, (e.g., Ko‘olau vs.
Wai‘anae on O‘ahu). Separate distribution lists were included for the small
islands of Ka‘ula and Lehua. Lehua distribution records were infrequently
Likewise, family classifications for ferns and lycophytes follows that
used in Palmer (2005). More modern classifications (e.g., Smith et al. 2006,
Christenhusz et al. 2011) will be adopted in due time. For now, the reader
can refer to the Smithsonian’s latest supplement (mentioned earlier), which
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Hawaiian Native & Naturalized Vascular Plant Checklist
included in the Manual, and the bulk of distribution records included in this
checklist were derived from Wood & LeGrande (2006). Distribution
records from Ka‘ula were not included in the Manual, and the records listed
here were included on the basis on voucher specimens stored in Herbarium
Pacificum, Bishop Museum. Voucher records from Molokini, and islet of
similar size off East Maui, were not included in this edition of the checklist.
Main Hawaiian Islands
EM
East Maui
H
Hawai‘i
K
Kaua‘i
Ka
Kaho‘olawe
Kl
Ka‘ula Rock
L
Läna‘i
Le
Lehua
M
Maui
Mo
Moloka‘i
Ni
Ni‘ihau
O
O‘ahu
WM West Maui
Maui as forestry plantings with no vouchered record of naturalization]. By
keeping these taxa in the checklist for now, the user can remain aware of
them, and their status can perhaps more readily be clarified.
• Explanation of categories included in the Citation field:
COMBNOV (combinatio nova, new combination) = a new combination
created when a taxon is transferred from one genus to another and the same
specific epithet is used. Ex. Canthium odoratum (G. Forst.) Seem. =
Psydrax odorata (G. Forst.) A. C. Sm. & S. P. Darwin; Lobelia
gaudichaudii (A. DC.) subsp. koolauensis (Hosaka & Fosberg) = L.
koolauense (Hosaka & Fosberg) Lammers [also a STATNOV, in this case].
Northwestern Hawaiian Islands
FF
French Frigate Shoals
GP
Gardner Pinnacles
Ku
Kure
La
Laysan
Li
Lisianski
Mi
Midway
N
Nihoa
Ne
Necker
PH
Pearl & Hermes
DESCR (description) = the cited source includes a description of the taxon;
the quality may range from a few essential diagnostic characters to a
detailed botanical description.
EXTINCT? = published record announcing the likelihood that a taxon is
extinct, either throughout its range or on a particular island. It is not a
primary role of this checklist to monitor all reports of likely plant
extinctions, a function more efficiently monitored locally by rare plant
organizations.
• Citations
The citation field provides a chronological, citation-based account of
changes in island distribution and scientific names for each taxon dating
back to Wagner et al. (1990 [for dicot and monocot taxa]) and Palmer
(2003 [for ferns and lycophytes]), adding new island records, new
naturalized records, and newly described native taxa as they become
known. In most cases, a specific page number for the citation is included to
aid quick access to information.
KEY = the cited source includes a key that treats the taxon; the key may
range in scope from a few closely related taxa in a genus to a regional key
including extra-Hawaiian taxa.
misapplied = a scientific name that at some point in time is incorrectly
applied in a sense different from that of the original author. Such names
begin as misidentifications but can become misapplied by virtue of
subsequent widespread adoption or perpetuation in publications. The
misapplication can be widespread in scope (as with some popular but
misidentified cultivated plants) or restricted to a small region (e.g.,
misidentification of the Hawaiian endemic tree fern Cibotium menziesii as
C. chamissoi by Skottsberg in 1942). For simplification here, any
misidentifications that are published locally (not in widely available
publications) and later corrected are treated here as misapplied names, using
the format incorrect name sensu author/year citation, non author citation
for authentic taxon.
An added feature of this edition of the checklist is the inclusion of nonnative taxa that are not currently considered naturalized, but that have, in
the recent or distant past, been noted as such in the literature, or as
displaying tendencies toward becoming naturalized. These taxa are
designated by a “?” in the status column if they are not currently
documented as being truly naturalized throughout their range in the state. If
the taxon is naturalized on at least one island, but questionably naturalizing
on others, the “?” immediately follows the island designation, followed by a
parenthetical explanation for the designation [e.g., Eucalyptus cladocalyx
(Myrtaceae, page A-122) is naturalized on Hawai‘i, but noted on O‘ahu and
new island record = these are generally noted by island abbreviations in
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Hawaiian Native & Naturalized Vascular Plant Checklist
parentheses following a bibliographic citation (e.g., Herbst & Wagner
1996:9 (WM) notes a new island record from West Maui on page 9 of their
paper).
undulata subsp. kauaiensis becoming D. kauaiensis (Lammers 2005).
SSPNOV = a newly described subspecies.
NEWNAT (new naturalization) = a newly documented naturalized record
for the state; includes both records for taxa previously known from in
Hawai‘i but not in a naturalized condition (i.e., new naturalized record), and
those previously unknown in the state (i.e., new state record).
Syn. (synonymum, synonym) = a taxonomic name rejected in favor of
another name. Two kinds: nomenclatural synonyms are based on the same
original type material. Ex. Evodia molokaiensis (Hillebr.) Drake, Pelea
molokaiensis Hillebr., and Melicope molokaiensis (Hillebr.) T. G. Hartley &
B. C. Stone are all differently named combinations for the same plant
originally described by Hillebrand in 1888; if any of the three is currently
considered the accepted name, the other two automatically become
nomenclatural synonyms. Taxonomic synonyms are based on different
original type material, and their synonymization is based on the botanist’s
taxonomic judgement. By botanical code, the oldest available validly
published legitimate name must be used, and all others become taxonomic
synonyms. Using the previous example, Pelea foetida H. Lév. (published
1911) and P. stellata H. St. John (publ. 1944) are taxonomic synonyms of
Pelea molokaiensis Hillebr. (publ. 1888) [which itself is now a
nomenclatural synonym of Melicope molokaiensis (Hillebr.) T. G.Hartley &
B. C.Stone].
NOM = general indicator that a name change was made, usually a
synonymization or correction of a misapplied name.
NOMNOV (nomen novum, new name) = a newly coined name necessitated
for nomenclatural reasons; it is not a new taxon. Ex. When the genus
Rollandia was synonymized into Cyanea, the binomial Rollandia
angustifolia (Hillebr.) Rock should have become a COMBNOV as Cyanea
angustifolia. Because the latter name had already been published for a
different species [C. angustifolia (Cham.) Hillebr.], however, Lammers
(1993) was able to coin a new epithet for the taxon, C. koolauensis
Lammers, Givnish & Sytsma.
REDISCOVER = published record noting that a taxon thought to be
extinct has been relocated; could refer to rediscovery on a single island of a
multi-island taxon. However, it is not a primary role of this checklist to
report on all rare plant rediscoveries, a function more comprehensively
monitored locally by rare plant organizations.
VARNOV = a newly described variety.
•
Navigating the Citation field
The citation field included in Appendices A & B is intended to provide
a running history of a taxon in the Hawaiian Islands, focusing especially on
changes in scientific names and island distribution. Because the starting
point for this checklist is Wagner et al. (1990), the initial citation for most
of the dicot and monocot taxa is from the Manual; similarly, Palmer (2005)
is the first mentioned citation for most of the fern and lycophyte taxa. The
first page on which the taxon is treated is given, along with its island
distribution.
RESURRECT = a taxon previously treated as a synonym is restored to
recognized status; restored status may involve a COMBNOV or
STATNOV. Ex. Stenogyne sherffii O. Deg., a synonym of S. kaalae Wawra
in Wagner et al. (1990), was resurrected (Wagner & Weller 1999) as a
subspecies, S. kaalae subsp. sherffii (O. Deg.) W. L. Wagner & Weller
(both a COMBNOV and STATNOV).
Ex. 1. Wagner et al. 1990:786 (K, O)
SPNOV (species nova, new species) = a newly described species, based on
an original type.
This is a standard opening citation for taxa treated in the Manual. This
entry, for Scaevola glabra (Dicot, Goodeniaceae, p. A-99), indicates that
the species description starts on page 786, and that it occurs on Kaua‘i and
O‘ahu. No name or distributional changes have occurred since 1990.
STATNOV (status novus, new status) = indicates that a name change
involves a change in the rank (status) of a described taxon, such as a change
from subspecific or varietal rank to full species, or vice versa. Ex. Delissea
Palmer 2003:234 (K)
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Hawaiian Native & Naturalized Vascular Plant Checklist
Standard opening citation for taxa treated in Palmer’s fern guide. This
entry, for Sadleria unisora (Fern, Dicksoniaceae, p. A-235), is described on
page 234 and occurs on Kaua‘i. Again, no name or distributional changes
have occurred since 1990.
Ex. 5. Wagner et al. 1990:451 (as Cyanea grimesiana var. mauiensis,
syn. of C. grimesiana subsp. grimesiana); Lammers 1998:31
(RESURRECT/M, COMBNOV & STATNOV:Syn. C. grimesiana var.
mauiensis)
All such Wagner et al. (1990) and Palmer (2003) entries are accompanied
by full descriptions and relevant keys, thus those citations are not annotated
with “DESCR” or “KEY.” (but see Ex. 3).
In this entry, the currently accepted species (Cyanea mauiensis, Dicot,
Campanulaceae, p. A-48), was originally a synonym in the Manual under a
different name (C. grimesiana var. mauiensis). When Lammers resurrected
the taxon in 1998, he elevated it to full species status under a new
combination.
Ex. 2. Wagner et al. 1990:234 (O [as Tetraplasandra gymnocarpa]);
Lowry & Plunkett 2010:74 (COMBNOV:Syn. T. gymnocarpa)
Ex. 6. Oppenheimer 2003:5 (NEWNAT/H, DESCR); Staples & Herbst
2005:99 (KEY), 101 (DESCR)
This entry for Polyscias gymnocarpa (Dicot, Araliaceae, p. A-11) notes that
this species from O‘ahu was known as Tetraplasandra gymnocarpa in the
Manual, but was later recombined in 2010 under the genus Polyscias.
Celosia argentea (Dicot, Amaranthaceae, p. A-5), was published as a new
naturalization record for the Big Island and the state in 2003, accompanied
by a description. The species was later treated in A Tropical Garden Flora
(Staples & Herbst 2005), in which a key and description were included. The
latter citation is a good indicator that the taxon is cultivated in the Hawaiian
Islands.
Ex. 3. Wagner et al. 1990:1554 (Mo?, L? [adventive], DESCR);
Wagner & Herbst 1995:24 (NEWNAT/Mo, L); Herbst & Clayton 1998:28
(KEY)
This entry, for Hyparrhenia hirta (Monocot, Poaceae, p. A-216), introduces
a category of plants that were included in the Manual but not given full
treatment. .This situation applies when all island distributions in the Manual
citation are accompanied by question marks, for reasons usually provided in
brackets. In the case above, the species was considered to be adventive (not
fully naturalized) on Moloka‘i and Läna‘i. Later, in 1995, populations on
those two islands were published as the first records of naturalization of H.
hirta in the state. It was included in a species key in the 1998 Herbst &
Clayton paper. If diagnostic characters are provided for these kinds of
Manual records, they are annotated with “DESCR.”
Ex. 7. Wagner & Lorence 1998:311 (SPNOV/K, DESCR)/Herbst &
Wagner 1999:31 [as Hedyotis flynnii]; Terrell et al. 2005:833/Kennedy et
al. 2010:24 (COMBNOV:Syn. H. flynnii)
Hedyotis flynnii (Dicot, Rubiaceae, p. A-149) was described as a new
species from Kaua‘i in 1998 (secondarily reported by Herbst & Wagner in
1999). In 2005, the species was transferred to Kadua (secondarily reported
by Kennedy et al. in 2010).
Ex. 8. St. John 1987b:344 (SPNOV/K [as Delissea kolekoleensis],
DESCR); Lammers 1992:130 (COMBNOV:Syn. D. kolekoleensis, KEY,
DESCR)/ Herbst & Wagner 1999:16; Wood 2012:92 (EXTINCT?,
DESCR)
Ex. 4. Wagner et al. 1990:170 (O [as Blechum brownei]); Wagner &
Herbst 1995:14 (H, NOM:Syn. B. brownei = B. pyramidatum); Tripp et al.
2009:900/Wagner et al. 2012:3 (NOM:Syn. B. pyramidatum)
This entry for Ruellia blechum (Dicot, Acanthaceae, p. A-2) involves
multiple name changes since it was published in the Manual as Blechum
brownei. The name was subsequently synonymized with B. pyramidatum,
and the latter was yet again synonymized into Ruellia blechum. The formal
proposal to transfer Blechum into Ruellia is made in Tripp et al. (2009);
Wagner et al. (2012) verifies the acceptance of this name change for
Hawaiian material.
This citation, for Cyanea kolekoleensis (Dicot, Campanulaceae, p. A-46),
showcases one of Harold St. John’s newly published species from 1987, too
late for the Manual to pass judgement on the taxon. Lammers in 1992
accepted the species, but tranferred it from Delissea to Cyanea. In 2012,
Wood speculated that the species might be extinct.
Ex. 9. Wagner et al. 1990:1080 (K); St. John 1987g:44 (SPNOV/K [as
Lysimachia kahiliensis], DESCR); Marr & Bohm 1997:265 ([as L.
5
Hawaiian Native & Naturalized Vascular Plant Checklist
daphnoides] KEY, DESCR, 272 [as L. kahiliensis] KEY, DESCR); Herbst
& Wagner 1999:30 (NOM:Syn. L. kahiliensis)
Owing to the sorting routines used, hybrid taxa with an “x” immediately
preceding the specific epithet have sorted ahead of all non-hybrid species,
and in no particular alphabetical sequence (e.g., Asplenium, p. A-230–231).
An index to accepted families and species is provided, beginning on page
16, preceding the appendices. Pagination is provided for the first page on
which an accepted family or genus is treated in Appendices A and B, the
main body of checklist, which may assist the user in navigating the
checklist.
In a variation of Ex. 8, Lysimachia daphnoides (Dicot, Primulaceae, p. A140) was treated in the Manual. One of St. John’s new species from 1987
was L. kahiliensis, published too late for consideration in the Manual
treatment. In a 1997 revision of Hawaiian Lysimachia, Marr & Bohm
accepted L. kahiliensis as a good species. Herbst & Wagner (1999)
disagreed, considering L. kahiliensis a synonym of L. daphnoides.
Table 1. Numbers of families, genera, and taxa by major group
GymnoFerns/
Dicots Monocots
sperms Lycophytes
Families
134
33
5
29
Genera
695
227
10
71
Total taxa
2130
583
19
251
Ex. 10. Herbst & Clayton 1998:36 (NEWNAT/L [as Sporobolus
elongatus], KEY; NEWNAT/H [as S. pyramidalis], KEY); Snow 2008:41
(O [NOM: S. pyramidatus misapplied, in part], H [NOM: S. pyramidalis
misapplied])
This entry for Sporobolus elongatus (Monocot, Poaceae, p. A-226) includes
misapplied names. In 1998, Sporobolus elongatus (on Läna‘i) and S.
pyramidalis (on Hawai‘i) were both treated as new naturalizations in the
state. In 2008, Snow added O‘ahu to the distribution of S. elongatus, based
on a specimen collected on the University of Hawai‘i, Mänoa campus,
previously misidentified as S. pyramidatus (authentic S. pyramidatus
remaining a part of the naturalized Hawaiian flora). Snow also added
Hawai‘i island to the distribution of S. elongatus, based on reidentification
of the specimen misidentified as S. pyramidalis earlier in the entry.
Sporobolus pyramidalis was thus removed as a naturalized element in the
Hawaiian flora. Associated entries in Appendices C and D provide more
details for the misapplied names.
•
Total
201
1003
2983
Table 2. Origin status of included taxa by major group
GymnoFerns/
Dicots Monocots
sperms Lycophytes Total
end/end?
taxa
963
113
0
168
1244
ind/ind?
taxa
61
38
0
43
142
nat/nat?
taxa
1010
375
16
41
1442
pol/pol?
taxa
14
13
0
0
27*
? taxa
81
44
3
0
128
Total taxa
2130
583
19
251
2983
*Total includes 3 species that have yet to be documented as naturalizing:
Artocarpus altilis, Dioscorea alata, and Lagenaria siceraria.
Additional abbreviations
coll(s.) = collection(s)
ex. = example
f. = filius
fma. = forma
hwy = highway
incl. = including
sp. = species (singular)
spm(s.) = specimen(s)
spp. = species (plural)
subsp. = subspecies (singular)
subspp. = subspecies (plural)
var. = variety
Table 3. Numbers of native vs. non-native taxa by major group
GymnoFerns/
Dicots Monocots sperms Lycophytes Total
native taxa
1024
151
0
211
1386
non-native
taxa
1106
432
19
40
1597
Total taxa
2130
583
19
251
2983
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Hawaiian Native & Naturalized Vascular Plant Checklist
ACKNOWLEDGMENTS
Work on this checklist update and revision was supported by National
Science Foundation Biological Research Collections grant DEB-1057453
and by the Elizabeth Loy McCandless Marks Endowment Fund. Warren L.
Wagner, Smithsonian Institution, is thanked for continuing to maintain the
excellent Flora of the Hawaiian Islands website, the source for many of the
updates included in this checklist. George Staples and Derral Herbst
provided much-appreciated botanical mentoring over the years. In-house
support from the hard-working Herbarium Pacificum staff, led by Barbara
Kennedy, Shelley James, and Napua Harbottle was invaluable, as was front
office support from Tracie Mackenzie and Leslie Santos. Bishop Museum
Research Associates Alex Lau and Danielle Frohlich are thanked for
reviewing the manuscript. Special thanks to Neal Evenhuis, co-editor of
Hawaii Biological Survey publications, for encouraging publication of this
checklist.
This checklist update was made possible only because of the
outstanding fieldwork, research, and taxonomic interpretation provided by
the list of authors included in the bibliography. All are here acknowledged
in appreciation for helping to advance understanding of the unique and
fragile Hawaiian flora. Locally, much fieldwork has been conducted
statewide on the distribution of naturalized plants subsequent to publication
of the Manual. The past and present list of field biologists most busily
documenting them include Randy Bartlett, Keali‘i Bio, Pat Conant, Curt
Daehler, Tim Flynn, Danielle Frohlich, Derral Herbst, Guy Hughes, Shelley
James, Barbara Kennedy, Alex Lau, Lloyd Loope, Dave Lorence, J. Scott
Meidell, Ken Nagata, Hank Oppenheimer, James Parker, Bobby Parsons,
Linda Pratt, Neil Snow, George Staples, Forest & Kim Starr, Warren L.
Wagner, Ken Wood, and Mike Wysong. The staffs of the O‘ahu Invasive
Species Committee, O‘ahu Army Natural Resources Program, and the Plant
Pest Control Branch of the Hawai‘i State Department of Agriculture are
thanked for bringing to light many new naturalizing plant taxa.
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Hawaiian Native & Naturalized Vascular Plant Checklist
Brummitt, R. K. and C. E. Powell (eds.). 1992. Authors of Plant Names.
Kew, Royal Botanic Gardens, 732 pp.
Bruneau, A., M. Mercure, G. P. Lewis and P. S. Herendeen. 2008.
Phylogenetic patterns and diversification in caesalpinioid legumes.
Botany 86: 697–718.
Burney, D. A., H. F. James, L. P. Burney, S. L. Olson, W. Kikuchi, W. L.
Wagner, M. Burney, D. McCloskey, D. Kikuchi, F. V. Grady, R. Gage,
II and R. Nishek. 2001. Fossil evidence for a diverse biota from Kaua'i
and its transformation since human arrival. Ecological Monographs
71(4): 615–641.
Carr, G. D. 1995. A fully fertile intergeneric hybrid derivative from
Argyroxiphium sandwicense ssp. macrocephalum x Dubautia menziesii
(Asteraceae) and its relevance to plant evolution in the Hawaiian
Islands. American Journal of Botany 82(12): 1574–1581.
Carr, G. D. 1998. Another new species of Dubautia (Asteraceae--Madiinae)
from Kaua‘i, Hawaiian Islands. Novon 8(1): 8–11.
Carr, G. D. 1999. A reassessment of Dubautia (Asteraceae: HeliantheaeMadiinae) on Kaua‘i. Pacific Science 53(2): 144–158.
Carr, G. D. and D. H. Lorence. 1998. A new species of Dubautia
(Asteraceae--Madiinae) from Kaua‘i, Hawaiian Islands. Novon 8(1):
4–7.
Catalán, P., R. J. Soreng and P. M. Peterson. 2009. Festuca aloha and F.
molokaiensis (Poaceae: Loliinae), two new species from Hawai‘i.
Journal of the Botanical Research Institute of Texas 3(1): 51–58.
Chemisquy, M. A., L. M. Giussani, M. A. Scataglini, E. A. Kellogg and O.
Morrone. 2010. Phylogenetic studies favour the unification of
Pennisetum, Cenchrus, and Odontelytrum (Poaceae): a combined
nuclear, plastid and morphological analysis, and nomenclatural
combinations in Cenchrus. Annals of Botany, London 106: 107–130.
Chinnock, R. J. 2007. Eremophila and allied genera -- Monograph of the
Myoporaceae. Australia, Rosenberg Publishing Pty Ltd., 677 pp.
Christenhusz, M. J. M. 2011. (2034) Proposal to conserve Blechnum
hawaiianum against B. norfolkense (Blechnaceae, Pteridophyta). Taxon
60(6): 1772.
Christenhusz, M. J. M., X.-c. Zhang and H. Schneider. 2011. A linear
sequence of extant families and genera of lycophytes and ferns.
Phytotaxa 19: 7–54.
Clayton, W. D. and N. Snow. 2010. A Key to Pacific Grasses. Royal
Botanic Gardens, Kew, Kew Publishing, 107 pp.
Bibliographic citations
The author-year citations in the “Citation” field are linked to the following
alphabetical list (by author) of full bibliographic citations. Many of the
papers appearing in the Bishop Museum Occasional Papers series are
available as pdfs at http://hbs.bishopmuseum.org/hbspubs.html, in the
Records of the Hawaii Biological Survey section.
Ackerman, J. D., W. Falcon and W. Recart. 2011. New records of
naturalized orchids for the Hawaiian Islands. Bishop Museum
Occasional Papers 110: 1–4.
Aedo, C. and F. M. Garmendia. 1997. Twelve new names in Geranium L.
(Geraniaceae). Kew Bulletin 52(3): 725–727.
Aguraiuja, R. and K. R. Wood. 2003. Diellia mannii (D. C. Eaton) Robins.
(Aspleniaceae) rediscovered in Hawai‘i. American Fern Journal 93(3):
154–156.
Alford, M. H. 2012. New records of Gamochaeta (Asteraceae) in the
Hawaiian archipelago. Bishop Museum Occasional Papers 113: 1–6.
Al-Shehbaz, I. A. 1985. The genera of Brassiceae (Cruciferae;
Brassicaceae) in the southeastern United States. Journal of the Arnold
Arboretum 66: 279–351.
Al-Shehbaz, I. A. and R. A. Price. 1998. Delimitation of the genus
Nasturtium (Brassicaceae). Novon 8: 124–126.
Baldwin, B. G. and G. D. Carr. 2005. Dubautia kalalauensis, a new species
of the Hawaiian silversword alliance (Compositae, Madiinae) from
northwestern Kaua‘i, U.S.A. Novon 15(2): 259–263.
Baldwin, B. G. and E. A. Friar. 2010. Dubautia carrii and D. hanaulaensis,
new species of the Hawaiian silversword alliance (Compositae,
Madiinae) from Moloka‘i and Maui. Novon 20(1): 1–8.
Barkworth, M. E. 1990. Nassella (Gramineae, Stipeae): revised
interpretation and nomenclatural changes. Taxon 39: 597–614.
Bruegmann, M. M. 1996. Endangered and threatened wildlife and plants;
determination of endangered status for three plant species (Cyanea
dunbarii, Lysimachia maxima, and Schiedea sarmentosa) from the
Island of Molokai, Hawaii. Federal Register 61(198): 53130–53137.
Bruegmann, M. M. 1999. New records of flowering plants on Midway
Atoll. Bishop Museum Occasional Papers 59: 1–2.
Brummitt, R. K. 2011. Report of the nomenclature committee for vascular
plants: 63; proposals to conserve or reject names; (1890) To conserve
Bonamia menziesii A. Gray with conserved type (Convolvulaceae)
Taxon 60(4): 1202–1203.
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Hawaiian Native & Naturalized Vascular Plant Checklist
Clemants, S. E. and S. L. Mosyakin. 2003. Dysphania. Flora of North
America 4: 267–275.
Conant, P. 1996. New Hawaiian pest plant records for 1995. Bishop
Museum Occasional Papers 46: 1–2.
Conant, P. 1998. Range extensions for melastomes in Hawai‘i. Bishop
Museum Occasional Papers 56: 1–2.
Coppen d’Eeckenbrugge, G., V. E. Barney, P. M. Jorgensen and J. M.
MacDougal. 2001. Passiflora tarminiana, a new cultivated species of
Passiflora subgenus Tacsonia (Passifloraceae). Novon 11(1): 8–15.
Daehler, C.C. and R.F. Baker. 2006. New records of naturalized and
naturalizing plants around Lyon Arboretum, Manoa Valley, O‘ahu.
Bishop Museum Occasional Papers 87: 3–18.
Darbyshire, S. J., H. E. Connor and B. Ertter. 2010. The genus
Rytidosperma (Poaceae) in the United States of America. Journal of the
Botanical Research Institute of Texas 4(2): 663–676.
DeFelice, R. C. 1999. A new distributional record for Halophila hawaiiana
Doty & B. Stone (Hydrocharitaceae) in Hawai‘i. Bishop Museum
Occasional Papers 59: 2–3.
Dougill, S. J. and P. C. Banko. 2002. A new genus of alien grass discovered
in Hawai‘i: Oryzopsis (Poaceae). Newsletter of the Hawaiian Botanical
Society 41(1, 2): 8.
Dragon, J. A. and D. S. Barrington. 2009. Systematics of the Carex
aquatilis and C. lenticularis lineages: geographically and ecologically
divergent sister clades of Carex section Phacocystis (Cyperaceae).
American Journal of Botany 96: 1896–1906.
Du Puy, D. J., J.-N. Labat and B. D. Schrire. 1993. The separation of two
previously confused species in the Indigofera spicata complex
(Leguminosae: Papilionoideae). Kew Bulletin 48: 727–733.
Ebihara, A., J.-Y. Dubuisson, K. Iwatsuki, S. Hennequin and M. Ito. 2006.
A taxonomic revision of Hymenophyllaceae. Blumea 51: 221–280.
Evenhuis, N.L. and L.G. Eldredge (eds.). 2004. Natural History of Nihoa
and Necker Islands. Bishop Museum Press, Honolulu, HI, 220 pp.
Ezcurra, C. 1993. Systematics of Ruellia (Acanthaceae) in southern South
America. Annals of the Missouri Botanical Garden 80: 787–845.
Fantz, P. R. 1996. Taxonomic notes on the Centrosema pubescens Bentham
complex in Central America (Leguminosae: Phaseoleae: Clitoriinae).
Sida 17(2): 321–332.
Florence, J. 1997. Flore de la Polynésie Française, vol. 1. ORSTOM, Paris,
393 pp.
Flynn, T. and D. H. Lorence. 1998. New naturalized plant records for the
Hawaiian Islands. Bishop Museum Occasional Papers 56: 5–6.
Flynn, T. and D. H. Lorence. 2002. Additions to the flora of the Hawaiian
Islands. Bishop Museum Occasional Papers 69: 14–16.
Frohlich, D. and A. Lau. 2007. New plant records from O‘ahu for 2006.
Bishop Museum Occasional Papers 96: 8–13.
Frohlich, D. and A. Lau. 2008. New plant records from O‘ahu for 2007.
Bishop Museum Occasional Papers 100: 3–12.
Frohlich, D. and A. Lau. 2010. New plant records from O‘ahu for 2008.
Bishop Museum Occasional Papers 107: 3–18.
Frohlich, D. and A. Lau. 2012. New plant records for the Hawaiian Islands
2010–2011. Bishop Museum Occasional Papers 113: 27–54.
Fryxell, P. A. 2001. Talipariti (Malvaceae), a segregate from Hibiscus.
Contributions from the University of Michigan Herbarium 23: 225–
270.
Gemmill, C. E. C. 1998. A new narrow, endemic species of Pritchardia
(Arecaceae) from Kaua‘i, Hawaiian Islands. Novon 8(1): 18–22.
Govaerts, R. and A. Radcliffe-Smith. 1996. New names and combinations
in Euphorbiaceae - Phyllanthoideae. Kew Bulletin 5(1): 175–178.
Govaerts, R., D. G. Frodin and A. Radcliffe-Smith. 2000. World Checklist
and Bibliography of Euphorbiaceae (with Pandaceae). Kew, England,
Royal Botanic Gardens, Kew.
Grayer, R. J., M. R. Eckert, A. Lever, N. C. Veitch, G. C. Kite and A. J.
Paton. 2010. Distribution of exudate flavonoids in the genus
Plectranthus. Biochemical Systematics and Ecology 38(3): 335–341.
Green, P. S. 1991. The Indo-Pacific strand Scaevola (Goodeniaceae) - yet
again! Taxon 40(1): 118–122.
Green, P. S. 1994. Asteraceae. Flora of Australia 49: 362–403.
Green, P. S. and G. E. Wickens. 1989. The Olea europaea complex. The
Davis & Hedge festchhrift: plant taxonomy, phytogeography and
related subjects. K. Tan. Edinburgh, University Press: pp. 287–299.
Grey-Wilson, C. 1982. Balsaminaceae. Flora of Tropical East Africa 45.
Groeninckx, I., S. Dessein, H. Ochoterena, C. Persson, T. J. Motley, J.
Kårehed, B. Bremer, S. Huysmans and E. Smets. 2009. Phylogeny of
the herbaceous tribe Spermacoceae (Rubiaceae) based on plastid DNA
data. Annals of the Missouri Botanical Garden 96(1): 109–132.
Gu, H. and P. C. Hoch. 1997. Systematics of Kalimeris (Asteraceae:
Astereae). Annals of the Missouri Botanical Garden 84: 762–814.
Harbaugh, D. T., H. L. Oppenheimer, K. R. Wood and W. L. Wagner.
2010. Taxonomic revision of the endangered Hawaiian red-flowered
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Hawaiian Native & Naturalized Vascular Plant Checklist
sandalwoods (Santalum) and discovery of an ancient hybrid species.
Systematic Botany 35(4): 827–838.
Harley, R. and A. Paton. 2001. Leonurus japonicus Houtt. (Labiatae): the
correct name for a common tropical weed. Kew Bulletin 56: 243–244.
Hartley, T. G. 2001. On the taxonomy and biogeography of Euodia and
Melicope (Rutaceae). Allertonia 8(1): 1–319.
Hartley, T. G. and B. C. Stone. 1989. Reduction of Pelea with new
combinations in Melicope (Rutaceae). Taxon 38(1): 119–123.
Havran, J. C., H. Oppenheimer, J. Keaton and K. Piotrowski. 2012.
Interisland range expansion of Viola lanaiensis (Violaceae:
Malpighiales), an endangered Hawaiian violet. Pacific Science 66(4):
447–456.
Hawley, B. M. and B. L. Pyle. 1995. New Hawaiian plant records. I. Bishop
Museum Occasional Papers 42: 1.
Haynes, R. R., D. H. Les and M. Kral. 1998. Two new combinations in
Stuckenia, the correct name for Coleogeton (Potamogetonaceae).
Novon 8(3): 241.
Heads, M. J. 1996. Biogeography, taxonomy and evolution in the Pacific
genus Coprosma (Rubiaceae). Candollea 51(2): 381–405.
Herbarium Pacificum Staff. 1996. New Hawaiian plant records for 1995.
Bishop Museum Occasional Papers 46: 3–8.
Herbarium Pacificum Staff. 1997. New Hawaiian plant records for 1996.
Bishop Museum Occasional Papers 49: 18–19.
Herbarium Pacificum Staff. 1998. New Hawaiian plant records for 1997.
Bishop Museum Occasional Papers 56: 8–15.
Herbarium Pacificum Staff. 1999. New Hawaiian plant records for 1998.
Bishop Museum Occasional Papers 58: 3–11.
Herbst, D. R. 1998. New records for Hawaiian plants. I. Bishop Museum
Occasional Papers 56: 2–4.
Herbst, D. R. and W. D. Clayton. 1998. Notes on the grasses of Hawai‘i:
new records, corrections, and name changes. Bishop Museum
Occasional Papers 55: 17–38.
Herbst, D. R., G. W. Staples and C. T. Imada. 2004. New Hawaiian plant
records for 2002–2003. Bishop Museum Occasional Papers 78: 3–12.
Herbst, D. R. and W. L. Wagner. 1996. Contributions to the flora of
Hawai‘i. V. Bishop Museum Occasional Papers 46: 8–12.
Herbst, D. R. and W. L. Wagner. 1999. Contributions to the flora of
Hawai‘i: VII. Bishop Museum Occasional Papers 58: 12–36.
Hill, K. D. and L. A. S. Johnson. 1995. Systematic studies in the eucalypts
7. A revision of the bloodwoods, genus Corymbia (Myrtaceae).
Telopea 6(2–3): 185–504.
Hodel, D. R. 2007. A review of the genus Pritchardia. Palms 51(4): Special
Supplement S-1–53.
Hodel, D. R. 2009. A new species of Pritchardia and the rediscovery of P.
lowreyana on Oahu, Hawaii. Palms 53(4): 173–179.
Hovenkamp, P.H. and F. Miyamoto. 2005. A conspectus of the native and
naturalized species of Nephrolepis (Nephrolepidaceae) in the world.
Blumea 50: 279–322.
Huber, H. 1987. Piperaceae, pp. 272–300 in Dassanayake, M. D., and F. R.
Fosberg (eds.), A Revised Handbook to the Flora of Ceylon, vol. 6.
New Delhi, Amerind Publ. Co.
Hughes, G. D. 1995. New Hawaiian plant records. II. Bishop Museum
Occasional Papers 42: 1–10.
Imada, C. T. 2007. New Hawaiian plant records for 2005–2006. Bishop
Museum Occasional Papers 96: 34–41.
Imada, C. T., S. A. James and B. H. Kennedy. 2008. New Hawaiian plant
records from Herbarium Pacificum for 2007. Bishop Museum
Occasional Papers 100: 12–16.
Imada, C. T., G. W. Staples and D. R. Herbst. 2000. New Hawaiian plants
records for 1999. Bishop Museum Occasional Papers 63: 9–16.
Imada, C. T., W. L. Wagner and D. R. Herbst. 1989. Checklist of native and
naturalized flowering plants of Hawai‘i. Bishop Museum Occasional
Papers 29: 31–87.
Jankalski, S. 2008. Subgenera and new combinations in Dracaena.
Sansevieria 18: 17–21.
Jeffrey, C. 1986. The Senecioneae in East Tropical Africa. Notes on
Compositae: IV. Kew Bulletin 41(4): 873–943.
Kato, M. 2001. Deparia cataracticola (Woodsiaceae), a new species from
Hawaii. Acta Phytotaxonomica et Geobotanica 52(1): 1–9.
Kennedy, B. H., S. A. James and C. T. Imada. 2010. New Hawaiian plant
records from Herbarium Pacificum for 2008. Bishop Museum
Occasional Papers 107: 19–26.
Khan, R. and C. E. Jarvis. 1989. The correct name for the plant known as
Pluchea symphytifolia (Miller) Gillis (Asteraceae). Taxon 38(4): 659–
662.
Kiehn, M. and C. T. Imada. 2010. Psychotria greenwelliae Fosberg — not
on O‘ahu! Bishop Museum Occasional Papers 107: 27–28.
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Hawaiian Native & Naturalized Vascular Plant Checklist
King, R. M. and H. Robinson. 1987. The genera of the Eupatorieae
(Asteraceae). Monographs in Systematic Botany from the Missouri
Botanical Garden 22: i–ix, 1–581.
Koopman, W.J.M., M. J. Zevenbergen, and R. G. van den Berg. 2001.
Species relationships in Lactuca s.l. (Lactuceae, Asteraceae) inferred
from AFLP fingerprints. American Journal of Botany 88: 1881–1887.
Koyama, T., B. H. Kennedy and C. T. Imada. 2010. On the identity of
Gahnia lanaiensis O. Deg, I. Deg. & J. Kern (Cyperaceae) of Hawai‘i.
Bishop Museum Occasional Papers 107: 29–32.
Kraus, F. 2003. New records of alien plants and animals in Hawai‘i. Bishop
Museum Occasional Papers 74: 76–78.
Labiak, P. H. 2011. Stenogrammitis, a new genus of grammitid ferns
segregated from Lellingeria (Polypodiaceae). Brittonia 63(1): 139–149.
Lammers, T. G. 1991. Systematics of Clermontia (CampanulaceaeLobelioideae). Systematic Botany Monographs 32: 1–97.
Lammers, T. G. 1992. Two new combinations in the endemic Hawaiian
genus Cyanea (Campanulaceae: Lobelioideae). Novon 2(2): 129–131.
Lammers, T. G. 1996. A new linear-leaved Cyanea (Campanulaceae:
Lobelioideae) from Kaua‘i, and the “rediscovery” of Cyanea
linearifolia. Brittonia 48(2): 237–240.
Lammers, T. G. 1998. New names and new combinations in
Campanulaceae. Novon 8(1): 31–35.
Lammers, T. G. 1999. A new Lobelia from Mexico, with additional new
combinations in world Campanulaceae. Novon 9(3): 381–389.
Lammers, T. G. 2004. Five new species of the endemic Hawaiian genus
Cyanea (Campanulaceae: Lobelioideae). Novon 14(1): 84–101.
Lammers, T. G. 2005. Revision of Delissea (Campanulaceae-Lobelioideae).
Systematic Botany Monographs 73: 1–75.
Lammers, T. G. 2007. Revision of Lobelia sect. Galeatella
(Campanulaceae: Lobelioideae). Journal of the Botanical Research
Institute of Texas 1(2): 789–810.
Lammers, T. G. 2009. Revision of the endemic Hawaiian genus
Trematolobelia (Campanulaceae: Lobelioideae). Brittonia 61(2): 126–
143.
Lammers, T. G., T. J. Givnish and K. J. Sytsma 1993. Merger of the
endemic Hawaiian genera Cyanea and Rollandia (Campanulaceae:
Lobelioideae). Novon 3(4): 437–441.
Lammers, T. G. and D. H. Lorence. 1993. A new species of Cyanea
(Campanulaceae: Lobelioideae) and resurrection of C. remyi. Novon
3(4): 431–436.
Lau, A. and D. Frohlich. 2012. New plant records from O‘ahu for 2009.
Bishop Museum Occasional Papers 113: 7–26.
Lau, J. Q. C. 1995. New Hawaiian plant records. III. Bishop Museum
Occasional Papers 42: 10.
Les, D. H. and R. R. Haynes. 1996. Coleogeton (Potamogetonaceae), a new
genus of pondweeds. Novon 6(4): 389–391.
Loope, L. and A. C. Medeiros. 1992. A new and invasive pampas grass on
Maui. Newsletter of the Hawaiian Botanical Society 31(1): 7–8.
Lorence, D. H. 1996. A new species of Nototrichium (Amaranthaceae) from
Kaua‘i, Hawaiian Islands. Novon 6(1): 64–67.
Lorence, D. H. and T. Flynn. 1997. New naturalized plant records for
Kaua‘i. Bishop Museum Occasional Papers 49: 9–13.
Lorence, D. H. and T. Flynn. 1999. New naturalized plant records for the
Hawaiian Islands. Bishop Museum Occasional Papers 59: 3–6.
Lorence, D. H. and T. Flynn. 2006. New naturalized plant records for
Kaua‘i and Hawai‘i. Bishop Museum Occasional Papers 88: 1–5.
Lorence, D. H., T. Flynn and W. L. Wagner. 1995. Contributions to the
flora of Hawai‘i. III. Bishop Museum Occasional Papers 41: 19–58.
Lorence, D. H. and C. E. C. Gemmill. 2004. Pritchardia flynnii
(Arecaceae), a new endemic species from Kaua‘i, Hawaiian Islands.
Novon 14(2): 185–189.
Lorence, D. H. and S. P. Perlman. 2007. A new species of Cyrtandra
(Gesneriaceae) from Hawai‘i, Hawaiian Islands. Novon 17(3): 357–
361.
Lorence, D. H. and W. L. Wagner. 1995. Another new, nearly extinct
species of Hibiscadelphus (Malvaceae) from the Hawaiian Islands.
Novon 5(2): 183–187.
Lorence, D. H. and W. L. Wagner. 1996. A new species of Chamaesyce
section Sclerophyllae (Euphorbiaceae ) from Kauai. Novon 6(1): 68–
71.
Lorence, D. H., W. L. Wagner and W. G. Laidlaw. 2010. Kadua haupuensis
(Rubiaceae: Spermacoceae), a new endemic species from Kaua‘i,
Hawaiian Islands. Brittonia 62(2): 137–144.
Lorence, D. H. and K. R. Wood. 1994. Kanaloa, a new genus of Fabaceae
(Mimosoideae) from Hawaii. Novon 4(2): 137–145.
Lowry II, P. P. and G. M. Plunkett. 2010. Recircumscription of Polyscias
(Araliaceae) to include six related genera, with a new infrageneric
classification and a synopsis of species. Plant Diversity and Evolution
128(1–2): 55–84.
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Hawaiian Native & Naturalized Vascular Plant Checklist
Lowry II, P. P., & K. R. Wood. 2000. A new, threatened species of
Tetraplasandra (Araliaceae) from Kaua‘i, Hawaiian Islands, and notes
on its conservation status. Novon 10(1): 40–44.
Lye, K. A. 2003. Schoenoplectiella Lye, gen. nov. (Cyperaceae). Lidia 6:
20–29.
Mabberley, D. J. 2008. Mabberley’s Plant-Book: a Portable Dictionary of
Plants, Their Classification and Uses. Cambridge University Press,
Cambridge, 1021 pp.
Mansion, G. 2004. A new classification of the polyphyletic genus
Centaurium Hill (Chironiinae, Gentianaceae): description of the New
World endemic Zeltnera, and reinstatement of Gyrandra Griseb. and
Schenkia Griseb. Taxon 53(3): 719–740.
Marr, K. L. and B. A. Bohm. 1997. A taxonomic revision of the endemic
Hawaiian Lysimachia (Primulaceae) including three new species.
Pacific Science 51(3): 254–287.
Masuyama, S. and Y. Watano. 2010. Cryptic species in the fern
Ceratopteris thalictroides (L.) Brongn. (Parkeriaceae). IV. Taxonomic
revision. Acta Phytotaxonomica et Geobotanica 61(2): 75–86.
McDermid, K. J., M. C. Gregoritza and D. W. Freshwater. 2002. A new
record of a second seagrass species from the Hawaiian Archipelago:
Halophila decipiens Ostenfeld. Aquatic Botany 74: 257–262.
McDermid, K. J., M. C. Gregoritza, J. W. Reeves and D. W. Freshwater.
2003. Morphological and genetic variation in the endemic seagrass
Halophila hawaiiana (Hydrocharitaceae) in the Hawaiian Archipelago.
Pacific Science 57(2): 199–209.
Medeiros, A. C., L. L. Loope, P. Conant and S. McElvaney. 1997. Status,
ecology, and management of the invasive plant Miconia calvescens
DC. (Melastomataceae) in the Hawaiian Islands. Bishop Museum
Occasional Papers 48: 23–36.
Meidell, J. S., H. L. Oppenheimer and R. T. Bartlett. 1997. New plant
records from Pu‘u Kukui watershed and adjacent areas. Bishop
Museum Occasional Papers 49: 17–18.
Meidell, J. S., H. L. Oppenheimer and R. T. Bartlett. 1998. New plant
records from West Maui. Bishop Museum Occasional Papers 56: 6–8.
Middleton, D.J. 2002. Revision of Alyxia (Apocynaceae). Part 2: Pacific
islands and Australia. Blumea 47: 1–93.
Miller, S. E. 2003. Begonia fusca (Begoniaceae), a new naturalized species
for Hawai‘i. Bishop Museum Occasional Papers 74: 22–23.
Molvray, M. 1997. A synopsis of Korthalsella (Viscaceae). Novon 7(3):
268–273.
Morden, C. W. and M. C. Gregoritza. 2005. Population varition and
phylogeny in the endangered Chamaesyce skottsbergii (Euphorbiaceae)
based on RAPD and ITS analyses. Conservation Genetics 6: 969–979.
Mort, M. E., D. E. Soltis, P. S. Soltis, J. Francisco-Ortega and A. Santos
Guerra. 2001. Phylogenetic relationships and evolution of Crassulaceae
inferred from matK sequence data. American Journal of Botany 88:
76–91.
Mosyakin, S. L. and W. L. Wagner. 1998. Notes on two alien taxa of
Rumex L. (Polygonaceae) naturalized in the Hawaiian Islands. Bishop
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Motley, T. J. 1995. Rediscovery of Labordia triflora (Loganiaceae). Pacific
Science 49(3): 221–226.
Motley, T. J. 2005. Tetraplasandra lydgatei (Araliaceae): taxonomic
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Pacific Science 59(1): 105–110.
Munir, A. A. 1992. A taxonomic revision of the genus Stachytarpheta Vahl
(Verbenaceae) in Australia. Journal of the Adelaide Botanic Gardens
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Nagata, K. M. 1995. New Hawaiian plant records. IV. Bishop Museum
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Namoff, S., Q. Luke, F. Jiménez, A. Veloz, C. E. Lewis, V. Sosa, M.
Maunder and J. Francisco-Ortega. 2010. Phylogenetic analyses of
nucleotide sequences confirm a unique plant intercontinental
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Nesom, G. L. 1994. Review of the taxonomy of Aster sensu lato
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Phytologia 77(3): 141–297.
Nesom, G. L. 2000. Generic conspectus of the tribe Astereae (Asteraceae)
in North American and Central America, the Antilles, and Hawaii.
Sida, Botanical Miscellany 20: 1–100.
Nesom, G. L. 2001a. Taxonomic notes on Keysseria and Pytinicarpa
(Asteraceae: Astereae, Lageniferinae). Sida 19(3): 513–518.
Nesom, G. L. 2001b. New records in Pseudognaphalium (Asteraceae:
Gnaphalieae) for the United States. Sida 19(4): 1185–1190.
Nesom, G. L. 2002. Euchiton (Asteraceae: Gnaphalieae) in North American
and Hawaii. Sida 20(2): 515–521.
Nesom, G. L. 2005. Taxonomy of the Symphyotrichum (Aster) subulatum
group and Symphyotrichum (Aster) tenuifolium (Asteraceae: Stereae).
Sida 21: 2125–2140.
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Hawaiian Native & Naturalized Vascular Plant Checklist
Nicolson, D. H. 1996. (1233) Proposal to conserve the name Lagenophora
(Compositae) with a conserved spelling. Taxon 45(May): 341-342.
Oppenheimer, H. L. 2002. The spread of gymnosperms on Maui: a
neglected element of the modern Hawaiian flora. Bishop Museum
Occasional Papers 68: 19–23.
Oppenheimer, H. L. 2003. New plant records from Maui and Hawai‘i
counties. Bishop Museum Occasional Papers 73: 3–30.
Oppenheimer, H. L. 2004. New Hawaiian plant records for 2003. Bishop
Museum Occasional Papers 79: 8–20.
Oppenheimer, H. L. 2006. New Hawai‘i plant records for 2004. Bishop
Museum Occasional Papers 88: 10–15.
Oppenheimer, H. L. 2007. New plant records from Moloka‘i, Läna‘i, Maui,
and Hawai‘i for 2006. Bishop Museum Occasional Papers 96: 17–34.
Oppenheimer, H. L. 2008. New Hawaiian plant records for 2007. Bishop
Museum Occasional Papers 100: 22–38.
Oppenheimer, H. L. 2010. New Hawaiian plant records from Maui County
for 2008. Bishop Museum Occasional Papers 107: 33–40.
Oppenheimer, H. L. 2011. New Hawaiian plant records for 2009. Bishop
Museum Occasional Papers 110: 5–10.
Oppenheimer, H. L. and R. T. Bartlett. 2000. New plant records from Maui,
O‘ahu, and Hawai‘i islands. Bishop Museum Occasional Papers 64: 1–
10.
Oppenheimer, H. L. and R. T. Bartlett. 2002. New plant records from the
main Hawaiian Islands. Bishop Museum Occasional Papers 69: 1–14.
Oppenheimer, H. L., J. S. Meidell and R. T. Bartlett. 1999. New plant
records for Maui and Moloka‘i. Bishop Museum Occasional Papers 59:
7–11.
Oudejans, R.C.H.M. 1989. New names and new combinations in the genus
Euphorbia (Euphorbiaceae). Phytologia 67(1): 43–49.
Palmer, D. D. 2003. Hawai‘i’s Ferns and Fern Allies, University of
Hawai‘i Press, Honolulu, 324 pp.
Palmer, D. D. 2005. Pneumatopteris pendens (Thelypteridaceae) a new
Hawaiian endemic species of Pneumatopteris from Hawaii. American
Fern Journal 95(2): 80–83.
Parker, J. L. and B. Parsons. 2010. New plant records from the Big Island
for 2008. Bishop Museum Occasional Papers 107: 41–43.
Parker, J. L. and B. Parsons. 2012a. New plant records from the Big Island
for 2009. Bishop Museum Occasional Papers 113: 55–63.
Parker, J. L. and B. Parsons. 2012b. New plant records from the Big Island
for 2010–2011. Bishop Museum Occasional Papers 113: 65–74.
Parris, B. S. 2007. Five new genera and three new species of
Grammitidaceae (Filicales) and the re-establishment of Oreogrammitis.
Gardens’ Bulletin Singapore 58(2): 233–274.
Peterson, P. M., K. Romaschenko, N. Snow and G. Johnson. 2012. A
molecular phylogeny and classification of Leptochloa (Poaceae:
Chlorideae) sensu lato and related genera. Annals of Botany, London
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Peterson, P. M., R. J. Soreng, G. Davidse, T. S. Filgueiras, F. O. Zuloaga
and E. J. Judziewicz. 2001. Catalogue of New World grasses
(Poaceae): II. Subfamily Chloridoideae. Contributions from the U.S.
National Herbarium 41: 1–255.
Pfeil, B. E., C. L. Brubaker, L. A. Craven and M. D. Crisp. 2002.
Phylogeny of Hibiscus and the tribe Hibisceae (Malvaceae) using
chloroplast DNA sequences of ndhF and the rpl16 intron. Systematic
Botany 27: 333–350.
Pratt, L. W. and K. F. Bio. 2012. New plant records from Hawai‘i island.
Bishop Museum Occasional Papers 113: 75–80.
Puttock, C. F. 1999. Revision of Atractocarpus (Rubiaceae: Gardenieae) in
Australia and new combinations for some extra-Australian taxa.
Australian Systematic Botany 12(2): 271–309.
Ranker, T. A. 2008. A new combination in Adenophorus (Polypodiaceae).
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and taxonomic notes in the tribe Morindeae (Rubiaceae). Adansonia
33: 283–309.
Resslar, P. M. 2010. Caladium bicolor naturalized on the island of Hawai‘i.
Bishop Museum Occasional Papers 107: 44–45.
Rico-Arce, M. L. 2007. A Checklist and Synopsis of American Species of
Acacia (Leguminosae: Mimosoideae), Conabio & Royal Botanic
Gardens, Kew, 207 pp.
Robinson, H., S. C. Keeley, J. J. Skvarla and R. Chan. 2008. Studies on the
Gymnantheminae (Vernonieae: Asteraceae) III. Restoration of the
genus Strobocalyx and the new genus Tarlmounia. Proceedings of the
Biological Society of Washington 121(1): 19–33.
Robson, N. K. B. 1990. Studies in the genus Hypericum L. (Guttiferae) 8.
Sections 29. Brathys (part 2) and 30. Trigynobrathys. Bulletin of the
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Ronse Decraene, L.-P. A., J.R. 1988. Generic limits in Polygonum and
related genera (Polygonaceae) on the basis of floral characters.
Botanical Journal of the Linnean Society 98: 321–371.
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Hawaiian Native & Naturalized Vascular Plant Checklist
Russell, C. and M. M. Bruegmann. 1996a. Endangered and threatened
wildlife and plants; determination of endangered or threatened status
for fourteen plant taxa from the Hawaiian Islands. Federal Register
61(198): 53108–53124.
Russell, C. and M. M. Bruegmann. 1996b. Endangered and threatened
wildlife and plants; determination of endangered status for twenty-five
plant species from the Island of Oahu, Hawaii. Federal Register
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Sato, P. 1998. A new naturalized plant species in windward O‘ahu,
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Saunders, J. G. 2011. Resurrection of the Maui endemic Waltheria
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Shannon, R. K. and W. L. Wagner. 1996. New records of Hawaiian
flowering plants primarily from the United States National Herbarium.
Bishop Museum Occasional Papers 46: 13–15.
Shannon, R. K., W. L. Wagner and D. H. Nicolson. 1997. (1322) Proposal
to conserve the name Scaevola taccada (Goodeniaceae) with a
conserved type. Taxon 46: 801–802.
Shaw, R. B., B. F. Close and L. Schnell. 1995. Rediscovery of Solanum
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Shaw, R. B. and P. P. Douglas. 1995. Carex inversa R. Br. (Cyperaceae),
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97–102.
Snow, N. 2008. Notes on grasses (Poaceae) in Hawai‘i. Bishop Museum
Occasional Papers 100: 38–43.
Snow, N. 2011. A new name in Asplenium (Aspleniaceae) from Hawai‘i.
Harvard Papers in Botany 16(1): 161.
Snow, N. and G. Davidse. 2011. Notes on grasses (Poaceae) in Hawai‘i: 3.
Bishop Museum Occasional Papers 110: 17–22.
Snow, N. and A. Lau. 2010. Notes on grasses (Poaceae) in Hawai‘i: 2.
Bishop Museum Occasional Papers 107: 46–60.
Snow, N., T. A. Ranker and D. H. Lorence. 2011. Taxonomic changes in
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Soreng, R. J. and E. A. Cope. 1991. On the taxonomy of cultivated species
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St. John, H. 1987a. Diagnoses of Phyllostegia species (Labiatae): Hawaiian
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St. John, H. 1987b. Diagnoses of Delissea species (Lobeliaceae) from
Kauai: Hawaiian plant studies 145. Phytologia 63(5): 339–349.
St. John, H. 1987c. Diagnoses of Panicum species (Gramineae): Hawaiian
plant studies 149. Phytologia 63(5): 368–395.
St. John, H. 1987d. Section Lobicalyces of Cyrtandra (Gesneriaceae):
Hawaiian plant studies 154. Phytologia 63(6): 476-484.
St. John, H. 1987e. Diagnoses of Cyrtandra species (Gesneriaceae) section
Microcalyces: Hawaiian plant studies 155. Phytologia 63(6): 485–493.
St. John, H. 1987f. Diagnoses of Cyrtandra species (Gesneriaceae) section
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St. John, H. 1987g. Diagnoses of Lysimachia species (Primulaceae).
Hawaiian plant studies 159. Phytologia 64(1): 43–50
St. John, H. 1987h. Diagnoses of new species of Sicyocarya
(Cucurbitaceae): Hawaiian plant studies 160. Phytologia 64(1): 51–54.
St. John, H. 1987i. Diagnosis of Pleomele stenophylla (Liliaceae):
Hawaiian plant studies 161. Phytologia 64(1): 55.
St. John, H. 1988a. Diagnoses of some Phanerogamae: Hawaiian plant
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St. John, H. 1988b. Diagnoses of new Cyrtandra species (Gesneriaceae):
Hawaiian plant studies 144. Phytologia 65(3): 195–206.
Staples, G. W. 2009. (1890) Proposal to conserve the name Bonamia
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651.
Staples, G. W. and D. R. Herbst. 2005. A Tropical Garden Flora: Plants
Cultivated in the Hawaiian Islands and Other Tropical Places. Bishop
Museum Press, Honolulu, 908 pp.
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Hawaiian Native & Naturalized Vascular Plant Checklist
Staples, G. W., D. R. Herbst and C. T. Imada. 2006. New Hawaiian plant
records for 2004. Bishop Museum Occasional Papers 88: 6–9.
Staples, G. W., C. T. Imada and D. R. Herbst. 2002. New Hawaiian plant
records for 2000. Bishop Museum Occasional Papers 68: 3–18.
Staples, G. W., C. T. Imada and D. R. Herbst. 2003. New Hawaiian plant
records for 2001. Bishop Museum Occasional Papers 74: 7–21.
Staples, G. W. and K. R. Woolliams. 1997. An overlooked naturalized aroid
for the Hawaiian flora. Bishop Museum Occasional Papers 49: 13–16.
Starr, F. and K. Martz. 2000. New plant records for Midway Atoll. Bishop
Museum Occasional Papers 64: 10–12.
Starr, F., K. Martz and L. L. Loope. 1999. New plant records from East
Maui for 1998. Bishop Museum Occasional Papers 59: 11–15.
Starr, F., K. Martz and L. L. Loope. 2002. New plant records for the
Hawaiian archipelago. Bishop Museum Occasional Papers 69: 16–27.
Starr, F. and K. Starr. 2011. New plant records from Midway Atoll, Maui,
and Kaho‘olawe. Bishop Museum Occasional Papers 110: 23–35.
Starr, F. and K. Starr. 2012. New plant records from Maui and Kaho‘olawe.
Bishop Museum Occasional Papers 113: 87–90.
Starr, F., K. Starr and L. L. Loope. 2003. New plant records from the
Hawaiian archipelago. Bishop Museum Occasional Papers 74: 23–34.
Starr, F., K. Starr and L. L. Loope. 2004. New plant records from the
Hawaiian archipelago. Bishop Museum Occasional Papers 79: 20–30.
Starr, F., K. Starr and L. L. Loope. 2006. New plant records from the
Hawaiian archipelago. Bishop Museum Occasional Papers 87: 31–43.
Starr, F., K. Starr and L. L. Loope. 2008. New plant records from the
Hawaiian archipelago. Bishop Museum Occasional Papers 100: 44–49.
Starr, F., K. Starr and L. L. Loope. 2010. New plant records from the
Hawaiian Archipelago. Bishop Museum Occasional Papers 107: 61–68.
Steinmann, V. W. and J. M. Porter. 2002. Phylogenetic relationships in
Euphorbieae (Euphorbiaceae) based on ITS and ndhF sequence data.
Annals of the Missouri Botanical Garden 89: 453–490.
Strong, M. T. 2004. Proposal to reject the name Scirpus miliaceus
(Cyperaceae). Taxon 53(4): 1069–1070.
Strong, M. T. and W. L. Wagner. 1997. New and noteworthy Cyperaceae
from the Hawaiian Islands. Bishop Museum Occasional Papers 48: 37–
50.
Sun, B. and S. M. Phillips. 2006. Ischaemum. Flora of China 22: 609–613.
Sundberg, S. D. 2004. New combinations in North American
Symphyotrichum subgenus Astropolium (Asteraceae: Astereae). Sida
21: 903–910.
Swenson, U., J. Munzinger and I. V. Bartish. 2007. Molecular phylogeny of
Planchonella (Sapotaceae) and eight new species from New Caledonia.
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Terrell, E. E. and W. H. Lewis. 1990. Oldenlandiopsis (Rubiaceae), a new
genus from the Caribbean Basin, based on Oldenlandia callitrichoides
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Terrell, E. E., H. E. Robinson, W. L. Wagner and D. H. Lorence. 2005.
Resurrection of genus Kadua for Hawaiian Hedyotidinae (Rubiaceae),
with emphasis on seed and fruit characters and notes on South Pacific
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Thompson, J. 1989. A revision of the genus Leptospermum (Myrtaceae).
Telopea 3(3): 291–449.
Thomson, J. A. 2004. Towards a taxonomic revision of Pteridium
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Tripp, E. A., T. F. Daniel, J. C. Lendemer and L. A. McDade. 2009. New
molecular and morphological insights prompt transfer of Blechum to
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Tucker, G. C. 1994. Revision of the Mexican species of Cyperus
(Cyperaceae). Systematic Botany Monographs 43: 1–213.
Wagner, W. L. 1996. Scaevola hobdyi (Goodeniaceae), an enigmatic new
species from West Maui, Hawaiian Islands. Novon 6(2): 225–227.
Wagner, W. L. 1999a. Nomenclator and review of Phyllostegia
(Lamiaceae). Novon 9(2): 265–279.
Wagner, W. L. 1999b. A new species of Hawaiian Phyllostegia
(Lamiaceae) from Kaua‘i and recognition of a Wai‘anae Mountain,
O‘ahu, endangered variety of Phyllostegia parviflora. Novon 9(2):
280–283.
Wagner, W. L. and E. M. Harris. 2000. A unique Hawaiian Schiedea
(Caryophyllaceae: Alsinoideae) with only five fertile stamens.
American Journal of Botany 87(2): 153–160.
Wagner, W. L. and D. R. Herbst. 1995. Contributions to the flora of
Hawai‘i. IV. New records and name changes. Bishop Museum
Occasional Papers 42: 13–27.
Wagner, W. L. and D. R. Herbst. 1999. Supplement to the Manual of the
Flowering Plants of Hawai‘i, pp. 1885–1918 in Wagner, W. L., D. R.
Herbst and S. H. Sohmer, Manual of the Flowering Plants of Hawai‘i,
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Wagner, W. L. and D. R. Herbst. 2003. Supplement to the Manual of the
Flowering Plants of Hawai‘i, version 3.1 (12 Dec 2003).
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Hawaiian Native & Naturalized Vascular Plant Checklist
http://botany.si.edu/pacificislandbiodiversity/hawaiianflora/ManualSup
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Wagner, W. L., D. R. Herbst, N. Khan and T. Flynn. 2012. Hawaiian
vascular plant updates: a supplement to the Manual of the Flowering
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Wagner, W. L., D. R. Herbst, and D. H. Lorence. 2005-. Flora of the
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Wagner, W. L., D. R. Herbst and S. H. Sohmer. 1990. Manual of the
Flowering Plants of Hawai‘i. University of Hawaii Press and Bishop
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Wagner, W. L. and D. H. Lorence. 1998. A new, dioecious species of
Hedyotis (Rubiaceae) from Kaua‘i, Hawaiian Islands, and the
taxonomy of Kaua‘i Hedyotis schlechtendahliana resolved. Novon
8(3): 311–317.
Wagner, W. L. and D. H. Lorence. 2000. A reassessment of Cyrtandra
kealiae and C. limahuliensis (Gesneriaceae). Bishop Museum
Occasional Papers 63: 17–20.
Wagner, W. L. and H. Robinson. 2001[2002]. Lipochaeta and Melanthera
(Asteraceae: Heliantheae subtribe Ecliptinae): establishing their natural
limits and a synopsis. Brittonia 53(4): 539–561.
Wagner, W. L. and R. K. Shannon. 1998. Notes on Hawaiian Melicope
(Rutaceae). Bishop Museum Occasional Papers 56: 15–17.
Wagner, W. L. and R. K. Shannon. 1999. Nomenclator of Hawaiian Sicyos
(Cucurbitaceae). Novon 9(3): 441–447.
Wagner, W. L., R. K. Shannon and D. R. Herbst. 1997. Contributions to the
flora of Hawai‘i. VI. Bishop Museum Occasional Papers 48: 51–65.
Wagner, W. L. and S. G. Weller. 1991. Resurrection of a Kaua‘i Stenogyne:
S. kealiae. Pacific Science 45(1): 50–54.
Wagner, W. L. and S. G. Weller. 1999. Rediscovery and assessment of
Stenogyne sherffii Degener (Lamiaceae). Novon 9(3): 448–449.
Wagner, W. L., S. G. Weller and A. K. Sakai. 1994. Description of a rare
new cliff-dwelling species from Kaua‘i, Schiedea attenuata
(Caryophyllaceae). Novon 4(2): 187–190.
Wagner, W. L., S. G. Weller and A. K. Sakai. 2005. Monograph of
Schiedea (Caryophyllaceae-Alsinoideae). Systematic Botany
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Wagner, W. L., S. G. Weller, A. K. Sakai and A. C. Medeiros. 1999. An
autogamous rainforest species of Schiedea (Caryophyllaceae) from
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Wagner, W. L., K. R. Wood and D. H. Lorence. 2001. A new species of
Cyrtandra (Gesneriaceae) from Kaua‘i, Hawaiian Islands. Novon
11(1): 146–152.
Warren, S. D. 1993. Sporobolus indicus R. Br.—new record for
Kahoolawe. Newsletter of the Hawaiian Botanical Society 31(3/4): 43.
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Newsletter of the Hawaiian Botanical Society 33(1): 1, 3.
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Pacific Basin. Brittonia 45(4): 275–285.
Wilson, K. A. 2003. New records of alien pteridophytes from Hawai‘i.
Bishop Museum Occasional Papers 74: 5–7.
Wilson, K. L. 1990. Some widespread species of Persicaria (Polygonaceae)
and their allies. Kew Bulletin 45: 621–636.
Wood, K. R. 2006. New plant records and rediscoveries within the
Hawaiian Islands. Bishop Museum Occasional Papers 88: 15–19.
Wood, K. R. 2007. New plant records, rediscoveries, range extensions, and
possible extinctions within the Hawaiian Islands. Bishop Museum
Occasional Papers 96: 13–17.
Wood, K. R. 2012. Possible extinctions, rediscoveries, and new plant
records within the Hawaiian Islands. Bishop Museum Occasional
Papers 113: 91–102.
Wood, K. R. and M. Kiehn. 2011. Pittosporum halophilum Rock
(Pittosporaceae: Apiales): rediscovery, taxonomic assessment, and
conservation status of a critically endangered endemic species from
Moloka‘i, Hawaiian Islands. Pacific Science 65(4): 465–476.
Wood, K. R. and M. LeGrande. 2006. An annotated checklist and new
island records of flowering plants from Lehua Islet, Ni‘ihau, Hawai‘i.
Bishop Museum Occasional Papers 87: 19–29.
Wood, K. R. and H. L. Oppenheimer. 2008. Stenogyne kauaulaensis
(Lamiaceae), a new species from Maui, Hawaiian Islands. Novon
18(4): 544–549.
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Hawaiian Native & Naturalized Vascular Plant Checklist
Wood, K. R., W. L. Wagner and T. J. Motley. 2007. Labordia lorenciana
(Loganiaceae): a new critically endangered species from Kaua‘i,
Hawaiian Islands with comments on its conservation. Systematic
Botany 32(1): 195–199.
Wysong, M., G. Hughes and K. R. Wood. 2007. New Hawaiian plant
records for the island of Moloka‘i. Bishop Museum Occasional Papers
96: 1–8.
Yang, Y. and P. E. Berry. 2011. Phylogenetics of the Chamaesyce clade
(Euphorbia, Euphorbiaceae): reticulate evolution and long-distance
dispersal in a prominent c4 lineage. American Journal of Botany 98:
1486–1503.
Yesilyurt, J. C. 2005. The fern genus Doryopteris (CheilanthoideaePteridaceae) in the Hawaiian Islands. Kew Bulletin 60: 547–558.
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Hawaiian Native & Naturalized Vascular Plant Checklist
Index of accepted families and genera
This index is a finding tool for locating the initial mention of family and
accepted generic names in Appendices A and B, where complete listings of
accepted native and naturalized vascular plant taxa are presented for the
main Hawaiian Islands (Appendix A) and the Northwestern Hawaiian
Islands (Appendix B). Generic names are followed by their family
Abelmoschus (Malvaceae) A-110
Abrus (Fabaceae) A-78
Abutilon (Malvaceae) A-110, B-9
Acacia (Fabaceae) A-78
Acaena (Rosaceae) A-143
Acanthaceae A-1, B-1
Acanthocereus (Cactaceae) A-40
Acanthospermum (Asteraceae) A-14
Achillea (Asteraceae) A-14
Achyranthes (Amaranthaceae) A-4, B-1
Adenanthera (Fabaceae) A-79
Adenophorus (Grammitidaceae) A-239
Adenostemma (Asteraceae) A-14
Adiantum (Pteridaceae) A-246
Aechmea (Bromeliaceae) A-184
Aeschynomene (Fabaceae) A-79
Afrocarpus (Podocarpaceae) A-177
Agavaceae A-178
Agave (Agavaceae) A-178
Ageratina (Asteraceae) A-14
Ageratum (Asteraceae) A-14
Aglaonema (Araceae) A-180
Agrostis (Poaceae) A-202
Ailanthus (Simaroubaceae) A-163
Aira (Poaceae) A-202
Aizoaceae A-3, B-1
Albizia (Fabaceae) A-79, B-7
Alectryon (Sapindaceae) A-160
Aleurites (Euphorbiaceae) A-70
designation. If you cannot find a generic name here that you are familiar
with from the Manual of the Flowering Plants of Hawai‘i (Wagner et al.
1990), refer to Appendix D, where the alphabetical list of synonyms can
clarify the present disposition of the name in question.
Alismataceae A-179
Allamanda (Apocynaceae) A-8
Allophyllus (Sapindaceae) A-160
Alnus (Betulaceae) A-35
Alocasia (Araceae) A-180
Aloe (Aloeaceae) A-179
Aloeaceae A-179
Alphitonia (Rhamnaceae) A-142
Alpinia (Zingiberaceae) A-230
Alstonia (Apocynaceae) A-8
Alternanthera (Amaranthaceae) A-4
Alysicarpus (Fabaceae) A-80
Alyxia (Apocynaceae) A-8
Amaranthaceae A-4, B-1
Amaranthus (Amaranthaceae) A-4, B-1
Ambrosia (Asteraceae) A-14
Ammannia (Lythraceae) A-109
Ammophila (Poaceae) A-202, B-13
Amsinckia (Boraginaceae) A-36
Anacardiaceae A-6, B-1
Anacardium (Anacardiaceae) A-6
Anagallis (Primulaceae) A-140, B-11
Andropogon (Poaceae) A-202, B-13
Anemone (Ranunculaceae) A-142
Anethum (Apiaceae) A-7, B-2
Angiopteris (Marattiaceae) A-243
Angophora (Myrtaceae) A-121
Annona (Annonaceae) A-6
Annonaceae A-6
18
Anoectochilus (Orchidaceae) A-200
Anredera (Basellaceae) A-34
Anthemis (Asteraceae) A-14
Anthephora (Poaceae) A-203
Anthoxanthum (Poaceae) A-203
Anthurium (Araceae) A-180
Antiaris (Moraceae) A-117
Antidesma (Euphorbiaceae) A-70
Antigonon (Polygonaceae) A-137, B-10
Antirrhinum (Scrophulariaceae) A-161
Aphelandra (Acanthaceae) A-1
Apiaceae A-7, B-2
Apocynaceae A-8, B-2
Aptenia (Aizoaceae) A-3
Aquifoliaceae A-10
Araceae A-180
Arachniodes (Dryopteridaceae) A-236
Araliaceae A-10
Araucaria (Araucariaceae) A-176
Araucariaceae A-176
Archontophoenix (Arecaceae) A-181
Arctium (Asteraceae) A-14
Arctotheca (Asteraceae) A-15
Ardisia (Myrsinaceae) A-119
Arecaceae A-181, B-12
Arenaria (Caryophyllaceae) A-55
Arenga (Arecaceae) A-181
Argemone (Papaveraceae) A-131
Argyreia (Convolvulaceae) A-63
Hawaiian Native & Naturalized Vascular Plant Checklist
Argyroxiphium (Asteraceae) A-15
Aristea (Iridaceae) A-196
Aristida (Poaceae) A-203
Aristolochia (Aristolochiaceae) A-13
Aristolochiaceae A-13
Arrhenatherum (Poaceae) A-203
Artabotrys (Annonaceae) A-7
Artemisia (Asteraceae) A-15
Arthraxon (Poaceae) A-203
Arthrostemma (Melastomataceae) A-115
Artocarpus (Moraceae) A-117
Arundina (Orchidaceae) A-200
Arundo (Poaceae) A-203
Asclepiadaceae A-13
Asclepias (Asclepiadaceae) A-13
Asparagus (Liliaceae) A-198
Aspleniaceae A-230
Asplenium (Aspleniaceae) A-230
Astelia (Liliaceae) A-198
Asteraceae A-14, B-2
Asystasia (Acanthaceae) A-1, B-1
Athyriaceae A-233
Athyrium (Athyriaceae) A-233
Atractocarpus (Rubiaceae) A-146
Atriplex (Chenopodiaceae) A-60, B-5
Avena (Poaceae) A-203
Axonopus (Poaceae) A-203
Azadirachta (Meliaceae) A-116
Azolla (Azollaceae) A-234
Azollaceae A-234
Bacopa (Scrophulariaceae) A-161, B-11
Balsaminaceae A-34
Baltimora (Asteraceae) A-15
Banksia (Proteaceae) A-141
Barleria (Acanthaceae) A-1
Basella (Basellaceae) A-34, B-3
Basellaceae A-34, B-3
Bassia (Chenopodiaceae) A-61
Bastardia (Malvaceae) A-110
Bataceae A-34
Batis (Bataceae) A-34
Bauhinia (Fabaceae) A-80
Beaumontia (Apocynaceae) A-9
Begonia (Begoniaceae) A-34
Begoniaceae A-34
Bellis (Asteraceae) A-15
Betulaceae A-35
Bidens (Asteraceae) A-16, B-2
Bignoniaceae A-35
Bischofia (Euphorbiaceae) A-71
Bixa (Bixaceae) A-36
Bixaceae A-36
Blechnaceae A-234
Blechnum (Blechnaceae) A-234
Blumea (Asteraceae) A-18
Blutaparon (Amaranthaceae) A-5
Bobea (Rubiaceae) A-146
Bocconia (Papaveraceae) A-131
Boehmeria (Urticaceae) A-169
Boerhavia (Nyctaginaceae) A-127, B-9
Bolboschoenus (Cyperaceae) A-187
Bonamia (Convolvulaceae) A-63
Boraginaceae A-36, B-3
Bothriochloa (Poaceae) A-204, B-13
Bothriospermum (Boraginaceae) A-36
Botrychiaceae A-235
Botrychium (Botrychiaceae) A-235
Bouteloua (Poaceae) A-204
Brassica (Brassicaceae) A-38, B-4
Brassicaceae A-38, B-4
Brexia (Grossulariaceae) A-100
Breynia (Euphorbiaceae) A-71
Brighamia (Campanulaceae) A-41
Briza (Poaceae) A-204
19
Bromeliaceae A-184
Bromus (Poaceae) A-204, B-14
Broussaisia (Hydrangeaceae) A-100
Broussonetia (Moraceae) A-118
Brugmansia (Solanaceae) A-163
Bruguiera (Rhizophoraceae) A-143
Buchnera (Scrophulariaceae) A-161
Buddleja (Buddlejaceae) A-40
Buddlejaceae A-40
Bulbostylis (Cyperaceae) A-187
Cactaceae A-40
Caesalpinia (Fabaceae) A-80, B-7
Cajanus (Fabaceae) A-80
Caladium (Araceae) A-180
Calamagrostis (Poaceae) A-205
Calathea (Marantaceae) A-199
Calliandra (Fabaceae) A-80
Callisia (Commelinaceae) A-186
Callistopteris (Hymenophyllaceae) A-240
Callitris (Cupressaceae) A-176
Caloncoba (Flacourtiaceae) A-93
Calophyllum (Clusiaceae) A-62
Calopogonium (Fabaceae) A-81
Calotropis (Asclepiadaceae) A-13
Calyptocarpus (Asteraceae) A-18, B-2
Camellia (Theaceae) A-167
Campanulaceae A-41
Canavalia (Fabaceae) A-81
Canna (Cannaceae) A-186
Cannabaceae A-55
Cannabis (Cannabaceae) A-55
Cannaceae A-186
Capparaceae A-55, B-4
Capparis (Capparaceae) A-55, B-4
Caprifoliaceae A-55
Capsella (Brassicaceae) A-38, B-4
Capsicum (Solanaceae) A-163
Hawaiian Native & Naturalized Vascular Plant Checklist
Cardamine (Brassicaceae) A-38
Cardiospermum (Sapindaceae) A-160
Carduus (Asteraceae) A-18
Carex (Cyperaceae) A-187
Carica (Caricaceae) A-55
Caricaceae A-55
Carissa (Apocynaceae) A-9
Carmona (Boraginaceae) A-37
Caryophyllaceae A-55, B-4
Caryota (Arecaceae) A-181
Cassia (Fabaceae) A-81
Cassytha (Lauraceae) A-107, B-9
Castilleja (Scrophulariaceae) A-161
Casuarina (Casuarinaceae) A-60, B-5
Casuarinaceae A-60, B-5
Catalpa (Bignoniaceae) A-35
Catha (Celastraceae) A-60
Catharanthus (Apocynaceae) A-9, B-2
Cayratia (Vitaceae) A-175
Cecropia (Cecropiaceae) A-60
Cecropiaceae A-60
Cedrela (Meliaceae) A-117
Celastraceae A-60
Celosia (Amaranthaceae) A-5
Cenchrus (Poaceae) A-205, B-14
Centaurea (Asteraceae) A-18
Centaurium (Gentianaceae) A-94, B-8
Centella (Apiaceae) A-7
Centranthus (Valerianaceae) A-170
Centratherum (Asteraceae) A-18
Centrosema (Fabaceae) A-81
Cerastium (Caryophyllaceae) A-55
Ceratophyllaceae A-60
Ceratophyllum (Ceratophyllaceae) A-60
Ceratopteris (Pteridaceae) A-246
Cereus (Cactaceae) A-40
Cestrum (Solanaceae) A-164
Chamaecrista (Fabaceae) A-82
Charpentiera (Amaranthaceae) A-5
Cheilanthes (Pteridaceae) A-246
Cheilocostus (Costaceae) A-187
Cheirodendron (Araliaceae) A-10
Chenopodiaceae A-60, B-5
Chenopodium (Chenopodiaceae) A-61, B-5
Chloris (Poaceae) A-207, B-14
Chromolaena (Asteraceae) A-18
Chrysobalanaceae A-62
Chrysobalanus (Chrysobalanaceae) A-62
Chrysophyllum (Sapotaceae) A-161
Chrysopogon (Poaceae) A-208
Cibotium (Dicksoniaceae) A-236
Ciclospermum (Apiaceae) A-7, B-2
Cinchona (Rubiaceae) A-146
Cinnamomum (Lauraceae) A-107
Cirsium (Asteraceae) A-18
Cissus (Vitaceae) A-175
Citharexylum (Verbenaceae) A-171
Cladium (Cyperaceae) A-188
Claoxylon (Euphorbiaceae) A-71
Cleome (Capparaceae) A-55
Clermontia (Campanulaceae) A-42
Clerodendrum (Verbenaceae) A-171
Clethra (Clethraceae) A-62
Clethraceae A-62
Clidemia (Melastomataceae) A-115
Clitoria (Fabaceae) A-82
Clusia (Clusiaceae) A-62
Clusiaceae A-62
Coccinia (Cucurbitaceae) A-67, B-6
Coccoloba (Polygonaceae) A-137, B-10
Coccothrinax (Arecaceae) A-181
Cocculus (Menispermaceae) A-117
Cocos (Arecaceae) A-181, B-12
Coffea (Rubiaceae) A-146
20
Coix (Poaceae) A-208
Collomia (Polemoniaceae) A-137
Colocasia (Araceae) A-180
Colubrina (Rhamnaceae) A-142
Combretaceae A-63
Commelina (Commelinaceae) A-186, B-12
Commelinaceae A-186, B-12
Coniogramme (Pteridaceae) A-246
Conocarpus (Combretaceae) A-63
Convolvulaceae A-63, B-5
Convolvulus (Convolvulaceae) A-63
Conyza (Asteraceae) A-18, B-2
Coprosma (Rubiaceae) A-146
Corchorus (Tiliaceae) A-168
Cordia (Boraginaceae) A-37
Cordyline (Agavaceae) A-178
Coreopsis (Asteraceae) A-19
Coriandrum (Apiaceae) A-7
Cornaceae A-66
Cornus (Cornaceae) A-66
Coronilla (Fabaceae) A-82
Coronopus (Brassicaceae) A-38, B-4
Cortaderia (Poaceae) A-208
Corymbia (Myrtaceae) A-121
Corynocarpaceae A-66
Corynocarpus (Corynocarpaceae) A-66
Cosmos (Asteraceae) A-19
Costaceae A-187
Costus (Costaceae) A-187
Cotoneaster (Rosaceae) A-143
Cotula (Asteraceae) A-19
Crassocephalum (Asteraceae) A-19
Crassula (Crassulaceae) A-66
Crassulaceae A-66
Crepidomanes (Hymenophyllaceae) A-241
Crepis (Asteraceae) A-19
Cressa (Convolvulaceae) A-63
Hawaiian Native & Naturalized Vascular Plant Checklist
Crocosmia (Iridaceae) A-196
Crotalaria (Fabaceae) A-82, B-7
Croton (Euphorbiaceae) A-71
Cryptocarya (Lauraceae) A-107
Cryptomeria (Taxodiaceae) A-178
Cryptostegia (Asclepiadaceae) A-13
Cryptotaenia (Apiaceae) A-7
Ctenitis (Dryopteridaceae) A-236
Cucumis (Cucurbitaceae) A-67
Cucurbitaceae A-67, B-6
Cupaniopsis (Sapindaceae) A-160
Cuphea (Lythraceae) A-109
Cupressaceae A-176
Cupressus (Cupressaceae) A-176
Curcuma (Zingiberaceae) A-230
Cuscuta (Cuscutaceae) A-69
Cuscutaceae A-69
Cyanea (Campanulaceae) A-44
Cyanthillium (Asteraceae) A-19
Cyatheaceae A-235
Cyclosorus (Thelypteridaceae) A-248
Cymbalaria (Scrophulariaceae) A-162
Cymbidium (Orchidaceae) A-200
Cymbopogon (Poaceae) A-208
Cynanchum (Asclepiadaceae) A-13
Cynodon (Poaceae) A-208, B-14
Cynoglossum (Boraginaceae) A-37
Cyperaceae A-187, B-12
Cyperus (Cyperaceae) A-188, B-12
Cyrtandra (Gesneriaceae) A-95
Cyrtomium (Dryopteridaceae) A-236
Cystopteris (Athyriaceae) A-233
Cytisus (Fabaceae) A-83
Dacrycarpus (Podocarpaceae) A-177
Dactylis (Poaceae) A-208
Dactyloctenium (Poaceae) A-209, B-14
Dalea (Fabaceae) A-83
Datura (Solanaceae) A-164
Daucus (Apiaceae) A-7
Davallia (Davalliaceae) A-235
Davalliaceae A-235
Delairea (Asteraceae) A-19
Delissea (Campanulaceae) A-50
Delonix (Fabaceae) A-83
Dendrobium (Orchidaceae) A-200
Dennstaedtiaceae A-235
Deparia (Athyriaceae) A-233
Derris (Fabaceae) A-83
Deschampsia (Poaceae) A-209
Descurainia (Brassicaceae) A-38
Desmanthus (Fabaceae) A-83, B-7
Desmodium (Fabaceae) A-83, B-7
Dianella (Liliaceae) A-199
Dianthus (Caryophyllaceae) A-55
Dichanthelium (Poaceae) A-209
Dichanthium (Poaceae) A-209
Dichelachne (Poaceae) A-210
Dichondra (Convolvulaceae) A-63
Dicksonia (Dicksoniaceae) A-236
Dicksoniaceae A-236
Dicliptera (Acanthaceae) A-1
Dicranopteris (Gleicheniaceae) A-239
Dieffenbachia (Araceae) A-180
Dietes (Iridaceae) A-196
Digitaria (Poaceae) A-210, B-14
Dillenia (Dilleniaceae) A-69
Dilleniaceae A-69
Dimerocostus (Costaceae) A-187
Dioclea (Fabaceae) A-84
Diodia (Rubiaceae) A-147
Dioscorea (Dioscoreaceae) A-195
Dioscoreaceae A-195
Diospyros (Ebenaceae) A-69
Diplachne (Poaceae) A-211, B-14
21
Diplazium (Athyriaceae) A-234
Diplopterygium (Gleicheniaceae) A-239
Dipsacaceae A-69
Disakisperma (Poaceae) A-211
Dissochondrus (Poaceae) A-211
Dissotis (Melastomataceae) A-115
Distichlis (Poaceae) A-211
Dodonaea (Sapindaceae) A-160
Doodia (Blechnaceae) A-234
Dopatrium (Scrophulariaceae) A-162
Doryopteris (Pteridaceae) A-246
Dovyalis (Flacourtiaceae) A-93
Drosera (Droseraceae) A-69
Droseraceae A-69
Drymaria (Caryophyllaceae) A-56
Dryopteridaceae A-236
Dryopteris (Dryopteridaceae) A-236
Dubautia (Asteraceae) A-19
Duranta (Verbenaceae) A-171
Dysphania (Chenopodiaceae) A-61
Dyssodia (Asteraceae) A-22
Ebenaceae A-69
Echinochloa (Poaceae) A-211, B-14
Echinodorus (Alismataceae) A-179
Eclipta (Asteraceae) A-22
Egeria (Hydrocharitaceae) A-195
Ehrharta (Poaceae) A-212
Eichhornia (Pontederiaceae) A-228
Elaeagnaceae A-70
Elaeagnus (Elaeagnaceae) A-70
Elaeocarpaceae A-70
Elaeocarpus (Elaeocarpaceae) A-70
Elaphoglossaceae A-238
Elaphoglossum (Elaphoglossaceae) A-238
Eleocharis (Cyperaceae) A-192
Elephantopus (Asteraceae) A-22
Eleusine (Poaceae) A-212, B-15
Hawaiian Native & Naturalized Vascular Plant Checklist
Embelia (Myrsinaceae) A-120
Emex (Polygonaceae) A-137
Emilia (Asteraceae) A-22, B-2
Encelia (Asteraceae) A-22
Enchylaena (Chenopodiaceae) A-61
Enneapogon (Poaceae) A-212
Entada (Fabaceae) A-84
Enterolobium (Fabaceae) A-84
Entolasia (Poaceae) A-212
Epacridaceae A-70
Epidendrum (Orchidaceae) A-201
Epilobium (Onagraceae) A-129
Epipremnum (Araceae) A-180
Eragrostis (Poaceae) A-212, B-15
Erechtites (Asteraceae) A-22
Erica (Ericaceae) A-70
Ericaceae A-70
Erigeron (Asteraceae) A-23
Eriobotrya (Rosaceae) A-144
Eriocaulaceae A-195
Eriocaulon (Eriocaulaceae) A-195
Eriochloa (Poaceae) A-214, B-15
Erodium (Geraniaceae) A-94
Eryngium (Apiaceae) A-7
Erythrina (Fabaceae) A-84
Escallonia (Grossulariaceae) A-100
Eschscholzia (Papaveraceae) A-131
Etlingera (Zingiberaceae) A-230
Eucalyptus (Myrtaceae) A-121
Euchiton (Asteraceae) A-23
Eugenia (Myrtaceae) A-123
Eupatorium (Asteraceae) A-23
Euphorbia (Euphorbiaceae) A-71, B-6
Euphorbiaceae A-70, B-6
Eurya (Theaceae) A-167
Eustachys (Poaceae) A-215, B-15
Evolvulus (Convolvulaceae) A-63
Exocarpos (Santalaceae) A-159
Fabaceae A-78, B-7
Fagaceae A-93
Falcataria (Fabaceae) A-85
Fallopia (Polygonaceae) A-138
Fatoua (Moraceae) A-118
Festuca (Poaceae) A-215, B-15
Ficus (Moraceae) A-118, B-9
Filicium (Sapindaceae) A-160
Fimbristylis (Cyperaceae) A-192, B-13
Fitchia (Asteraceae) A-23
Flacourtia (Flacourtiaceae) A-93
Flacourtiaceae A-93
Flaveria (Asteraceae) A-23, B-2
Flemingia (Fabaceae) A-85
Flindersia (Rutaceae) A-153
Florestina (Asteraceae) A-23
Flueggea (Euphorbiaceae) A-76
Foeniculum (Apiaceae) A-7
Fragaria (Rosaceae) A-144
Frankenia (Frankeniaceae) A-93, B-8
Frankeniaceae A-93, B-8
Fraxinus (Oleaceae) A-128
Freycinetia (Pandanaceae) A-201
Fuchsia (Onagraceae) A-130
Fumaria (Fumariaceae) A-94
Fumariaceae A-94
Furcraea (Agavaceae) A-178
Gahnia (Cyperaceae) A-193
Gaillardia (Asteraceae) A-23, B-2
Galinsoga (Asteraceae) A-23
Galium (Rubiaceae) A-147
Gamochaeta (Asteraceae) A-23
Gardenia (Rubiaceae) A-147
Garnotia (Poaceae) A-215
Gastridium (Poaceae) A-215
Gentianaceae A-94, B-8
22
Geraniaceae A-94
Geranium (Geraniaceae) A-94
Gesneriaceae A-95
Gibasis (Commelinaceae) A-186
Gilia (Polemoniaceae) A-137
Gladiolus (Iridaceae) A-196
Gleicheniaceae A-239
Gliricidia (Fabaceae) A-85
Glyceria (Poaceae) A-215
Glycine (Fabaceae) A-85
Gomphrena (Amaranthaceae) A-5
Gonocarpus (Haloragaceae) A-100
Goodeniaceae A-99, B-8
Gossypium (Malvaceae) A-111
Gouania (Rhamnaceae) A-143
Grammitidaceae A-239
Grevillea (Proteaceae) A-141
Grewia (Tiliaceae) A-168
Grossulariaceae A-100
Guaiacum (Zygophyllaceae) A-176
Gunnera (Gunneraceae) A-100
Gunneraceae A-100
Guzmania (Bromeliaceae) A-184
Gynochthodes (Rubiaceae) A-147
Habenaria (Orchidaceae) A-201
Hackelochloa (Poaceae) A-215
Haematoxylum (Fabaceae) A-85
Halophila (Hydrocharitaceae) A-196, B-13
Haloragaceae A-100
Haplopteris (Vittariaceae) A-249
Haplostachys (Lamiaceae) A-100
Harrisia (Cactaceae) A-40
Hebe (Scrophulariaceae) A-162
Hedera (Araliaceae) A-11
Hedychium (Zingiberaceae) A-230
Helianthus (Asteraceae) A-24
Helichrysum (Asteraceae) A-24
Hawaiian Native & Naturalized Vascular Plant Checklist
Heliconia (Heliconiaceae) A-195
Heliconiaceae A-195
Heliocarpus (Tiliaceae) A-168
Heliotropium (Boraginaceae) A-37, B-3
Hemarthria (Poaceae) A-216
Hemigraphis (Acanthaceae) A-1
Herissantia (Malvaceae) A-111
Heritiera (Sterculiaceae) A-167
Hesperocnide (Urticaceae) A-169
Hesperomannia (Asteraceae) A-24
Heterocentron (Melastomataceae) A-115
Heteromeles (Rosaceae) A-144
Heteropogon (Poaceae) A-216
Heterotheca (Asteraceae) A-24
Hibiscadelphus (Malvaceae) A-111
Hibiscus (Malvaceae) A-111, B-9
Hillebrandia (Begoniaceae) A-35
Hippeastrum (Liliaceae) A-199
Hippobroma (Campanulaceae) A-52
Hiptage (Malpighiaceae) A-110
Hirschfeldia (Brassicaceae) A-38
Holcus (Poaceae) A-216
Homalanthus (Euphorbiaceae) A-77
Hordeum (Poaceae) A-216, B-15
Hoya (Asclepiadaceae) A-13
Hunnemannia (Papaveraceae) A-131
Huperzia (Lycopodiaceae) A-242
Hydrangeaceae A-100
Hydrocharitaceae A-195, B-13
Hydrocotyle (Apiaceae) A-7
Hydrophyllaceae A-100, B-8
Hylocereus (Cactaceae) A-41
Hymenophyllaceae A-240
Hymenophyllum (Hymenophyllaceae) A-241
Hyparrhenia (Poaceae) A-216
Hypericum (Clusiaceae) A-62
Hypochoeris (Asteraceae) A-24
Hypoestes (Acanthaceae) A-1
Hypolepis (Dennstaedtiaceae) A-235
Hypolytrum (Cyperaceae) A-193
Hyptis (Lamiaceae) A-101
Ilex (Aquifoliaceae) A-10
Impatiens (Balsaminaceae) A-34
Indigofera (Fabaceae) A-85, B-7
Inga (Fabaceae) A-85
Ipomoea (Convolvulaceae) A-63, B-5
Iridaceae A-196
Isachne (Poaceae) A-216
Ischaemum (Poaceae) A-216
Isodendrion (Violaceae) A-173
Isoetaceae A-241
Isoëtes (Isoetaceae) A-241
Ixophorus (Poaceae) A-217
Jacaranda (Bignoniaceae) A-35
Jacquemontia (Convolvulaceae) A-65
Jasminum (Oleaceae) A-129
Jatropha (Euphorbiaceae) A-77
Joinvillea (Joinvilleaceae) A-197
Joinvilleaceae A-197
Juncaceae A-197
Juncus (Juncaceae) A-197
Juniperus (Cupressaceae) A-177
Justicia (Acanthaceae) A-1
Kadua (Rubiaceae) A-148
Kalanchoe (Crassulaceae) A-66
Kalimeris (Asteraceae) A-25
Kanaloa (Fabaceae) A-85
Keysseria (Asteraceae) A-25
Khaya (Meliaceae) A-117
Koeleria (Poaceae) A-217
Koelreuteria (Sapindaceae) A-160
Kokia (Malvaceae) A-113
Korthalsella (Viscaceae) A-174
Kummerowia (Fabaceae) A-86
23
Kyllinga (Cyperaceae) A-193
Lablab (Fabaceae) A-86
Labordia (Loganiaceae) A-108
Lachnagrostis (Poaceae) A-217
Lactuca (Asteraceae) A-25
Lagenaria (Cucurbitaceae) A-68
Lamarckia (Poaceae) A-217
Lamiaceae A-100, B-8
Lamium (Lamiaceae) A-101
Landoltia (Lemnaceae) A-198
Lantana (Verbenaceae) A-172, B-12
Lapsana (Asteraceae) A-25
Lathyrus (Fabaceae) A-86
Lauraceae A-107, B-9
Leea (Leeaceae) A-107
Leeaceae A-107
Lemna (Lemnaceae) A-198
Lemnaceae A-198
Lentibulariaceae A-107
Leonotis (Lamiaceae) A-101
Leonurus (Lamiaceae) A-101
Lepechinia (Lamiaceae) A-101
Lepidium (Brassicaceae) A-38, B-4
Lepisorus (Polypodiaceae) A-244
Leptecophylla (Epacridaceae) A-70
Leptochloa (Poaceae) A-217
Leptospermum (Myrtaceae) A-124
Lepturus (Poaceae) A-217, B-15
Lespedeza (Fabaceae) A-86
Leucaena (Fabaceae) A-86, B-7
Leucanthemum (Asteraceae) A-25
Leymus (Poaceae) A-217
Licuala (Arecaceae) A-182
Ligustrum (Oleaceae) A-129
Liliaceae A-198
Linaceae A-107
Linaria (Scrophulariaceae) A-162
Hawaiian Native & Naturalized Vascular Plant Checklist
Lindernia (Scrophulariaceae) A-162
Lindsaea (Lindsaeaceae) A-241
Lindsaeaceae A-241
xLindsaeosoria (Lindsaeaceae) A-242
Linum (Linaceae) A-107
Liparis (Orchidaceae) A-201
Lipochaeta (Asteraceae) A-26
Liriodendron (Magnoliaceae) A-110
Livistona (Arecaceae) A-182
Lobelia (Campanulaceae) A-52
Lobularia (Brassicaceae) A-39, B-4
Loganiaceae A-108
Logfia (Asteraceae) A-26
Lolium (Poaceae) A-218
Lonicera (Caprifoliaceae) A-55
Lophospermum (Scrophulariaceae) A-162
Lophostemon (Myrtaceae) A-124
Lotus (Fabaceae) A-86
Ludwigia (Onagraceae) A-130
Lupinus (Fabaceae) A-86
Luzula (Juncaceae) A-197
Lycium (Solanaceae) A-164
Lycopodiaceae A-242
Lycopodiella (Lycopodiaceae) A-243
Lycopodium (Lycopodiaceae) A-243
Lygodium (Schizaeaceae) A-248
Lysimachia (Primulaceae) A-140
Lythraceae A-109
Lythrum (Lythraceae) A-109
Macadamia (Proteaceae) A-142
Macaranga (Euphorbiaceae) A-77
Macfadyena (Bignoniaceae) A-35
Machaerina (Cyperaceae) A-193
Maclura (Moraceae) A-119
Macroptilium (Fabaceae) A-87
Macrothelypteris (Thelypteridaceae) A-249
Macrotyloma (Fabaceae) A-87
Madia (Asteraceae) A-26
Magnoliaceae A-110
Majidea (Sapindaceae) A-160
Malachra (Malvaceae) A-113
Mallotus (Euphorbiaceae) A-77
Malpighiaceae A-110
Malva (Malvaceae) A-113, B-9
Malvaceae A-110, B-9
Malvastrum (Malvaceae) A-113, B-9
Malvaviscus (Malvaceae) A-114
Mangifera (Anacardiaceae) A-6
Manihot (Euphorbiaceae) A-77
Mansoa (Bignoniaceae) A-35
Marantaceae A-199
Marattia (Marattiaceae) A-243
Marattiaceae A-243
Marrubium (Lamiaceae) A-101
Marsilea (Marsileaceae) A-243
Marsileaceae A-243
Martyniaceae A-115
Maurandya (Scrophulariaceae) A-162
Mazus (Scrophulariaceae) A-162
Medicago (Fabaceae) A-87, B-7
Medinilla (Melastomataceae) A-115
Megaskepasma (Acanthaceae) A-2
Melaleuca (Myrtaceae) A-124
Melanthera (Asteraceae) A-26, B-3
Melastoma (Melastomataceae) A-116
Melastomataceae A-115
Melia (Meliaceae) A-117
Meliaceae A-116
Melicope (Rutaceae) A-153
Melilotus (Fabaceae) A-87, B-8
Melinis (Poaceae) A-218, B-15
Melochia (Sterculiaceae) A-167
Menispermaceae A-117
Mentha (Lamiaceae) A-101
24
Merremia (Convolvulaceae) A-65
Metrosideros (Myrtaceae) A-125
Mezoneuron (Fabaceae) A-87
Michelia (Magnoliaceae) A-110
Miconia (Melastomataceae) A-116
Microlepia (Dennstaedtiaceae) A-235
Microsorum (Polypodiaceae) A-244
Mimosa (Fabaceae) A-87
Mirabilis (Nyctaginaceae) A-127, B-10
Miscanthus (Poaceae) A-218
Mitracarpus (Rubiaceae) A-150
Modiola (Malvaceae) A-114
Molineria (Liliaceae) A-199
Molluginaceae A-117
Mollugo (Molluginaceae) A-117
Momordica (Cucurbitaceae) A-68, B-6
Monochoria (Pontederiaceae) A-228
Montanoa (Asteraceae) A-28
Moraceae A-117, B-9
Morella (Myricaceae) A-119
Morelotia (Cyperaceae) A-194
Morinda (Rubiaceae) A-150
Morus (Moraceae) A-119
Mucuna (Fabaceae) A-88
Muehlenbeckia (Polygonaceae) A-138
Muhlenbergia (Poaceae) A-218
Muntingia (Muntingiaceae) A-119
Muntingiaceae A-119
Murdannia (Commelinaceae) A-186
Murraya (Rutaceae) A-158
Musa (Musaceae) A-199
Musaceae A-199
Myoporaceae A-119
Myoporum (Myoporaceae) A-119
Myosotis (Boraginaceae) A-37
Myricaceae A-119
Myriophyllum (Haloragaceae) A-100
Hawaiian Native & Naturalized Vascular Plant Checklist
Myrsinaceae A-119
Myrsine (Myrsinaceae) A-120
Myrtaceae A-121
Nageia (Podocarpaceae) A-178
Najadaceae A-200
Najas (Najadaceae) A-200
Nama (Hydrophyllaceae) A-100, B-8
Nassella (Poaceae) A-218
Nasturtium (Brassicaceae) A-39
Neonotonia (Fabaceae) A-88
Nephrolepidaceae A-243, B-16
Nephrolepis (Nephrolepidaceae) A-243, B-16
Neraudia (Urticaceae) A-169
Nerium (Apocynaceae) A-9
Nertera (Rubiaceae) A-150
Nestegis (Oleaceae) A-129
Nicandra (Solanaceae) A-164
Nicotiana (Solanaceae) A-164, B-11
Noronhia (Oleaceae) A-129
Nothocestrum (Solanaceae) A-164
Nototrichium (Amaranthaceae) A-6
Nyctaginaceae A-127, B-9
Nymphaea (Nymphaeaceae) A-128
Nymphaeaceae A-128
Ochna (Ochnaceae) A-128
Ochnaceae A-128
Ochrosia (Apocynaceae) A-9
Ocimum (Lamiaceae) A-102
Odontonema (Acanthaceae) A-2
Oenothera (Onagraceae) A-130, B-10
Oldenlandia (Rubiaceae) A-150
Oldenlandiopsis (Rubiaceae) A-150
Olea (Oleaceae) A-129
Oleaceae A-128
Onagraceae A-129, B-10
Ophioglossaceae A-244
Ophioglossum (Ophioglossaceae) A-244
Oplismenus (Poaceae) A-218
Opuntia (Cactaceae) A-41
Orchidaceae A-200
Oreobolus (Cyperaceae) A-194
Oreogrammitis (Grammitidaceae) A-240
Orthocarpus (Scrophulariaceae) A-162
Osteomeles (Rosaceae) A-144
Osteospermum (Asteraceae) A-28
Otacanthus (Scrophulariaceae) A-162
Oxalidaceae A-131, B-10
Oxalis (Oxalidaceae) A-131, B-10
Oxyspora (Melastomataceae) A-116
Pachyrhizus (Fabaceae) A-88
Paederia (Rubiaceae) A-151
Palafoxia (Asteraceae) A-28
Palisota (Commelinaceae) A-186
Pandanaceae A-201
Pandanus (Pandanaceae) A-202
Panicum (Poaceae) A-219, B-15
Papaveraceae A-131
Paraserianthes (Fabaceae) A-88
Parentucellia (Scrophulariaceae) A-162
Parkinsonia (Fabaceae) A-88
Parthenium (Asteraceae) A-28
Paspalidium (Poaceae) A-220
Paspalum (Poaceae) A-220, B-15
Passiflora (Passifloraceae) A-131
Passifloraceae A-131
Paulownia (Scrophulariaceae) A-162
Pectis (Asteraceae) A-28
Pedilanthus (Euphorbiaceae) A-77
Pelargonium (Geraniaceae) A-95
Pellaea (Pteridaceae) A-247
Peltophorum (Fabaceae) A-88
Pentas (Rubiaceae) A-151
Peperomia (Piperaceae) A-133
Pereskia (Cactaceae) A-41
25
Perityle (Asteraceae) A-28
Perrottetia (Celastraceae) A-60
Persea (Lauraceae) A-107
Persicaria (Polygonaceae) A-138
Petasites (Asteraceae) A-28
Petrorhagia (Caryophyllaceae) A-56
Petroselinum (Apiaceae) A-8
Peucedanum (Apiaceae) A-8
Phaius (Orchidaceae) A-201
Phalaris (Poaceae) A-222
Philadelphus (Hydrangeaceae) A-100
Philodendron (Araceae) A-180
Phlebodium (Polypodiaceae) A-244
Phleum (Poaceae) A-222
Phoenix (Arecaceae) A-182
Phormium (Agavaceae) A-178
Photinia (Rosaceae) A-144
Phragmites (Poaceae) A-222
Phyla (Verbenaceae) A-172, B-12
Phyllanthus (Euphorbiaceae) A-78
Phyllostachys (Poaceae) A-223
Phyllostegia (Lamiaceae) A-102, B-8
Phymatosorus (Polypodiaceae) A-245
Physalis (Solanaceae) A-164
Phytolacca (Phytolaccaceae) A-133
Phytolaccaceae A-133
Picris (Asteraceae) A-28
Pilea (Urticaceae) A-170, B-12
Pimenta (Myrtaceae) A-126
Pinaceae A-177
Pinanga (Arecaceae) A-182
Pinus (Pinaceae) A-177
Piper (Piperaceae) A-134
Piperaceae A-133
Piptatherum (Poaceae) A-223
Pipturus (Urticaceae) A-170
Piscidia (Fabaceae) A-88
Hawaiian Native & Naturalized Vascular Plant Checklist
Pisonia (Nyctaginaceae) A-127, B-10
Pistia (Araceae) A-181
Pithecellobium (Fabaceae) A-88
Pittosporaceae A-135
Pittosporum (Pittosporaceae) A-135
Pityrogramma (Pteridaceae) A-247
Planchonella (Sapotaceae) A-161
Plantaginaceae A-136, B-10
Plantago (Plantaginaceae) A-136, B-10
Platanthera (Orchidaceae) A-201
Platycerium (Polypodiaceae) A-245
Platycodon (Campanulaceae) A-54
Platydesma (Rutaceae) A-158
Platymiscium (Fabaceae) A-89
Plectranthus (Lamiaceae) A-104
Pleiostachya (Marantaceae) A-199
Pleomele (Agavaceae) A-178
Pluchea (Asteraceae) A-28, B-3
Plumbaginaceae A-137
Plumbago (Plumbaginaceae) A-137
Poa (Poaceae) A-223, B-16
Poaceae A-202, B-13
Podocarpaceae A-177
Podocarpus (Podocarpaceae) A-178
Podranea (Bignoniaceae) A-35
Polemoniaceae A-137
Polyalthia (Annonaceae) A-7
Polycarpon (Caryophyllaceae) A-56
Polygala (Polygalaceae) A-137
Polygalaceae A-137
Polygonaceae A-137, B-10
Polygonum (Polygonaceae) A-138
Polypodiaceae A-244
Polypodium (Polypodiaceae) A-245
Polypogon (Poaceae) A-223, B-16
Polypremum (Loganiaceae) A-109
Polyscias (Araliaceae) A-11
Polystachya (Orchidaceae) A-201
Polystichum (Dryopteridaceae) A-238
Pontederiaceae A-228
Poranopsis (Convolvulaceae) A-65
Portulaca (Portulacaceae) A-139, B-11
Portulacaceae A-139, B-11
Potamogeton (Potamogetonaceae) A-229
Potamogetonaceae A-229
Primulaceae A-140, B-11
Pritchardia (Arecaceae) A-182, B-12
Priva (Verbenaceae) A-172
Proboscidea (Martyniaceae) A-115
Procris (Urticaceae) A-170
Prosopis (Fabaceae) A-89, B-8
Proteaceae A-141
Prunella (Lamiaceae) A-105
Prunus (Rosaceae) A-144
Pseudelephantopus (Asteraceae) A-29
Pseuderanthemum (Acanthaceae) A-2
Pseudognaphalium (Asteraceae) A-29, B-3
Pseudogynoxys (Asteraceae) A-29
Pseudophegopteris (Thelypteridaceae) A-249
Psidium (Myrtaceae) A-126
Psilotaceae A-245, B-17
Psilotum (Psilotaceae) A-245, B-17
Psychotria (Rubiaceae) A-151
Psydrax (Rubiaceae) A-152
Pteralyxia (Apocynaceae) A-9
Pteridaceae A-246
Pteridium (Dennstaedtiaceae) A-236
Pteris (Pteridaceae) A-247
Pterolepis (Melastomataceae) A-116
Ptychosperma (Arecaceae) A-184
Pueraria (Fabaceae) A-89
Pyracantha (Rosaceae) A-144
Pyrostegia (Bignoniaceae) A-35
Pyrrosia (Polypodiaceae) A-245
26
Quercus (Fagaceae) A-93
Radermachera (Bignoniaceae) A-36
Ranunculaceae A-142
Ranunculus (Ranunculaceae) A-142
Raphanus (Brassicaceae) A-39
Rapistrum (Brassicaceae) A-39
Rauvolfia (Apocynaceae) A-9
Reichardia (Asteraceae) A-29
Remya (Asteraceae) A-29
Rhamnaceae A-142
Rhamnus (Rhamnaceae) A-143
Rhizophora (Rhizophoraceae) A-143
Rhizophoraceae A-143
Rhodomyrtus (Myrtaceae) A-126
Rhus (Anacardiaceae) A-6
Rhynchosia (Fabaceae) A-89
Rhynchospora (Cyperaceae) A-194
Richardia (Rubiaceae) A-152
Ricinus (Euphorbiaceae) A-78
Rivina (Phytolaccaceae) A-133
Rorippa (Brassicaceae) A-40
Rosa (Rosaceae) A-145
Rosaceae A-143
Roystonea (Arecaceae) A-184
Rubiaceae A-146
Rubus (Rosaceae) A-145
Ruellia (Acanthaceae) A-2, B-1
Rumex (Polygonaceae) A-138, B-10
Ruppia (Ruppiaceae) A-229
Ruppiaceae A-229
Ruta (Rutaceae) A-158
Rutaceae A-153
Rytidosperma (Poaceae) A-224
Saccharum (Poaceae) A-224
Sacciolepis (Poaceae) A-224
Sadleria (Blechnaceae) A-234
Sagina (Caryophyllaceae) A-56, B-4
Hawaiian Native & Naturalized Vascular Plant Checklist
Sagittaria (Alismataceae) A-179
Salicornia (Chenopodiaceae) A-61, B-5
Salsola (Chenopodiaceae) A-62
Salvia (Lamiaceae) A-105
Salvinia (Salviniaceae) A-248
Salviniaceae A-248
Samanea (Fabaceae) A-89
Sambucus (Caprifoliaceae) A-55
Sanchezia (Acanthaceae) A-3
Sandoricum (Meliaceae) A-117
Sanicula (Apiaceae) A-8
Sansevieria (Agavaceae) A-179
Santalaceae A-159, B-11
Santalum (Santalaceae) A-159, B-11
Sapindaceae A-160
Sapindus (Sapindaceae) A-161
Sapotaceae A-161
Sauropus (Euphorbiaceae) A-78
Sauvagesia (Ochnaceae) A-128
Scabiosa (Dipsacaceae) A-69
Scaevola (Goodeniaceae) A-99, B-8
Schedonnardus (Poaceae) A-224
Schedonorus (Poaceae) A-224
Schefflera (Araliaceae) A-12
Schenkia (Gentianaceae) A-94
Schiedea (Caryophyllaceae) A-56, B-4
Schinus (Anacardiaceae) A-6, B-1
Schizachyrium (Poaceae) A-224
Schizaea (Schizaeaceae) A-248
Schizaeaceae A-248
Schizostachyum (Poaceae) A-225
Schlumbergera (Cactaceae) A-41
Schoenoplectiella (Cyperaceae) A-194
Schoenoplectus (Cyperaceae) A-194
Schoenus (Cyperaceae) A-195
Scleria (Cyperaceae) A-195
Scrophulariaceae A-161, B-11
Sechium (Cucurbitaceae) A-68
Selaginella (Selaginellaceae) A-248
Selaginellaceae A-248
Selenicereus (Cactaceae) A-41
Senecio (Asteraceae) A-30
Senna (Fabaceae) A-89
Serissa (Rubiaceae) A-153
Sesbania (Fabaceae) A-90, B-8
Sesuvium (Aizoaceae) A-3, B-1
Setaria (Poaceae) A-225, B-16
Sherardia (Rubiaceae) A-153
Sicyos (Cucurbitaceae) A-68, B-6
Sida (Malvaceae) A-114, B-9
Sidastrum (Malvaceae) A-114
Sideroxylon (Sapotaceae) A-161
Sigesbeckia (Asteraceae) A-30
Silene (Caryophyllaceae) A-59
Simaroubaceae A-163
Sinapis (Brassicaceae) A-40
Sisymbrium (Brassicaceae) A-40
Sisyrinchium (Iridaceae) A-196
Smilacaceae A-229
Smilax (Smilacaceae) A-229
Solanaceae A-163, B-11
Solandra (Solanaceae) A-165
Solanum (Solanaceae) A-165, B-11
Solidago (Asteraceae) A-30
Soliva (Asteraceae) A-30
Sonchus (Asteraceae) A-30, B-3
Sophora (Fabaceae) A-90
Sorghum (Poaceae) A-225
Spartium (Fabaceae) A-91
Spathodea (Bignoniaceae) A-36
Spathoglottis (Orchidaceae) A-201
Spergula (Caryophyllaceae) A-59
Spergularia (Caryophyllaceae) A-59, B-5
Spermacoce (Rubiaceae) A-153
27
Spermolepis (Apiaceae) A-8
Sphaeropteris (Cyatheaceae) A-235
Sphagneticola (Asteraceae) A-30, B-3
Sphenomeris (Lindsaeaceae) A-242
Sphenopholis (Poaceae) A-225
Spirodela (Lemnaceae) A-198
Sporobolus (Poaceae) A-226, B-16
Stachys (Lamiaceae) A-105
Stachytarpheta (Verbenaceae) A-172, B-12
Stapelia (Asclepiadaceae) A-14
Stellaria (Caryophyllaceae) A-59, B-5
Stemmadenia (Apocynaceae) A-9
Stenogrammitis (Polypodiaceae) A-245
Stenogyne (Lamiaceae) A-105
Stenotaphrum (Poaceae) A-226, B-16
Stephania (Menispermaceae) A-117
Sterculia (Sterculiaceae) A-167
Sterculiaceae A-167, B-11
Sticherus (Gleicheniaceae) A-239
Stictocardia (Convolvulaceae) A-65
Streblus (Moraceae) A-119
Streptosolen (Solanaceae) A-166
Stromanthe (Marantaceae) A-199
Strongylodon (Fabaceae) A-91
Stuckenia (Potamogetonaceae) A-229
Stylosanthes (Fabaceae) A-91
Swietenia (Meliaceae) A-117
Syagrus (Arecaceae) A-184
Symphyotrichum (Asteraceae) A-31
Syncarpia (Myrtaceae) A-126
Synedrella (Asteraceae) A-31
Syngonium (Araceae) A-181
Syzygium (Myrtaceae) A-126
Tabebuia (Bignoniaceae) A-36
Tabernaemontana (Apocynaceae) A-9
Tacca (Taccaceae) A-229
Taccaceae A-229
Hawaiian Native & Naturalized Vascular Plant Checklist
Tagetes (Asteraceae) A-31
Talinum (Portulacaceae) A-140
Tamaricaceae A-167
Tamarindus (Fabaceae) A-91
Tamarix (Tamaricaceae) A-167
Tanacetum (Asteraceae) A-31
Taraxacum (Asteraceae) A-31
Taxodiaceae A-178
Tecoma (Bignoniaceae) A-36
Tectaria (Dryopteridaceae) A-238
Tephrosia (Fabaceae) A-91
Terminalia (Combretaceae) A-63
Tetragonia (Aizoaceae) A-3, B-1
Tetramolopium (Asteraceae) A-31
Tetrapanax (Araliaceae) A-12
Tetrastigma (Vitaceae) A-176
Tetrazygia (Melastomataceae) A-116
Theaceae A-167
Thelypteridaceae A-248
Thelypteris (Thelypteridaceae) A-249
Themeda (Poaceae) A-226
Thespesia (Malvaceae) A-115
Thevetia (Apocynaceae) A-10, B-2
Thunbergia (Acanthaceae) A-3
Thymelaeaceae A-167
Thysanolaena (Poaceae) A-226
Tibouchina (Melastomataceae) A-116
Tiliaceae A-168
Tillandsia (Bromeliaceae) A-185
Tithonia (Asteraceae) A-33
Toona (Meliaceae) A-117
Torenia (Scrophulariaceae) A-163
Torilis (Apiaceae) A-8
Touchardia (Urticaceae) A-170
Tournefortia (Boraginaceae) A-37, B-4
Tradescantia (Commelinaceae) A-186
Tragopogon (Asteraceae) A-33
Tragus (Poaceae) A-227
Trema (Ulmaceae) A-169
Trematolobelia (Campanulaceae) A-54
Trembleya (Melastomataceae) A-116
Trianthema (Aizoaceae) A-3
Tribulus (Zygophyllaceae) A-176, B-12
Trichospermum (Tiliaceae) A-168
Tridax (Asteraceae) A-33, B-3
Trifolium (Fabaceae) A-91
Trimezia (Iridaceae) A-196
Triodanis (Campanulaceae) A-54
Triphasia (Rutaceae) A-158
Triplaris (Polygonaceae) A-139
Trisetum (Poaceae) A-227
Triticum (Poaceae) A-227
Triumfetta (Tiliaceae) A-169
Tropaeolaceae A-169
Tropaeolum (Tropaeolaceae) A-169
Turnera (Turneraceae) A-169
Turneraceae A-169
Typha (Typhaceae) A-229
Typhaceae A-229
Ulex (Fabaceae) A-92
Ulmaceae A-169
Uncinia (Cyperaceae) A-195
Urena (Malvaceae) A-115
Urera (Urticaceae) A-170
Urochloa (Poaceae) A-227, B-16
Urtica (Urticaceae) A-170
Urticaceae A-169, B-12
Utricularia (Lentibulariaceae) A-107
Vaccaria (Caryophyllaceae) A-59
Vaccinium (Ericaceae) A-70
Vachellia (Fabaceae) A-92, B-8
Valerianaceae A-170
Vallisneria (Hydrocharitaceae) A-196
Vanda (Orchidaceae) A-201
28
Vandenboschia (Hymenophyllaceae) A-241
Vanilla (Orchidaceae) A-201
Verbascum (Scrophulariaceae) A-163
Verbena (Verbenaceae) A-172, B-12
Verbenaceae A-171, B-12
Verbesina (Asteraceae) A-33, B-3
Vernonia (Asteraceae) A-33
Veronica (Scrophulariaceae) A-163
Vicia (Fabaceae) A-92
Vigna (Fabaceae) A-92
Vinca (Apocynaceae) A-10
Viola (Violaceae) A-173
Violaceae A-173
Viscaceae A-174
Vitaceae A-175
Vitex (Verbenaceae) A-173
Vitis (Vitaceae) A-176
Vittariaceae A-249
Wahlenbergia (Campanulaceae) A-54
Waltheria (Sterculiaceae) A-167, B-11
Washingtonia (Arecaceae) A-184
Watsonia (Iridaceae) A-197
Werauhia (Bromeliaceae) A-185
Wikstroemia (Thymelaeaceae) A-167
Wilkesia (Asteraceae) A-33
Wolffia (Lemnaceae) A-198
Xanthium (Asteraceae) A-33, B-3
Xanthosoma (Araceae) A-181
Xylosma (Flacourtiaceae) A-93
Xyridaceae A-229
Xyris (Xyridaceae) A-229
Youngia (Asteraceae) A-33
Zantedeschia (Araceae) A-181
Zanthoxylum (Rutaceae) A-158
Zephyranthes (Liliaceae) A-199
Zeuxine (Orchidaceae) A-201
Zingiber (Zingiberaceae) A-230
Hawaiian Native & Naturalized Vascular Plant Checklist
Zingiberaceae A-230
Zinnia (Asteraceae) A-33
Zizania (Poaceae) A-228
Ziziphus (Rhamnaceae) A-143
Zornia (Fabaceae) A-93
Zoysia (Poaceae) A-228
Zygophyllaceae A-176, B-12
29
APPENDIX A. Main Hawaiian Islands Native and Naturalized Vascular Plants
Scientific Name
Dicot
Acanthaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Aphelandra aurantiaca (Scheidw.) Lindl.
nat
Asystasia gangetica (L.) T.Anderson
nat
K
O Mo L
M
H
Wagner et al. 1990:168 (K, O, Mo, M); Oppenheimer & Bartlett 2000:1 (H);
Staples & Herbst 2005:81 (KEY), 82 (DESCR); Oppenheimer 2007:18 (L)
Barleria cristata L.
nat
K
O Mo
M
H
Wagner et al. 1990:168 (K, O, H); Oppenheimer & Bartlett 2002:1 (WM);
Staples & Herbst 2005:81 (KEY), 82 (DESCR); Starr et al. 2006:31 (Mo)
Barleria lupulina Lindl.
O
Citation
?
O?
Barleria repens Nees
nat
O
Dicliptera chinensis (L.) Juss.
nat
Hemigraphis alternata (Burm.f.) T.Anderson
nat
Hemigraphis reptans (G.Forst.) T.Anderson
nat
Hypoestes phyllostachya Baker
nat
Justicia betonica L.
nat
Justicia carnea Lindl.
nat
K
Staples & Herbst 2005:81 (KEY, DESCR [as var. aurantiaca]); Daehler &
Baker 2006:4 (NEWNAT/O, DESCR)
Wagner et al. 1990:170 (O? [cultivated, perhaps escaping at Hälawa Beach
Park], DESCR); Staples & Herbst 2005:81 (KEY), 83 (DESCR)
L
O Mo
M
M
Staples et al. 2002:3 (NEWNAT/O, DESCR); Oppenheimer 2003:3 (L, EM,
WM, DESCR); Staples & Herbst 2005:81 (KEY), 83 (DESCR)
H
O
K
K
Wagner et al. 1990:171 (K, O); Staples et al. 2002:3 (H); Oppenheimer
2003:3 (WM); Oppenheimer 2004:8 (EM); Oppenheimer 2007:18 (Mo)
Wagner et al. 1990:171 (O); Staples & Herbst 2005:80 (KEY), 85 (DESCR)
O Mo
O Mo L
M
H
Wagner et al. 1990:171 (K, O, H); Oppenheimer et al. 1999:7 (EM);
Oppenheimer & Bartlett 2002:1 (WM); Staples et al. 2002:3 (Mo); Staples &
Herbst 2005:80 (KEY), 86 (DESCR)
H
Herbst & Wagner 1999:12 (NEWNAT/H); Staples & Herbst 2005:80 (KEY),
86 (DESCR)
M
H
Wagner et al. 1990:172 (K, O, Mo, L, M, H); Staples & Herbst 2005:81, 87
(KEY), 87 (DESCR)
M
H
Oppenheimer 2004:8 (NEWNAT/EM, DESCR); Staples & Herbst 2005:81,
87 (KEY), 87 (DESCR); Parker & Parsons 2012b:65 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 1
Scientific Name
Dicot
Acanthaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O Mo
Citation
Justicia spicigera Schltdl.
nat
Wagner et al. 1990:172 (Mo? [possibly sparingly naturalized in Hälawa
Valley]); Staples et al. 2002:4 (NEWNAT/O, Mo, DESCR); Staples & Herbst
2005:81, 87 (KEY), 87 (DESCR)
Megaskepasma erythrochlamys Lindau
nat
K
O
Odontonema cuspidatum (Nees) Kuntze
nat
K
O
Pseuderanthemum variabile (R.Br.) Radlk.
nat
K
O
Ruellia blechum L.
nat
Ruellia brevifolia (Pohl) C.Ezcurra
nat
K
O Mo L
M
Wagner et al. 1990:174 (K, O [as Ruellia graecizans]); Ezcurra
1993:802/Herbst & Wagner 1999:13 (NOM:Syn. R. graecizans);
Oppenheimer 2003:4 (Mo, EM); Staples & Herbst 2005:80, 81, 91 (KEY), 91
(DESCR); Starr et al. 2010:61 (L)
Ruellia brittoniana Leonard
nat
K
O
M
Wagner et al. 1990:174 (K, O); Oppenheimer & Bartlett 2002:2 (WM);
Staples & Herbst 2005:80, 81, 91 (KEY), 91 (DESCR)
Ruellia devosiana E.Morren
nat
K
Ruellia prostrata Poir.
nat
M
H
Staples & Herbst 2005:81 (KEY), 88 (DESCR); Frohlich & Lau 2010:3
(NEWNAT/O, DESCR); Frohlich & Lau 2012:27 (K); Parker & Parsons
2012a:55 (H)
H
Wagner et al. 1990:167 (K?, O?, H? [as Odontonema strictum; non-cultivated
plants not appearing to set fruit, thus not truly naturalized]); Lorence et al.
1995:20 (NEWNAT/K, DESCR, NOM: O. strictum misapplied);
Oppenheimer & Bartlett 2002:2 (EM); Staples et al. 2002:4 (H); Staples &
Herbst 2005:81 (KEY), 88 (DESCR); Frohlich & Lau 2007:9 (O)
Herbarium Pacificum Staff 1998:8 (NEWNAT/O [as Pseuderanthemum
fasciculatum], DESCR); Herbst & Wagner 1999:12 (NOM: P. fasciculatum
misapplied); Staples et al. 2003:7 (K)
O
H
Wagner et al. 1990:170 (O [as Blechum brownei]); Wagner & Herbst
1995:14 (H, NOM:Syn. Blechum brownei = B. pyramidatum); Tripp et al.
2009:900/Wagner et al. 2012:3 (NOM:Syn. B. pyramidatum)
Lorence et al. 1995:20 (NEWNAT/K, DESCR); Staples & Herbst 2005:80,
81, 91 (KEY), 91 (DESCR)
O Mo
M
Wagner et al. 1990:174 (O); Oppenheimer 2003:4 (Mo, WM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 2
Scientific Name
Dicot
Acanthaceae
Status
Kl Le Ni K
O Mo L
Sanchezia parvibracteata Sprague & Hutch.
nat
Sanchezia speciosa Leonard
nat
K
Thunbergia alata Bojer ex Sims
nat
K
Mo L
Thunbergia fragrans Roxb.
nat
K
Thunbergia grandiflora Roxb.
nat
Thunbergia laurifolia Lindl.
M Ka H
Citation
H
Staples & Herbst 2005:80 (KEY), 92 (DESCR); Parker & Parsons 2012a:55
(NEWNAT/H)
H
Lorence et al. 1995:20 (NEWNAT/K, DESCR); Herbst & Wagner 1999:13
(author correction); Staples & Herbst 2005:80 (KEY), 92 (DESCR); Parker &
Parsons 2012b:65 (H)
M
H
Wagner et al. 1990:175 (K, Mo, EM, H); Oppenheimer 2003:4 (WM);
Staples & Herbst 2005:81, 93 (KEY), 93 (DESCR); Starr et al. 2010:61 (L)
O Mo L
M
H
Wagner et al. 1990:175 (K, O, M, H); Staples & Herbst 2005:81, 93 (KEY),
95 (DESCR); Starr et al. 2006:31 (Mo); Starr et al. 2010:61 (L)
K
O
M
H
Wagner et al. 1990:175 (K, O, H); Starr et al. 2002:16 (EM); Staples &
Herbst 2005:81, 93 (KEY), 94 (DESCR)
nat
K
O
M
H
Wagner et al. 1990:175 (K, O); Starr et al. 1999:11 (EM); Oppenheimer
2003:4 (H); Staples & Herbst 2005:81, 93 (KEY), 94 (DESCR); Starr et al.
2006:31 (L)
?
K?
L
Aizoaceae
Aptenia cordifolia (L.f.) Schwantes
Sesuvium portulacastrum (L.) L.
ind
Sesuvium verrucosum Raf.
nat
Tetragonia tetragonioides (Pall.) Kuntze
nat
Trianthema portulacastrum L.
nat
Le Ni
K
Staples & Herbst 2005:96 (KEY, DESCR); Frohlich & Lau 2012:49 (K?
[adventive])
O Mo L
M Ka
O Mo
M
K
O Mo L
M
K
O
M
H
Wagner et al. 1990:176 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:96 (KEY), 99 (DESCR); Wood & LeGrande 2006:19 (Le)
Frohlich & Lau 2012:27 (NEWNAT/O, Mo, EM, DESCR)
H
Wagner et al. 1990:178 (K, O, H); Oppenheimer et al. 1999:7 (WM); Starr et
al. 2003:23 (EM); Staples & Herbst 2005:96 (KEY), 98 (DESCR); Starr et al.
2006:31 (Mo); Oppenheimer 2010:33 (L)
Wagner et al. 1990:179 (O); Herbst & Wagner 1996:9 (WM); Lorence &
Flynn 2006:2 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 3
Scientific Name
Dicot
Amaranthaceae
Achyranthes aspera L. var. aspera
Achyranthes aspera L. var. pubescens (Moq.)
C.C.Towns.
Status
Kl Le Ni K
nat
O Mo L
M Ka H
O Mo
?
H
L?
Achyranthes atollensis H.St.John
end
Achyranthes mutica A.Gray
end
Achyranthes splendens Mart. ex Moq. var. rotundata
Hillebr.
end
O Mo L
Achyranthes splendens Mart. ex Moq. var. splendens
end
L
Alternanthera brasiliana (L.) Kuntze
nat
Alternanthera caracasana Kunth
nat
Alternanthera pungens Kunth
Citation
Wagner et al. 1990:181 (O, H); Staples et al. 2002:4 (Mo)
Wagner et al. 1990:181 (L? [single coll. in 1850s, likely no longer present],
DESCR)
Only in Northwestern Hawaiian Islands; see NWHI checklist
K
H
Wagner et al. 1990:181 (K, H? [presumed location for 1779 coll.]); Hawley
& Pyle 1995:1 (REDISCOVER/H)
Wagner et al. 1990:182 (O, Mo, L); Staples & Herbst 2005:99 (KEY), 103
(DESCR)
M
Wagner et al. 1990:182 (L, M)
O Mo
M?
Wagner et al. 1990:183 (Mo); Wagner & Herbst 1995:14 (O); Starr & Starr
2011:23 (EM); Wagner et al. 2012:5 (EM record adventive)
K
O Mo L
M
Wagner et al. 1990:183 (K, O, Mo, L, M)
nat
K
O Mo L
M
H
Wagner et al. 1990:183 (O, Mo, H); Lorence & Flynn 1999:3 (K);
Oppenheimer & Bartlett 2000:1 (WM); Oppenheimer & Bartlett 2002:2 (L);
Starr et al. 2002:17 (EM)
Alternanthera sessilis (L.) R.Br. ex DC.
nat
K
O Mo L
M
H
Wagner et al. 1990:185 (O, Mo, M, H); Lorence et al. 1995:21 (K); Staples &
Herbst 2005:99, 100 (KEY), 101 (DESCR); Imada 2007:34 (L)
Alternanthera tenella Colla
nat
Amaranthus brownii Christoph. & Caum
end
Amaranthus dubius Mart. ex Thell.
nat
O
Wagner et al. 1990:185 (O); Staples & Herbst 2005:99, 100 (KEY), 100
(DESCR)
Only in Northwestern Hawaiian Islands; see NWHI checklist
K
O
L
M
H
Wagner et al. 1990:186 (K, O, L, H); Staples & Herbst 2005:99 (KEY), 101
(DESCR); Starr et al. 2006:32 (EM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 4
Scientific Name
Dicot
Amaranthaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Amaranthus graecizans L.
nat
O
Amaranthus hybridus L. ssp. hybridus
nat
O
M
Amaranthus lividus L. ssp. polygonoides (Moq.) Probst
nat
O
M?
Amaranthus retroflexus L.
nat
Amaranthus spinosus L.
nat
Amaranthus viridis L.
nat
Blutaparon vermiculare (L.) Mears
nat
Celosia argentea L.
nat
Charpentiera densiflora Sohmer
end
K
Wagner et al. 1990:190 (K)
Charpentiera elliptica (Hillebr.) A.Heller
end
K
Wagner et al. 1990:190 (K)
Charpentiera obovata Gaudich.
end
K
Charpentiera ovata Gaudich. var. niuensis Sohmer
end
O
Charpentiera ovata Gaudich. var. ovata
end
O Mo
Charpentiera tomentosa Sohmer var. maakuaensis
Sohmer
end
O
Charpentiera tomentosa Sohmer var. tomentosa
end
O Mo L
Gomphrena celosioides Mart.
nat
K
Kl
Herbst et al. 2004:3 (NEWNAT/O, DESCR)
Wagner et al. 1990:188 (O, M)
H
Wagner et al. 1990:188 (K, O, M? [apparently collected once in 1869–1872],
H)
H
Wagner et al. 1997:51 (NEWNAT/H, DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:188 (K, O, Mo, M, Ka, H); Oppenheimer 2003:5 (L)
K
O Mo L
M Ka
H
Wagner et al. 1990:189 (K, O, L, M, Ka, H); Hughes 1995:1 (Mo)
O
Wagner & Herbst 1995:14 (NEWNAT/O, DESCR)
H
K
O Mo L
M
H
Oppenheimer 2003:5 (NEWNAT/H, DESCR); Staples & Herbst 2005:99
(KEY), 101 (DESCR)
Wagner et al. 1990:191 (K, O, Mo, L, M, H)
Wagner et al. 1990:191 (O)
M
H
Wagner et al. 1990:191 (O, Mo, M, H)
Wagner et al. 1990:191 (O)
O
M
H
Wagner et al. 1990:191 (O, Mo, L, M, H)
Wagner et al. 1990:192 (O); Wagner & Herbst 1995:15 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 5
Scientific Name
Dicot
Amaranthaceae
Status
Kl Le Ni K
Gomphrena globosa L.
nat
K
Nototrichium divaricatum Lorence
end
K
Nototrichium humile Hillebr.
end
Nototrichium sandwicense (A.Gray) Hillebr.
end
Ni
K
O Mo L
M Ka H
O
Citation
Wagner et al. 1990:192 (O); Lorence et al. 1995:21 (K); Staples & Herbst
2005:99 (KEY), 102 (DESCR)
Lorence 1996:64 (SPNOV/K, DESCR); Herbst & Wagner 1999:13 (DESCR)
O
M
O Mo L
M Ka
Wagner et al. 1990:193 (O, EM)
H
Wagner et al. 1990:194 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:99 (KEY), 103 (DESCR)
Anacardiaceae
Anacardium occidentale L.
?
Mangifera indica L.
nat
Rhus sandwicensis A.Gray
end
Rhus taitensis Guill.
nat
Schinus molle L.
nat
Schinus terebinthifolius Raddi
nat
K? O?
Ni
Wagner et al. 1990:196 (K?, O? [forestry plantings, not naturalized], DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:196 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:104 (KEY), 105 (DESCR)
K
O Mo
M
H
Wagner et al. 1990:197 (K, O, Mo, M, H)
O
K
Daehler & Baker 2006:4 (NEWNAT/O, DESCR)
O Mo L
M
H
Wagner et al. 1990:198 (EM, H); Staples & Herbst 2005:104 (KEY), 106
(DESCR)
M
H
Wagner et al. 1990:198 (K, O, Mo, L, M, H); Staples & Herbst 2005:104
(KEY), 106 (DESCR)
H
Staples et al. 2002:4 (NEWNAT/H, DESCR); Staples & Herbst 2005:108
(KEY), 109 (DESCR)
Annonaceae
Annona cherimola Mill.
nat
Annona muricata L.
nat
Mo
Staples & Herbst 2005:108 (KEY), 109 (DESCR); Wysong et al. 2007:2
(NEWNAT/Mo)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 6
Scientific Name
Dicot
Annonaceae
Status
Kl Le Ni K
O Mo L
M Ka H
M
H
Citation
Artabotrys hexapetalus (L.f.) Bhandari
nat
O
Nagata 1995:11 (NEWNAT/O, H); Starr et al. 2003:24 (EM, DESCR);
Staples & Herbst 2005:108 (KEY), 110 (DESCR)
Polyalthia suberosa (Roxb.) Thwaites
?
O?
Anethum graveolens L.
nat
O
M?
Centella asiatica (L.) Urb.
nat
K
O Mo L
M
H
Wagner et al. 1990:201 (K, O, Mo, L, M, H); Staples & Herbst 2005:112
(KEY), 115 (DESCR)
Ciclospermum leptophyllum (Pers.) Sprague ex Britton
& P.Wilson
nat
K
O Mo L
M
H
Wagner et al. 1990:201 (K, O, Mo, L, M, H)
Coriandrum sativum L.
nat
K
O
M
Cryptotaenia canadensis (L.) DC.
nat
K
O Mo
M
Daucus pusillus Michx.
nat?
K
O Mo L
Eryngium foetidum L.
?
Staples & Herbst 2005:108 (KEY), 111 (DESCR); Frohlich & Lau 2010:4
(NEWNAT/O, DESCR); Wagner et al. 2012:6 (O record adventive)
Apiaceae
Wagner et al. 1990:200 (O, M? [unvouchered Hillebrand (1888) citation]);
Staples & Herbst 2005:112 (KEY), 113 (DESCR)
Wagner et al. 1990:203 (O); Staples et al. 2002:5 (K); Oppenheimer 2003:5
(EM); Staples & Herbst 2005:112 (KEY), 113 (DESCR)
H
Wagner et al. 1990:203 (K, O, H); Oppenheimer et al. 1999:7 (Mo, WM);
Staples & Herbst 2005:112 (KEY), 114 (DESCR)
H
Wagner et al. 1990:204 (O, Mo, L, H); Lorence et al. 1995:22 (K)
O?
Wagner et al. 1990:199 (O? [cultivated, possibly naturalizing near Käne‘ohe],
DESCR); Staples & Herbst 2005:112 (KEY), 115 (DESCR)
Foeniculum vulgare Mill.
nat
K
O
L
M
H
Wagner et al. 1990:204 (K, O, L, M, H); Staples & Herbst 2005:112 (KEY),
115 (DESCR)
Hydrocotyle bowlesioides Mathias & Constance
nat
K
O
L
M
H
Wagner et al. 1990:205 (K, O, H); Oppenheimer & Bartlett 2002:2 (L, WM);
Starr et al. 2003:24 (EM)
Hydrocotyle sibthorpioides Lam.
nat
K
O
M
H
Wagner et al. 1990:205 (K, O, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 7
Scientific Name
Dicot
Apiaceae
Status
Hydrocotyle verticillata Thunb.
nat?
Petroselinum crispum (Mill.) A.W.Hill
Kl Le Ni K
Ni
O Mo L
M Ka H
Citation
K
O Mo
M
H
Wagner et al. 1990:205 (Ni, K, O, Mo, M, H)
nat
K
O
M
H
Wagner et al. 1990:207 (K, O, M, H); Staples & Herbst 2005:112 (KEY),
115 (DESCR)
Peucedanum sandwicense Hillebr.
end
K
O Mo
M
Sanicula kauaiensis H.St.John
end
K
Sanicula mariversa Nagata & S.M.Gon
end
O
Sanicula purpurea H.St.John & Hosaka
end
O
Sanicula sandwicensis A.Gray
end
Spermolepis hawaiiensis H.Wolff
end
K
Torilis arvensis (Huds.) Link
nat
K
Torilis nodosa (L.) Gaertn.
nat
Wagner et al. 1990:208 (K, Mo, EM, WM); Herbarium Pacificum Staff
1998:9 (O)
Wagner et al. 1990:209 (K)
Wagner et al. 1990:209 (O)
M
O Mo L
Wagner et al. 1990:210 (O, WM)
M
H
Wagner et al. 1990:210 (EM, H); Oppenheimer 2004:9 (WM)
M
H
Wagner et al. 1990:212 (K, O, Mo, M, H); Lorence et al. 1995:22
(REDISCOVER/K); Wagner & Herbst 1995:15 (L, REDISCOVER/Mo)
M
Staples et al. 2002:5 (NEWNAT/K, KEY); Starr & Starr 2012:87 (EM)
H
Wagner et al. 1990:213 (H); Staples et al. 2002:5 (KEY)
Apocynaceae
Allamanda schottii Pohl
?
Alstonia macrophylla Wall. ex G.Don
nat
Alyxia stellata (J.R.Forst. & G.Forst.) Roem. & Schult.
ind
K?
Staples & Herbst 2005:117 (KEY), 119 (DESCR); Frohlich & Lau 2012:49
(K? [adventive], DESCR)
O
Ni? K
H
O Mo L
M Ka? H
Wagner et al. 1997:51 (NEWNAT/H); Herbst et al. 2004:3 (O); Daehler &
Baker 2006:12 (O)
Wagner et al. 1990:214 (Ni?, K, O, Mo, L, M, Ka?, H [as Alyxia oliviformis;
not vouchered but likely formerly occurring on Ni & Ka]); Middleton
2002:56 (NOM:Syn. A. oliviformis, KEY, DESCR); Staples & Herbst
2005:117 (KEY), 128 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 8
Scientific Name
Dicot
Apocynaceae
Beaumontia multiflora Teijsm. & Binn.
Status
Kl Le Ni K
O Mo L
?
H?
Carissa macrocarpa (Eckl.) A.DC.
nat
Catharanthus roseus (L.) G.Don
nat
Nerium oleander L.
nat
Ochrosia compta K.Schum.
end
Ochrosia haleakalae H.St.John
end
Ochrosia kauaiensis H.St.John
end
Ochrosia kilaueaensis H.St.John
end
Pteralyxia kauaiensis Caum
end
Pteralyxia macrocarpa (Hillebr.) K.Schum.
end
Rauvolfia sandwicensis A.DC.
end
Rauvolfia vomitoria Afzel.
nat
Stemmadenia litoralis (Kunth) L.Allorge
nat
O
?
O?
Tabernaemontana pandacaqui Poir.
M Ka H
O
Ni
K
O Mo
M Ka
H
Wagner et al. 1990:215 (Ni, O, Mo, M, Ka, H); Lorence et al. 1995:22 (K);
Staples & Herbst 2005:117 (KEY), 120 (DESCR)
H
Staples & Herbst 2005:117 (KEY), 122 (DESCR); Parker & Parsons
2012a:55 (NEWNAT/H)
M
M
Wagner et al. 1990:218 (O, Mo); Oppenheimer 2006:10 (WM)
H
K
Wagner et al. 1990:218 (EM, H)
Wagner et al. 1990:218 (K)
H
K
Wagner et al. 1990:218 (H)
Wagner et al. 1990:219 (K)
O
K
Staples & Herbst 2005:117 (KEY), 128 (DESCR); Parker & Parsons
2012b:65 (H?, potential naturalization)
Staples & Herbst 2005:116 (KEY), 119 (DESCR); Lau & Frohlich 2012:7
(NEWNAT/O, DESCR)
O Mo
Ni
Citation
Wagner et al. 1990:220 (O)
O Mo L
M
H
Wagner et al. 1990:220 (Ni, K, O, Mo, L, M, H)
H
Kennedy et al. 2010:19 (NEWNAT/H, DESCR)
H
Staples & Herbst 2005:117 (KEY), 125 (DESCR); Lau & Frohlich 2012:7
(NEWNAT/O, DESCR); Parker & Parsons 2012a:56 (NEWNAT/H)
Staples & Herbst 2005:117 (KEY), 126 (DESCR); Frohlich & Lau 2010:4
(NEWNAT/O, DESCR); Wagner et al. 2012:6 (O record adventive)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 9
Scientific Name
Dicot
Apocynaceae
Status
Thevetia peruviana (Pers.) K.Schum.
nat
Vinca major L.
nat
Kl Le Ni K
K
O Mo L
O Mo L
M Ka H
Citation
M
Wagner et al. 1990:215 (K, O, M [as Cascabela thevetia]); Wagner & Herbst
1999:1858 (NOM:Syn. C. thevetia); Staples & Herbst 2005:116 (KEY), 127
(DESCR); Wysong et al. 2007:2 (Mo); Oppenheimer 2011:5 (L)
M
Starr et al. 2002:17 (NEWNAT/EM, DESCR)
Aquifoliaceae
Ilex anomala Hook. & Arn.
ind
K
O Mo L
Ilex aquifolium L.
nat
Ilex cassine L.
nat
O
Ilex paraguariensis St.-Hil.
nat
O
M
H
Wagner et al. 1990:222 (K, O, Mo, L, M, H)
M
H
Wagner et al. 1990:222 (H); Wagner & Herbst 1995:15 (EM); Staples &
Herbst 2005:129 (KEY), 130 (DESCR)
H
Wagner et al. 1990:222 (O?, H? [planted but not naturalized]); Imada et al.
2000:9 (NEWNAT/O, H); Staples & Herbst 2005:129 (KEY), 130 (DESCR)
Wagner et al. 1990:224 (O); Staples & Herbst 2005:129 (KEY), 130
(DESCR)
Araliaceae
Cheirodendron dominii Krajina
end
K
Wagner et al. 1990:226 (K)
Cheirodendron fauriei Hochr.
end
K
Wagner et al. 1990:226 (K)
Cheirodendron forbesii (Sherff) Lowry
end
K
Wagner et al. 1990:227 (K)
Cheirodendron platyphyllum (Hook. & Arn.) Seem. ssp.
kauaiense (Krajina) Lowry
end
K
Wagner et al. 1990:227 (K)
Cheirodendron platyphyllum (Hook. & Arn.) Seem. ssp.
platyphyllum
end
Cheirodendron trigynum (Gaudich.) A.Heller ssp.
helleri (Sherff) Lowry
end
O
Wagner et al. 1990:227 (O)
K
Wagner et al. 1990:228 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 10
Scientific Name
Dicot
Araliaceae
Status
Kl Le Ni K
Cheirodendron trigynum (Gaudich.) A.Heller ssp.
trigynum
end
Hedera helix L.
nat
K
Polyscias bisattenuata (Sherff) Lowry & G.M.Plunkett
end
K
Polyscias filicifolia (C.Moore ex E.Fourn.) L.H.Bailey
?
Polyscias flynnii (Lowry & K.R.Wood) Lowry &
G.M.Plunkett
end
Polyscias gymnocarpa (Hillebr.) Lowry & G.M.Plunkett
end
Polyscias hawaiensis (A.Gray) Lowry & G.M.Plunkett
end
Polyscias kavaiensis (H.Mann) Lowry & G.M.Plunkett
end
Polyscias lydgatei (Hillebr.) Lowry & G.M.Plunkett
end
Polyscias oahuensis (A.Gray) Lowry & G.M.Plunkett
end
Ni
O Mo L
M Ka H
O Mo L
M
H
Wagner et al. 1990:228 (Ni, O, Mo, L, M, H)
O
M
H
Wagner et al. 1990:228 (K, O, H); Starr et al. 2003:24 (EM); Staples &
Herbst 2005:131 (KEY), 133 (DESCR)
Wagner et al. 1990:235 (as Tetraplasandra bisattenuata, syn. of T. oahuensis);
Lowry & Plunkett 2010:73/Wagner et al. 2012:7 (RESURRECT/K,
COMBNOV:Syn. T. bisattenuata)
O?
Staples & Herbst 2005:132, 134 (KEY), 134 (DESCR); Frohlich & Lau
2012:50 (O? [adventive])
K
Lowry & Wood 2000:40 (SPNOV/K [as Tetraplasandra flynnii], KEY,
DESCR); Lowry & Plunkett 2010:74 (COMBNOV:Syn. T. flynnii)
O
K
O
Wagner et al. 1990:234 (O [as Tetraplasandra gymnocarpa]); Lowry &
Plunkett 2010:74 (COMBNOV:Syn. T. gymnocarpa)
Mo L
M
H
Wagner et al. 1990:234 (Mo, L, M, H [as Tetraplasandra hawaiensis]); Lowry
& Plunkett 2010:74 (COMBNOV:Syn. T. hawaiensis)
L
M
H
Wagner et al. 1990:235 (K, O, L, M, H [as Tetraplasandra kavaiensis]);
Lowry & Wood 2000:43 (KEY); Lowry & Plunkett 2010:74
(COMBNOV:Syn. T. kavaiensis)
O
K
Citation
Wagner et al. 1990:235 (as Tetraplasandra lydgatei, syn. of T. oahuensis);
Motley 2005:106 (RESURRECT/O, KEY, DESCR); Lowry & Plunkett
2010:74 (COMBNOV:Syn. T. lydgatei)
O Mo L
M
H
Wagner et al. 1990:235 (K, O, Mo, L, M, H [as Tetraplasandra oahuensis]);
Motley 2005:106 (KEY); Lowry & Plunkett 2010:74 (COMBNOV:Syn. T.
oahuensis)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 11
Scientific Name
Dicot
Araliaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Polyscias racemosa (C.N.Forbes) Lowry & G.M.Plunkett
end
Polyscias sandwicensis (A.Gray) Lowry & G.M.Plunkett
end
Polyscias waialealae (Rock) Lowry & G.M.Plunkett
end
K
Wagner et al. 1990:236 (K [as Tetraplasandra waialealae]); Lowry & Plunkett
2010:75 (COMBNOV:Syn. T. waialealae)
Polyscias waimeae (Wawra) Lowry & G.M.Plunkett
end
K
Wagner et al. 1990:236 (K [as Tetraplasandra waimeae]); Lowry & Plunkett
2010:75 (COMBNOV:Syn. T. waimeae)
Schefflera actinophylla (Endl.) Harms
nat
K
O Mo L
M
H
Wagner et al. 1990:232 (K, O, M, H); Staples et al. 2002:5 (Mo); Staples &
Herbst 2005:132 (KEY), 137 (DESCR); Oppenheimer 2007:19 (L)
Schefflera arboricola (Hayata) Merr.
nat
K
O?
M
H
Starr et al. 2003:24 (NEWNAT/EM, DESCR); Staples & Herbst 2005:132
(KEY), 137 (DESCR); Frohlich & Lau 2008:10 (O? [adventive]); Frohlich &
Lau 2010:4 (O); Frohlich & Lau 2012:28 (K); Parker & Parsons 2012a:56
(H); Wagner et al. 2012:7 (O record adventive)
Schefflera elegantissima (Veitch ex Masters) Lowry &
Frodin
K
Citation
Ni
Wagner et al. 1990:229 (K [as Munroidendron racemosum]); Herbst &
Wagner 1996:9 (REDISCOVER/K); Lowry & Plunkett 2010:74
(COMBNOV:Syn. M. racemosum)
O Mo L
M
H
Wagner et al. 1990:231 (Ni, O, Mo, L, M, H [as Reynoldsia sandwicensis]);
Lowry & Plunkett 2010:74 (COMBNOV:Syn. R. sandwicensis)
?
O?
Staples & Herbst 2005:132 (KEY), 138 (DESCR); Frohlich & Lau 2010:4
(NEWNAT/O, DESCR); Wagner et al. 2012:7 (O record adventive)
Schefflera heptaphylla (L.) Frodin
nat
O
Daehler & Baker 2006:13 (O? [adventive, as Schefflera taiwaniana]);
Frohlich & Lau 2012:28 (NEWNAT/O [as S. heptaphylla], DESCR, NOM:
S. taiwaniana misapplied)
Schefflera insularum (Seem.) Harms
nat
O
Frohlich & Lau 2012:29 (NEWNAT/O, DESCR)
Tetrapanax papyrifer (Hook.) K.Koch
nat
O
H
Staples & Herbst 2005:131 (KEY), 138 (DESCR); Lau & Frohlich 2012:8
(NEWNAT/O, DESCR); Parker & Parsons 2012a:56 (NEWNAT/H, DESCR)
Aristolochiaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 12
Scientific Name
Dicot
Aristolochiaceae
Aristolochia littoralis Parodi
Status
Kl Le Ni K
nat
O Mo L
M Ka H
Citation
K
O
M
Wagner et al. 1990:237 (O); Imada et al. 2000:10 (K); Starr et al. 2003:25
(EM); Staples & Herbst 2005:139 (DESCR); Oppenheimer 2007:19 (WM)
K
O Mo L
M Ka
H
Wagner et al. 1990:240 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:19 (Le? [unvouchered report by Caum in 1936])
K
O Mo L
M Ka
H
Wagner et al. 1990:240 (K, O, L, M, Ka, H); Oppenheimer 2007:19 (Mo)
O? Mo
M Ka? H?
Wagner et al. 1990:238 (Ka?, H? [reportedly escaping on Kaho‘olawe and
Kohala & Kona districts, Hawai‘i]); Starr et al. 2003:25 (NEWNAT/EM,
WM, DESCR); Starr et al. 2004:20 (K); Staples & Herbst 2005:140 (KEY,
DESCR); Wysong et al. 2007:2 (Mo); Frohlich & Lau 2008:3 (O); Starr &
Starr 2012:87 (remove K, WM from distribution, spms. reidentified as
Calotropis procera); Wagner et al. 2012:8 (O record adventive)
M Ka
Wagner et al. 1990:238 (H? [reported escaping by Keähole Airport]); Staples
& Herbst 2005:140 (KEY), 141 (DESCR); Wood & LeGrande 2006:19
(NEWNAT/Le); Wood 2006:15 (K); Oppenheimer 2008:23 (L); Parker &
Parsons 2012a:56 (H); Starr & Starr 2012:87 (EM, WM, Ka, NOM:
Calotropis gigantea misapplied, in part)
Asclepiadaceae
Asclepias curassavica L.
nat
Le? Ni
Asclepias physocarpa (E.Mey.) Schltr.
nat
Calotropis gigantea (L.) W.T.Aiton
nat
Calotropis procera (Aiton) W.T.Aiton
nat
Cryptostegia grandiflora Roxb. ex R.Br.
nat
O
Cryptostegia madagascariensis Bojer ex Decne.
nat
O Mo
Cynanchum gerrardii (Harv.) Liede
nat
O
Hoya australis R.Br. ex Traill
nat
Le
K
L
H
Staples & Herbst 2005:142 (DESCR); Lau & Frohlich 2012:8 (NEWNAT/O,
DESCR)
H
Staples & Herbst 2005:140 (KEY), 142 (DESCR); Staples et al. 2006:6
(NEWNAT/Mo, DESCR); Frohlich & Lau 2008:3 (O); Parker & Parsons
2012a:57 (H)
Frohlich & Lau 2012:29 (NEWNAT/O, DESCR)
M
Lorence & Flynn 1999:3 (NEWNAT/EM, DESCR); Staples & Herbst
2005:140, 142 (KEY), 142 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 13
Scientific Name
Dicot
Asclepiadaceae
Stapelia gigantea N.E.Br.
Status
Kl Le Ni K
nat
O Mo L
M Ka H
Citation
O Mo
M
H
Wagner et al. 1990:241 (O); Oppenheimer et al. 1999:7 (WM); Staples &
Herbst 2005:140 (KEY), 144 (DESCR, NOM:Syn. Stapelia nobilis
[mentioned in Wagner et al. 1990:241]); Wysong et al. 2007:2 (Mo);
Oppenheimer 2010:33 (EM); Parker & Parsons 2012a:57 (H)
O Mo L
M Ka
H
Wagner et al. 1990:252 (Ni, K, O, Mo, L, M, Ka, H)
Asteraceae
Acanthospermum australe (Loefl.) Kuntze
nat
Acanthospermum hispidum DC.
nat
Achillea millefolium L.
nat
Adenostemma viscosum J.R.Forst. & G.Forst.
ind
Ageratina adenophora (Spreng.) R.M.King & H.Rob.
nat
Ageratina riparia (Regel) R.M.King & H.Rob.
nat
Ageratum conyzoides L.
nat
Ageratum houstonianum Mill.
nat
Ambrosia artemisiifolia L.
nat
Anthemis cotula L.
nat
Arctium lappa L.
nat
Ni
K
Mo
K
Le Ni
Wagner et al. 1990:252 (Mo)
H
Wagner et al. 1990:252 (H); Staples & Herbst 2005:150 (KEY), 166
(DESCR)
H
Wagner et al. 1990:254 (K, O, Mo, L, M, H [as Adenostemma lavenia]);
King & Robinson 1987:60/Herbst & Wagner 1999:14 (NOM: A. lavenia
misapplied)
O Mo L
M
O Mo L
M
K
O Mo L
M
H
Wagner et al. 1990:255 (O, Mo, M, H); Herbst & Wagner 1996:9 (K);
Oppenheimer & Bartlett 2002:3 (L)
K
O Mo L
M Ka
H
Wagner et al. 1990:255 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:20 (Le)
K
O
M
H
Wagner et al. 1990:256 (K, O, H); Oppenheimer & Bartlett 2000:2 (EM);
Staples & Herbst 2005:148 (KEY), 153 (DESCR)
O Mo
M
H
Wagner et al. 1990:256 (O, Mo, M, H)
M
H
Wagner et al. 1990:258 (K, L, M, H)
H
Wagner et al. 1990:258 (O, L, H); Staples & Herbst 2005:148 (KEY), 167
(DESCR)
K
L
O
L
Wagner et al. 1990:254 (O, Mo, L, M)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 14
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Arctotheca calendula (L.) Levyns
nat
M
Starr & Starr 2011:23 (NEWNAT/EM, DESCR)
Argyroxiphium caliginis C.N.Forbes
end
M
Wagner et al. 1990:259 (WM)
Argyroxiphium grayanum (Hillebr.) O.Deg.
end
M
Wagner et al. 1990:260 (EM, WM)
Argyroxiphium kauense (Rock & M.Neal) O.Deg. &
I.Deg.
end
Argyroxiphium sandwicense DC. ssp. macrocephalum
(A.Gray) Meyrat
end
Argyroxiphium sandwicense DC. ssp. sandwicense
end
Argyroxiphium virescens Hillebr.
end
Artemisia australis Less.
end
Artemisia kauaiensis (Skottsb.) Skottsb.
end
Artemisia mauiensis (A.Gray) Skottsb.
end
Artemisia vulgaris L.
nat
Baltimora recta L.
nat
Bellis perennis L.
nat
H
M
Wagner et al. 1990:261 (EM)
H
M
Le Ni
K
O Mo L
M Ka
Wagner et al. 1990:261 (H)
Wagner et al. 1990:261 (EM)
H
K
Wagner et al. 1990:263 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:148 (KEY), 154 (DESCR); Wood & LeGrande 2006:20 (Le)
Wagner et al. 1990:263 (K)
M
K
Wagner et al. 1990:260 (H)
O
L
M
Wagner et al. 1990:265 (EM); Staples & Herbst 2005:148 (KEY), 154
(DESCR)
H
O
Wagner et al. 1990:265 (K, O, H); Wagner & Herbst 1995:15 (EM);
Oppenheimer 2004:9 (WM); Staples & Herbst 2005:148 (KEY), 154
(DESCR); Oppenheimer 2008:24 (L)
Wagner et al. 1990:267 (O) [Note: single coll. in 1981 from Makapu‘u
Lighthouse road]
K
H?
Wagner et al. 1990:267 (K, H? [uncertain location, no BISH voucher]);
Staples & Herbst 2005:149 (KEY), 167 (DESCR) [Note: BISH has single
cultivated Kaua‘i coll. in 1933 from Köke‘e]
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 15
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
K
O Mo L
nat
Bidens amplectens Sherff
end
O
Wagner et al. 1990:271 (O)
Bidens asymmetrica (H.Lév.) Sherff
end
O
Wagner et al. 1990:271 (O)
Bidens campylotheca Sch.Bip. ssp. campylotheca
end
Bidens campylotheca Sch.Bip. ssp. pentamera (Sherff)
Ganders & Nagata
end
M
Wagner et al. 1990:273 (EM); Lau 1995:10 (WM)
Bidens campylotheca Sch.Bip. ssp. waihoiensis
H.St.John
end
M
Wagner et al. 1990:273 (EM)
Bidens cervicata Sherff
end
Bidens conjuncta Sherff
end
Bidens cosmoides (A.Gray) Sherff
end
K
Bidens cynapiifolia Kunth
nat
K
Bidens forbesii Sherff ssp. forbesii
end
K
Wagner et al. 1990:275 (K)
Bidens forbesii Sherff ssp. kahiliensis Ganders & Nagata
end
K
Wagner et al. 1990:275 (K)
Bidens gardneri Baker
nat
Bidens hawaiensis A.Gray
end
H
Wagner et al. 1990:275 (H)
Bidens hillebrandiana (Drake) O.Deg. ssp.
hillebrandiana
end
H
Wagner et al. 1990:276 (H)
Ni
K
O
M Ka
L
H
Citation
Bidens alba (L.) DC. var. radiata (Sch.Bip.) Ballard ex
Melchert
K
O Mo L
M Ka H
H
O
Wagner et al. 1990:270 (K, O, M, Ka, H); Hughes 1995:2 (Mo);
Oppenheimer 2007:20 (L)
Wagner et al. 1990:273 (O, L, H); Lorence et al. 1995:22 (K)
Wagner et al. 1990:273 (Ni, K, O)
M
Wagner et al. 1990:273 (WM)
Wagner et al. 1990:274 (K)
O Mo L
M
H
Mo
Wagner et al. 1990:274 (O, Mo, L, M, H); Lorence et al. 1995:23 (K)
Wagner et al. 1990:275 (Mo)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 16
Scientific Name
Dicot
Asteraceae
Bidens hillebrandiana (Drake) O.Deg. ssp. polycephala
Nagata & Ganders
Bidens laevis (L.) Britton, Sterns & Poggenb.
Status
Kl Le Ni K
O Mo L
end
Mo
?
Bidens macrocarpa (A.Gray) Sherff
end
Bidens mauiensis (A.Gray) Sherff
end
Bidens menziesii (A.Gray) Sherff ssp. filiformis (Sherff)
Ganders & Nagata
end
Bidens menziesii (A.Gray) Sherff ssp. menziesii
end
Bidens micrantha Gaudich. ssp. ctenophylla (Sherff)
Nagata & Ganders
end
Bidens micrantha Gaudich. ssp. kalealaha Nagata &
Ganders
end
Bidens micrantha Gaudich. ssp. micrantha
end
Bidens molokaiensis (Hillebr.) Sherff
end
Bidens pilosa L.
nat
Bidens populifolia Sherff
end
Bidens sandvicensis Less. ssp. confusa Nagata &
Ganders
end
K
Bidens sandvicensis Less. ssp. sandvicensis
end
K
Bidens torta Sherff
end
M Ka H
M
Wagner et al. 1990:276 (Mo, M)
O? Mo?
Wagner et al. 1990:268 [Note: vouchers at BISH from O, Mo, but not since
1912]
O
Wagner et al. 1990:276 (O)
L
M Ka
Wagner et al. 1990:276 (L, M, Ka)
H
Mo
M
L
O Mo L
O
Wagner et al. 1990:279 (H)
M
Wagner et al. 1990:279 (L, EM, WM)
M
Wagner et al. 1990:279 (EM, WM)
O Mo
K
Wagner et al. 1990:278 (H); Staples & Herbst 2005:150 (KEY), 167
(DESCR)
Wagner et al. 1990:278 (Mo, WM)
H
Ni
Citation
Wagner et al. 1990:279 (O, Mo)
M Ka
H
Wagner et al. 1990:279 (Ni, K, O, Mo, L, M, Ka, H)
Wagner et al. 1990:281 (O)
Wagner et al. 1990:281 (K)
O
Wagner et al. 1990:281 (K, O)
O
Wagner et al. 1990:281 (O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 17
Scientific Name
Dicot
Asteraceae
Status
Bidens valida Sherff
end
Bidens wiebkei Sherff
end
Blumea laciniata (Roxb.) DC.
nat
Blumea sessiliflora Decne.
nat
Calyptocarpus vialis Less.
nat
Carduus pycnocephalus L.
nat
Centaurea cyanus L.
nat
Centaurea maculosa Lam.
Kl Le Ni K
O Mo L
M Ka H
K
Wagner et al. 1990:282 (K)
Mo
Wagner et al. 1990:282 (Mo)
O
H
M
K
O Mo L
M Ka
L
?
Wagner et al. 1990:283 (O, H)
Wagner et al. 1990:283 (M) [Note: known from single coll. in 1920, no
specific locality]
H
M
Ni
Citation
Wagner et al. 1990:284 (K, O, Mo, L, M); Oppenheimer 2006:10 (Ka);
Oppenheimer 2007:20 (H)
Wagner et al. 1990:284 (EM)
H
Wagner & Herbst 1995:15 (NEWNAT/L); Staples et al. 2003:8 (H); Staples
& Herbst 2005:148 (KEY), 168 (DESCR)
H?
Wagner et al. 1990:286 (H? [single coll. in 1965 from Kona], DESCR)
Centaurea melitensis L.
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:286 (Ni, K, O, Mo, L, M, Ka, H)
Centratherum punctatum Cass. ssp. punctatum
nat
K
O Mo L
M
H
Lorence et al. 1995:23 (NEWENAT/K, DESCR); Oppenheimer 2003:6 (H);
Starr et al. 2004:20 (EM); Staples & Herbst 2005:148 (KEY), 168 (DESCR);
Starr et al. 2006:32 (Mo); Starr et al. 2010:62 (L); Lau & Frohlich 2012:9 (O)
Chromolaena odorata (L.) R.M.King & H.Rob.
nat
Cirsium vulgare (Savi) Ten.
nat
Le?
K
O Mo L
M Ka
H
Wagner et al. 1990:287 (K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:20 (Le? [observed but unvouchered])
Conyza bonariensis (L.) Cronquist
nat
Le? Ni
K
O Mo L
M Ka
H
Wagner et al. 1990:288 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:20 (Le? [observed but unvouchered])
Conyza canadensis (L.) Cronquist var. canadensis
nat
M
H
Wagner et al. 1990:288 (M, H)
O
Frohlich & Lau 2012:30 (NEWNAT/O, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 18
Scientific Name
Dicot
Asteraceae
Status
Conyza canadensis (L.) Cronquist var. pusilla (Nutt.)
Cronquist
nat
Coreopsis lanceolata L.
nat
Coreopsis tinctoria Nutt.
?
Cosmos caudatus Kunth
?
Kl Le Ni K
Ni
K
O Mo L
O Mo L
L
M Ka H
Citation
M Ka
H
Wagner et al. 1990:288 (Ni, K, O, Mo, L, M, H); Starr et al. 2006:32 (Ka)
M
H
Wagner et al. 1990:289 (L, H); Meidell et al. 1997:17 (WM); Staples &
Herbst 2005:150, 151 (KEY), 156 (DESCR)
O?
Wagner et al. 1990:289 (O? [single coll. in 1916 from Schofield Barracks],
DESCR)
H
Wagner et al. 1990:289 (O, EM, H); Lorence et al. 1995:23 (K);
Oppenheimer 2003:6 (WM); Oppenheimer 2011:6 (L)
nat
K
O
Crassocephalum crepidioides (Benth.) S.Moore
nat
K
O Mo L
Crepis capillaris (L.) Wallr.
nat
H
Wagner et al. 1990:291 (H)
Crepis vesicaria L. ssp. taraxacifolia (Thuill.) Thuill.
nat
H
Staples et al. 2002:5 (NEWNAT/H)
Cyanthillium cinereum (L.) H.Rob.
nat
Delairea odorata Lem.
nat
Dubautia arborea (A.Gray) D.D.Keck
end
Dubautia carrii B.G.Baldwin & Friar
end
K
M
Wagner et al. 1990:242 (H? [Hillebrand (1888) reported it as naturalized, not
vouchered since], DESCR)
Cotula australis (Sieber ex Spreng.) Hook.f.
Ni
L
H?
M Ka? H
Wagner et al. 1990:291 (K, O, Mo, M, H); Nagata 1995:11 (L); Starr & Starr
2011:24 (Ka); Wagner et al. 2012:10 (Ka record adventive)
O Mo L
M Ka
H
Wagner et al. 1990:373 (Ni, K, O, Mo, L, M, Ka, H [as Vernonia cinerea var.
parviflora]); Wagner et al. 1997:52 (NOM:Syn. V. cinerea var. parviflora)
O
M
H
Wagner et al. 1990:356 (M, H [as Senecio mikanioides]); Jeffrey
1986:933/Herbst & Wagner 1999:14 (NOM:Syn. S. mikanioides); Staples &
Herbst 2005:148 (KEY), 168 (DESCR); Starr et al. 2010:62 (L); Lau &
Frohlich 2012:9 (O)
H
Wagner et al. 1990:295 (H)
L
Mo
Wagner et al. 1990:302 (Mo, WM [as Dubautia linearis subsp. opposita]);
Baldwin & Friar 2010:2 (SPNOV/Mo, NOM:Syn. D. linearis subsp. opposita,
DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 19
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Dubautia ciliolata (DC.) D.D.Keck ssp. ciliolata
end
H
Wagner et al. 1990:297 (H)
Dubautia ciliolata (DC.) D.D.Keck ssp. glutinosa
G.D.Carr
end
H
Wagner et al. 1990:297 (H)
Dubautia hanaulaensis B.G.Baldwin
end
Dubautia herbstobatae G.D.Carr
end
Dubautia imbricata H.St.John & G.D.Carr ssp. acronaea
G.D.Carr
end
K
Wagner et al. 1990:298 (K); Carr 1999:146 (KEY)
Dubautia imbricata H.St.John & G.D.Carr ssp. imbricata
end
K
Wagner et al. 1990:298 (K); Carr 1999:146 (KEY)
Dubautia kalalauensis B.G.Baldwin & G.D.Carr
end
K
Baldwin & Carr 2005:259 (SPNOV/K, DESCR)
Dubautia kenwoodii G.D.Carr
end
K
Carr 1998:8 (SPNOV/K, DESCR); Herbst & Wagner 1999:14 (DESCR);
Carr 1999:148 (KEY); Wood 2007:14 (EXTINCT?)
Dubautia knudsenii Hillebr. ssp. filiformis G.D.Carr
end
K
Wagner et al. 1990:298 (K); Carr 1999:148, 149 (KEY)
Dubautia knudsenii Hillebr. ssp. knudsenii
end
K
Wagner et al. 1990:299 (K); Carr 1999:148 (KEY)
Dubautia knudsenii Hillebr. ssp. nagatae (H.St.John)
G.D.Carr
end
K
Wagner et al. 1990:299 (K); Carr 1999:148, 149 (KEY)
Dubautia laevigata A.Gray
end
K
Wagner et al. 1990:299 (K); Carr 1999:150 (KEY)
Dubautia latifolia (A.Gray) D.D.Keck
end
K
Wagner et al. 1990:299 (K); Carr 1999:151 (KEY)
Dubautia laxa Hook. & Arn. ssp. bryanii (Sherff)
G.D.Carr
end
Dubautia laxa Hook. & Arn. ssp. hirsuta (Hillebr.)
G.D.Carr
end
M
O
Wagner et al. 1990:297 (O)
O
K
O
Baldwin & Friar 2010:5 (SPNOV/WM, DESCR)
Wagner et al. 1990:301 (O)
L
Wagner et al. 1990:301 (K, O, L); Carr 1999:152 (KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 20
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
K
O Mo L
O Mo
M Ka H
M
Citation
Dubautia laxa Hook. & Arn. ssp. laxa
end
Wagner et al. 1990:301 (O, Mo, M); Wood 2006:16 (K)
Dubautia laxa Hook. & Arn. ssp. pseudoplantaginea
(Skottsb.) G.D.Carr
end
Dubautia linearis (Gaudich.) D.D.Keck ssp. hillebrandii
(H.Mann) G.D.Carr
end
Dubautia linearis (Gaudich.) D.D.Keck ssp. linearis
end
Dubautia menziesii (A.Gray) D.D.Keck
end
Dubautia microcephala Skottsb.
end
K
Wagner et al. 1990:302 (K); Carr 1999:153 (KEY)
Dubautia paleata A.Gray
end
K
Wagner et al. 1990:303 (K); Carr 1999:154 (KEY)
Dubautia pauciflorula H.St.John & G.D.Carr
end
K
Wagner et al. 1990:303 (K); Carr 1999:154 (KEY)
Dubautia plantaginea Gaudich. ssp. humilis G.D.Carr
end
Dubautia plantaginea Gaudich. ssp. magnifolia (Sherff)
G.D.Carr
end
Dubautia plantaginea Gaudich. ssp. plantaginea
end
Dubautia platyphylla (A.Gray) D.D.Keck
end
Dubautia raillardioides Hillebr.
end
Dubautia reticulata (Sherff) D.D.Keck
end
Dubautia scabra (DC.) D.D.Keck ssp. leiophylla
(A.Gray) G.D.Carr
end
Dubautia scabra (DC.) D.D.Keck ssp. scabra
end
O
Wagner et al. 1990:301 (O)
H
L
Wagner et al. 1990:302 (H)
M
Wagner et al. 1990:302 (L, M)
M
Wagner et al. 1990:302 (EM)
M
Wagner et al. 1990:304 (WM)
K
Wagner et al. 1990:304 (K); Carr 1999:155 (KEY)
O Mo L
M
H
M
Wagner et al. 1990:304 (O, Mo, L, M, H)
Wagner et al. 1990:305 (M)
K
Wagner et al. 1990:305 (K); Carr 1999:155 (KEY)
M
Mo L
Wagner et al. 1990:305 (M)
M
H
Wagner et al. 1990:307 (Mo, L, M, H)
M
H
Wagner et al. 1990:308 (EM, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 21
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Dubautia sherffiana Fosberg
end
Wagner et al. 1990:308 (O)
Dubautia syndetica G.D.Carr & Lorence
end
K
Carr & Lorence 1998:4 (SPNOV/K, DESCR); Herbst & Wagner 1999:14
(DESCR); Carr 1999:156 (KEY)
Dubautia waialealae Rock
end
K
Wagner et al. 1990:308 (K); Carr 1999:156 (KEY)
Dubautia waianapanapaensis G.D.Carr
end
Dyssodia tenuiloba (DC.) B.L.Rob.
nat
Eclipta prostrata (L.) L.
nat
Elephantopus mollis Kunth
M
Wagner et al. 1990:297 (EM [syn. of Dubautia dolosa]); Carr
1995:1580/Wagner & Herbst 2003:7 (RESURRECT/EM, NOM:Syn. D.
dolosa)
Starr et al. 2002:18 (NEWNAT/O, EM, WM, DESCR); Staples & Herbst
2005:150 (KEY), 168 (DESCR); Starr et al. 2006:32 (Mo, Ka); Lorence &
Flynn 2006:2 (K); Starr et al. 2010:62 (L)
K
O Mo L
M Ka
K
O Mo L
M
H
Wagner et al. 1990:309 (Ni, K, O, Mo, M, H [as Eclipta alba]); Wagner &
Herbst 1995:16 (NOM:Syn. E. alba); Oppenheimer 2003:7 (L)
nat
K
O Mo L
M
H
Wagner et al. 1990:310 (K, O, Mo, M, H); Oppenheimer & Bartlett 2002:4
(L)
Emilia coccinea (Sims) G.Don
nat
K
O
Emilia fosbergii Nicolson
nat
K
O Mo L
M Ka
Emilia sonchifolia (L.) DC. var. javanica (Burm.f.)
Mattf.
nat
K
O
M
Emilia sonchifolia (L.) DC. var. sonchifolia
nat
K
O
Encelia farinosa A.Gray
nat
Erechtites hieracifolia (L.) Raf. ex DC.
nat
K
O Mo L
M
H
Wagner et al. 1990:313 (K, O, Mo, L, M, H)
Erechtites valerianifolia (Wolf) DC.
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:314 (K, O, Mo, L, M, H); Staples et al. 2003:8 (Ka)
Ni
Le Ni
L
L
Wagner et al. 1990:312 (O, L); Imada et al. 2000:10 (K)
M
H
Wagner et al. 1990:312 (Ni, K, O, Mo, L, M, Ka, H)
Wagner et al. 1990:312 (K, O); Wagner et al. 1997:52 (EM); Oppenheimer
2008:24 (L)
H
M
Wagner et al. 1990:312 (K, O, M); Oppenheimer & Bartlett 2002:4 (H)
Wagner et al. 1990:313 (EM, WM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 22
Scientific Name
Dicot
Asteraceae
Erigeron annuus (L.) Pers.
Status
Kl Le Ni K
O Mo L
M Ka H
?
Citation
H?
Wagner et al. 1990:314 (H? [single coll. in 1936 from Hämäkua])
Erigeron bellioides DC.
nat
K
O Mo L
M
H
Wagner et al. 1990:314 (O, M); Nagata 1995:11 (O); Oppenheimer & Bartlett
2000:2 (WM); Oppenheimer 2003:7 (H); Staples et al. 2003:8 (Mo); Starr et
al. 2004:21 (K); Oppenheimer 2008:24 (L)
Erigeron karvinskianus DC.
nat
K
O Mo
M
H
Wagner et al. 1990:315 (K, O, M, H); Hughes 1995:2 (Mo); Staples & Herbst
2005:150 (KEY), 168 (DESCR)
Euchiton gymnocephalus (DC.) Holub
nat
H
Nesom 2002:516, 518 (NEWNAT/H, KEY)
Euchiton japonicus (Thunb.) Anderb.
nat
M
H
Wagner et al. 1997:54 (NEWNAT/H, KEY); Herbst & Wagner 1999:14 (EM,
H); Staples et al. 2002:5 (EM); Nesom 2002:518 (KEY)
Euchiton sphaericus (Willd.) Anderb.
nat
M
H
Wagner et al. 1990:320 (Mo, L, M, H [as Gnaphalium japonicum]); Wagner
et al. 1997:54 (NOM: G. japonicum misapplied, KEY); Nesom 2002:516,
519 (KEY)
Eupatorium capillifolium (Lam.) Small
nat
H
Staples et al. 2003:9 (NEWNAT/H, DESCR)
Fitchia speciosa Cheeseman
nat
Flaveria trinervia (Spreng.) C.Mohr
nat
Florestina tripteris DC.
nat
Gaillardia pulchella Foug.
nat
K
Galinsoga parviflora Cav.
nat
K
Galinsoga quadriradiata Ruiz & Pav.
nat
Gamochaeta americana (Mill.) Weddell
nat
Mo L
O
K
Wagner et al. 1990:317 (O)
O
M Ka
Wagner et al. 1990:317 (O); Lorence et al. 1995:24 (K); Herbst & Wagner
1996:9 (Ka); Oppenheimer 2003:7 (EM, WM)
M
Oppenheimer & Bartlett 2000:2 (NEWNAT/WM, DESCR)
O Mo
M
Wagner et al. 1990:318 (O, M); Lorence et al. 1995:24 (K, Mo)
O Mo L
M Ka
H
Wagner et al. 1990:318 (K, O, M, Ka, H); Oppenheimer 2008:24 (L);
Oppenheimer 2011:6 (Mo)
O Mo
M
H
Wagner et al. 1990:320 (O, Mo, M, H)
M
Alford 2012:2, 5 (NEWNAT/EM, KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 23
Scientific Name
Dicot
Asteraceae
Status
Gamochaeta argyrinea G.L.Nesom
nat
Gamochaeta pensylvanica (Willd.) Cabrera
nat
Gamochaeta purpurea (L.) Cabrera
Kl Le Ni K
O Mo L
M Ka H
Citation
O
M
H
Alford 2012:3, 5 (NEWNAT/O, EM, WM, H, KEY)
O Mo L
M Ka
H
Wagner et al. 1990:321 (as Gnaphalium peregrinum, syn. of G. purpureum);
Alford 2012:2, 3, 4 (NEWNAT/Le, K, O, Mo, L, EM, WM, Ka, H, KEY,
resurrected from Gamochaeta purpurea)
nat
O Mo
M
H
Wagner et al. 1990:321 (K, O, Mo, L, M, Ka, H [as Gnaphalium
purpureum]); Wagner et al. 1997:54 (NOM:Syn. G. purpureum, KEY); Wood
& LeGrande 2006:20 (Le); Alford 2012:2, 5 (NOM: Gnaphalium peregrinum
[=Gamochaeta pensylvanica] resurrected from synonymy, remove Le, K, L,
Ka from distibution, KEY)
Gamochaeta simplicicaulis (Willd. ex Spreng.) Cabrera
nat
Mo
Helianthus annuus L.
nat
H
Wagner et al. 1990:322 (H); Staples & Herbst 2005:150 (KEY), 161
(DESCR)
Helichrysum foetidum (L.) Cass.
nat
H
Wagner et al. 1990:323 (H)
Hesperomannia arborescens A.Gray
end
O Mo L
M
Wagner et al. 1990:325 (O, Mo, L); Herbarium Pacificum Staff 1999:3 (WM)
Hesperomannia arbuscula Hillebr.
end
O
M
Wagner et al. 1990:325 (O, WM)
Hesperomannia lydgatei C.N.Forbes
end
K
Heterotheca grandiflora Nutt.
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:326 (K, O, L, M, Ka, H); Oppenheimer 2010:33 (Mo)
Hypochoeris glabra L.
nat
Le
K
O Mo L
M Ka
H
Wagner et al. 1990:327 (K, O, Mo, M, Ka, H); Herbst & Wagner 1996:10
(Le); Staples et al. 2003:9 (L)
Hypochoeris radicata L.
nat
Le
K
O Mo L
M Ka
H
Wagner et al. 1990:327 (K, Mo, L, M, Ka, H); Staples et al. 2002:6 (O);
Wood & LeGrande 2006:20 (Le)
Le
K
Alford 2012:4 (NEWNAT/Mo, KEY)
Wagner et al. 1990:325 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 24
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
O Mo L
M Ka H
O?
Citation
Kalimeris indica (L.) Sch.Bip. ssp. indica
nat
K
Wagner et al. 1990:242 (O? [as Boltonia indica; reported as adventive in
Käne‘ohe]); Nagata 1995:11 (NEWNAT/K); Gu & Hoch 1997:785/Herbst &
Wagner 1999:14 (NOM:Syn. B. indica); Staples & Herbst 2005:150 (KEY),
169 (DESCR)
Keysseria erici (C.N.Forbes) Cabrera
end
K
Wagner et al. 1990:329 (K [as Lagenifera erici]); Nicolson 1996:341
(Lagenophora proposed as conserved genus name); Nesom 2000:58/Nesom
2001b:514 (transfer of Hawaiian taxa to Keysseria, NOM:Syn. Lagenifera
erici)
Keysseria helenae (C.N.Forbes & Lydgate) Cabrera
end
K
Wagner et al. 1990:330 (K [as Lagenifera helenae]); Nicolson 1996:341
(Lagenophora proposed as conserved genus name); Nesom 2000:58/Nesom
2001b:514 (transfer of Hawaiian taxa to Keysseria, NOM:Syn. Lagenifera
helenae)
Keysseria maviensis (H.Mann) Cabrera
end
Lactuca sativa L.
nat
Lapsana communis L.
nat
Leucanthemum Xsuperbum (J.W.Ingram) D.H.Kent
nat
Leucanthemum vulgare Lam.
nat
Mo
K
O Mo L
M
Wagner et al. 1990:330 (Mo, EM, WM [as Lagenifera maviensis]); Nicolson
1996:341 (Lagenophora proposed as conserved genus name); Nesom
2000:58/Nesom 2001b:514 (transfer of Hawaiian taxa to Keysseria,
NOM:Syn. Lagenifera maviensis)
M Ka
H
Wagner et al. 1990:327 (O, L, M, H [as Lactuca serriola]); Wagner & Herbst
1995:16 (K); Koopman et al. 2001 (NOM:Syn. L. serriola); Oppenheimer
2003:7 (Mo); Staples & Herbst 2005:149 (KEY), 162 (DESCR); Starr et al.
2006:33 (Ka)
M
H
Wagner et al. 1990:331 (M, H)
H
Staples & Herbst 2005:151 (KEY), 163 (DESCR); Pratt & Bio 2012:75
(NEWNAT/H)
H
Wagner et al. 1990:286 (M, H [as Chrysanthemum leucanthemum]); Soreng
& Cope 1991:162/Herbst & Wagner 1999:15 (NOM:Syn. C. leucanthemum);
Staples & Herbst 2005:151 (KEY), 163 (DESCR)
M
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 25
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
Ni
O Mo L
Lipochaeta connata (Gaudich.) DC. ssp. acris (Sherff)
W.L.Wagner & H.Rob.
end
Lipochaeta connata (Gaudich.) DC. ssp. connata
end
Lipochaeta degeneri Sherff
end
Mo
Lipochaeta heterophylla A.Gray
end
Mo L
Lipochaeta lobata (Gaudich.) DC. ssp. leptophylla
(O.Deg. & Sherff) W.L.Wagner & H.Rob.
end
Lipochaeta lobata (Gaudich.) DC. ssp. lobata
end
Lipochaeta rockii Sherff
end
Lipochaeta succulenta (Hook. & Arn.) DC.
end
Logfia gallica (L.) Coss. & Germ.
nat
Madia sativa Molina var. sativa
nat
Melanthera bryanii (Sherff) W.L.Wagner & H.Rob.
end
Melanthera fauriei (H.Lév.) W.L.Wagner & H.Rob.
end
M Ka H
K
Wagner et al. 1990:334 (Ni, K [as Lipochaeta connata var. acris]); Wagner &
Robinson 2002:545 (COMBNOV & STATNOV:Syn. L. connata var. acris)
K
M
Wagner et al. 1990:334 (K, WM [as Lipochaeta connata var. connata]);
Wagner & Robinson 2002:545 (NOM:Syn. L. connata var. connata)
Wagner et al. 1990:334 (Mo); Wagner & Robinson 2002:545
M
Wagner et al. 1990:336 (Mo, L, M); Wagner & Robinson 2002:545
O
Ni
Wagner et al. 1990:338 (O [as Lipochaeta lobata var. leptophylla]); Wagner
& Robinson 2002:547 (COMBNOV & STATNOV:Syn. L. lobata var.
leptophylla)
O
Mo
Ni
K
Citation
O Mo
M
Wagner et al. 1990:338 (Ni, O, WM [as Lipochaeta lobata var. lobata]);
Wagner & Robinson 2002:546 (NOM:Syn. L. lobata var. lobata)
M Ka H?
Wagner et al. 1990:340 (Mo, M, Ka, H? [apparent anomalous coll. from
1779; not vouchered since on Hawai‘i]); Wagner & Robinson 2002:547
M Ka
H
Wagner et al. 1990:341 (Ni, K, O, Mo, M, Ka, H); Wagner & Robinson
2002:549
H
Wagner et al. 1990:315 (H [as Filago gallica]); Wagner et al. 1997:54
(NOM:Syn. F. gallica)
M
Wagner et al. 1990:345 (EM)
Ka
K
Wagner et al. 1990:334 (Ka [as Lipochaeta bryanii]); Wagner & Robinson
2002:552 (COMBNOV:Syn. L. bryanii)
Wagner et al. 1990:336 (K [as Lipochaeta fauriei]); Wagner & Robinson
2002:552 (COMBNOV:Syn. L. fauriei)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 26
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
Ni
K
O Mo L
O Mo L
M Ka H
M Ka
H
Citation
Melanthera integrifolia (Nutt.) W.L.Wagner & H.Rob.
end
Melanthera kamolensis (O.Deg. & Sherff) W.L.Wagner
& H.Rob.
end
Melanthera lavarum (Gaudich.) W.L.Wagner & H.Rob.
end
Melanthera micrantha (Nutt.) W.L.Wagner & H.Rob.
ssp. exigua (O.Deg. & Sherff) W.L.Wagner & H.Rob.
end
K
Wagner et al. 1990:338 (K [as Lipochaeta micrantha var. exigua]); Wagner &
Robinson 2002:554 (COMBNOV & STATNOV:Syn. L. micrantha var.
exigua)
Melanthera micrantha (Nutt.) W.L.Wagner & H.Rob.
ssp. micrantha
end
K
Wagner et al. 1990:338 (K [as Lipochaeta micrantha var. micrantha]);
Wagner & Robinson 2002:554 (COMBNOV:Syn. L. micrantha var.
micrantha)
Melanthera perdita (Sherff) W.L.Wagner & H.Rob.
end
Melanthera populifolia (Sherff) W.L.Wagner & H.Rob.
end
Melanthera remyi (A.Gray) W.L.Wagner & H.Rob.
end
Melanthera subcordata (A.Gray) W.L.Wagner & H.Rob.
end
Melanthera tenuifolia (A.Gray) W.L.Wagner & H.Rob.
end
M
Mo L
M Ka
Wagner et al. 1990:336 (Ni, K, O, Mo, L, M, Ka, H [as Lipochaeta
integrifolia]); Wagner & Robinson 2002:552 (COMBNOV:Syn. L.
integrifolia); Staples & Herbst 2005:150 (KEY), 169 (DESCR)
Wagner et al. 1990:337 (EM [as Lipochaeta kamolensis]); Wagner &
Robinson 2002:552 (COMBNOV:Syn. L. kamolensis)
H
Ni
Wagner et al. 1990:337 (Mo, L, M, Ka, H [as Lipochaeta lavarum]); Wagner
& Robinson 2002:553 (COMBNOV:Syn. L. lavarum)
Wagner et al. 1990:340 (Ni [as Lipochaeta perdita]); Wagner & Robinson
2002:554 (COMBNOV:Syn. L. perdita)
L
Wagner et al. 1990:341 (as Lipochaeta populifolia, syn. of L. subcordata);
Wagner & Robinson 2002:554 (RESURRECT/L, COMBNOV:Syn. L.
populifolia)
O
Wagner et al. 1990:340 (O [as Lipochaeta remyi]); Wagner & Robinson
2002:554 (COMBNOV:Syn. L. remyi)
H
O
Wagner et al. 1990:341 (K, L, H [as Lipochaeta subcordata]); Wagner &
Robinson 2002:554 (COMBNOV:Syn. L. subcordata, remove K [spms.
reidentified as Melanthera fauriei], remove L [to resurrected Läna‘i endemic
Melanthera populifolia])
Wagner et al. 1990:343 (O [as Lipochaeta tenuifolia]); Wagner & Robinson
2002:554 (COMBNOV:Syn. L. tenuifolia)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 27
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
O Mo L
M Ka H
Melanthera tenuis (O.Deg. & Sherff) W.L.Wagner &
H.Rob.
end
O
Melanthera venosa (Sherff) W.L.Wagner & H.Rob.
end
Melanthera waimeaensis (H.St.John) W.L.Wagner &
H.Rob.
end
K
Montanoa hibiscifolia (Benth.) Standl.
nat
K
Osteospermum calendulaceum L.f.
nat
Palafoxia callosa (Nutt.) Torr. & A.Gray
nat
Parthenium hysterophorus L.
nat
Wagner et al. 1990:343 (O [as Lipochaeta tenuis]); Wagner & Robinson
2002:555 (COMBNOV:Syn. L. tenuis)
Pectis linifolia L. var. linifolia
nat
Perityle emoryi Torr.
nat
Petasites japonicus (Siebold & Zucc.) Maxim. var.
japonicus
nat
Picris hieracioides L.
nat
Pluchea Xfosbergii Cooperr. & Galang
nat
Le
K
O Mo
M Ka
Pluchea carolinensis (Jacq.) G.Don
nat
Le Ni
K
O Mo L
M Ka
H
K
Citation
Wagner et al. 1990:344 (H [as Lipochaeta venosa]); Wagner & Robinson
2002:555 (COMBNOV:Syn. L. venosa)
Wagner et al. 1990:344 (K [as Lipochaeta waimeaensis]); Wagner &
Robinson 2002:555 (COMBNOV:Syn. L. waimeaensis)
O
L
M Ka
Mo
M
O Mo
M
H
Wagner et al. 1990:345 (K, O, L, M, H); Staples & Herbst 2005:149 (KEY),
169 (DESCR); Starr & Starr 2011:24 (Ka)
H
Wagner et al. 1990:346 (H)
Wagner et al. 1990:346 (Mo); Oppenheimer & Bartlett 2000:2 (WM)
H
Wagner et al. 1990:347 (K, Mo, M, H); Wagner & Herbst 1995:16 (O)
H
Wagner et al. 1990:347 (H)
Ka
Mo L
Starr et al. 2010:62 (NEWNAT/Ka, DESCR)
H
Wagner et al. 1990:349 (H)
H
Wagner et al. 1990:350 (L, H); Oppenheimer 2008:24 (Mo)
Wagner et al. 1990:350 (K, O, Mo, M); Wood & LeGrande 2006:21 (Le);
Starr et al. 2006:33 (Ka)
H
Wagner et al. 1990:351 (Ni, K, O, Mo, L, M, Ka, H [as Pluchea
symphytifolia]); Khan & Jarvis 1989:661/Wagner & Herbst 1995:16 (NOM:
P. symphytifolia misapplied); Wood & LeGrande 2006:20 (Le)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 28
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
Le Ni
O Mo L
Pluchea indica (L.) Less.
nat
Pseudelephantopus spicatus (B.Juss. ex Aubl.) C.F.Baker
nat
Pseudognaphalium attenuatum (DC.) Anderb.
nat
Pseudognaphalium sandwicensium (Gaudich.) Anderb.
var. hawaiiense (O.Deg. & Sherff) W.L.Wagner
end
Pseudognaphalium sandwicensium (Gaudich.) Anderb.
var. kilaueanum (O.Deg. & Sherff) W.L.Wagner
end
Pseudognaphalium sandwicensium (Gaudich.) Anderb.
var. molokaiense (O.Deg. & Sherff) W.L.Wagner
end
Pseudognaphalium sandwicensium (Gaudich.) Anderb.
var. sandwicensium
end
Pseudogynoxys chenopodioides (Kunth) Cabrera
nat
Reichardia picroides (L.) Roth
nat
O Mo
Reichardia tingitana (L.) Roth
nat
O Mo
Remya kauaiensis Hillebr.
end
M Ka H
K
O Mo L
M Ka
H
Wagner et al. 1990:351 (Ni, K, O, Mo, L, M, Ka); Oppenheimer & Bartlett
2002:4 (H); Wood & LeGrande 2006:21 (Le)
K
O
M
H
Wagner et al. 1990:310 (K, O, M, H [as Elephantopus spicatus]); Herbst &
Wagner 1999:15 (NOM:Syn. E. spicatus)
H
Nesom 2001a:1186 (NEWNAT/H, DESCR)/Herbst et al. 2004:4
H
Wagner et al. 1990:321 (EM, H [as Gnaphalium sandwicensium var.
hawaiiense]); Wagner et al. 1997:54 (COMBNOV:Syn. G. sandwicensium
var. hawaiiense, KEY)
H
Wagner et al. 1990:321 (H [as Gnaphalium sandwicensium var.
kilaueanum]); Wagner et al. 1997:54 (COMBNOV:Syn. G. sandwicensium
var. kilaueanum, KEY)
M
Ni
Citation
K
O Mo L
M
O Mo L
M
K
Wagner et al. 1990:321 (Mo [as Gnaphalium sandwicensium var.
molokaiense]); Wagner et al. 1997:54 (Mo, L, WM, COMBNOV:Syn. G.
sandwicensium var. molokaiense, KEY)
H
Wagner et al. 1990:321 (Ni, K, O, Mo, L, M, H [as Gnaphalium
sandwicensium var. sandwicensium]); Wagner et al. 1997:54
(COMBNOV:Syn. G. sandwicensium var. sandwicensium, KEY); Staples &
Herbst 2005:149 (KEY), 170 (DESCR)
H
Wagner et al. 1990:355 [as Senecio confusus; escaping cultivation but not
naturalized]; Staples & Herbst 2005:150 (KEY), 163 (NOM:Syn. S.
confusus, DESCR); Parker & Parsons 2012a:57 (NEWNAT/H)
Wagner et al. 1990:352 (O); Shannon & Wagner 1996:13 (Mo)
H
Wagner et al. 1990:352 (O, Mo, H)
Wagner et al. 1990:353 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 29
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
Remya mauiensis Hillebr.
end
Remya montgomeryi W.L.Wagner & D.R.Herbst
end
K
Senecio madagascariensis Poir.
nat
K
Senecio sylvaticus L.
nat
Senecio vulgaris L.
nat
Sigesbeckia orientalis L.
nat
Solidago canadensis L. var. scabra Torr. & A.Gray
nat
Soliva sessilis Ruiz & Pav.
nat
Sonchus asper (L.) Hill
nat
Sonchus oleraceus L.
nat
Sphagneticola trilobata (L.) Pruski
nat
O Mo L
M Ka H
M
Kl Le Ni
Wagner et al. 1990:353 (WM)
Wagner et al. 1990:355 (K)
O
L
O Mo
Le
Citation
M Ka
H
Lorence et al. 1995:24 (NEWNAT/K, H, DESCR); Starr et al. 1999:11 (EM);
Oppenheimer & Bartlett 2002:4 (WM); Herbst et al. 2004:4 (O); Starr et al.
2006:33 (L, Ka); Starr & Starr 2012:88 (EM, highest elev record)
M
H
Wagner et al. 1990:356 (O, Mo, M, H)
M
H
Wagner et al. 1990:356 (EM, H)
K
O Mo L
M Ka
H
Wagner et al. 1990:357 (K, O, Mo, M, Ka, H); Wood 2007:14 (Le);
Oppenheimer 2007:20 (L)
K
O
M
H
Wagner et al. 1990:357 (K, O, M, H); Staples & Herbst 2005:150 (KEY [as
Solidago canadensis]), 170 (DESCR)
H
Wagner & Herbst 1995:16 (NEWNAT/H, DESCR)
M
H
Wagner et al. 1990:358 (EM, H)
K
O Mo L
M Ka
H
Wagner et al. 1990:358 (K, O, Mo, L, M, Ka, H); Herbarium Pacificum Staff
1996:3 (Ni); Wood & LeGrande 2006:21 (Le)
K
O Mo L
M
H
Wagner et al. 1990:373 (K, O [as Wedelia trilobata]); Wagner et al. 1997:55
(COMBNOV:Syn. W. trilobata); Imada et al. 2000:10 (H); Oppenheimer &
Bartlett 2002:4 (EM, WM); Staples & Herbst 2005:150 (KEY), 165
(DESCR); Oppenheimer 2007:20 (Mo, L)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 30
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
Symphyotrichum expansum (Poepp. ex Spreng.)
G.L.Nesom
nat
Synedrella nodiflora (L.) Gaertn.
nat
Tagetes erecta L.
nat
O
Tagetes minuta L.
nat
O?
Tanacetum parthenium (L.) Sch.Bip.
?
Taraxacum laevigatum (Willd.) DC.
?
Taraxacum officinale W.W.Weber ex F.H.Wigg.
nat
Tetramolopium arenarium (A.Gray) Hillebr. ssp.
arenarium var. arenarium
end
Tetramolopium arenarium (A.Gray) Hillebr. ssp.
arenarium var. confertum Sherff
end
Tetramolopium arenarium (A.Gray) Hillebr. ssp. laxum
Lowrey
end
Ni
O Mo L
M Ka H
Citation
K
O Mo
M
H
Wagner et al. 1990:266 (Ni, K, O, Mo, M, H [as Aster subulatus var.
sandwicensis]); Nesom 1994:292 (NOM:Syn. A. subulatus var. sandwicensis
= Symphyotrichum squamatum); Sundberg 2004:907 (COMBNOV: S.
subulatum var. parviflorum [S. squamatum misapplied]; Nesom 2005:2133
(NOM:Syn. S. subulatum var. parviflorum, Aster subulatus var.
sandwicensis; S. squamatum misapplied, non (Spreng.) Nesom)
K
O Mo L
M Ka
H
Wagner et al. 1990:360 (K, O, Mo, M, Ka, H); Oppenheimer 2007:21 (L)
Staples & Herbst 2005:150 (KEY), 165 (DESCR); Kennedy et al. 2010:20
(NEWNAT/O)
H
M?
Wagner et al. 1990:286 (EM? [as Chrysanthemum parthenium; reported in
1988 as escape at Science City, Haleakalä, but voucher not seen], DESCR);
Herbst & Wagner 1999:15 (NOM:Syn. C. parthenium)
O?
K
Wagner et al. 1990:360 (O? [single coll. from Makiki in 1956, perhaps
cultivated], H)
Wagner et al. 1990:361 (O? [single coll. in 1981 from Mänoa Valley, perhaps
naturalizing], DESCR)
O Mo L
M
H
Wagner et al. 1990:361 (K, O, L, M, H); Oppenheimer 2003:7 (Mo); Staples
& Herbst 2005:149 (KEY), 170 (DESCR)
M
H
Wagner et al. 1990:363 (EM, H)
H
Wagner et al. 1990:363 (H)
M
Wagner et al. 1990:363 (EM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 31
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
O Mo L
M Ka H
M
Citation
Tetramolopium capillare (Gaudich.) H.St.John
end
Wagner et al. 1990:363 (WM); Herbst & Wagner 1996:10
(REDISCOVER/WM)
Tetramolopium consanguineum (A.Gray) Hillebr. ssp.
consanguineum
end
Tetramolopium consanguineum (A.Gray) Hillebr. ssp.
leptophyllum (Sherff) Lowrey var. kauense Lowrey
end
H
Wagner et al. 1990:364 (H)
Tetramolopium consanguineum (A.Gray) Hillebr. ssp.
leptophyllum (Sherff) Lowrey var. leptophyllum
end
H
Wagner et al. 1990:364 (H)
Tetramolopium conyzoides (A.Gray) Hillebr.
end
H
Wagner et al. 1990:364 (Mo, L, M, H)
Tetramolopium filiforme Sherff var. filiforme
end
O
Wagner et al. 1990:365 (O)
Tetramolopium filiforme Sherff var. polyphyllum
(Sherff) Lowrey
end
O
Wagner et al. 1990:365 (O)
Tetramolopium humile (A.Gray) Hillebr. ssp. haleakalae
Lowrey
end
M
Tetramolopium humile (A.Gray) Hillebr. ssp. humile
var. humile
end
M
Tetramolopium humile (A.Gray) Hillebr. ssp. humile
var. sublaeve Sherff
end
Tetramolopium lepidotum (Less.) Sherff ssp.
arbusculum (A.Gray) Lowrey
end
Tetramolopium lepidotum (Less.) Sherff ssp. lepidotum
end
Tetramolopium remyi (A.Gray) Hillebr.
end
K?
Wagner et al. 1990:364 (K? [locality doubtful])
Mo L
M
M
O
L
L
Wagner et al. 1990:367 (EM)
H
Wagner et al. 1990:367 (EM, H)
H
Wagner et al. 1990:367 (H)
Wagner et al. 1990:367 (EM)
Wagner et al. 1990:367 (O, L)
M
Wagner et al. 1990:367 (L, WM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 32
Scientific Name
Dicot
Asteraceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Tetramolopium rockii Sherff var. calcisabulorum
(H.St.John) Lowrey
end
Mo
Wagner et al. 1990:368 (Mo)
Tetramolopium rockii Sherff var. rockii
end
Mo
Wagner et al. 1990:368 (Mo)
Tetramolopium sylvae Lowrey
ind
Mo
Tetramolopium tenerrimum (Less.) Nees
end
Tithonia diversifolia (Hemsl.) A.Gray
nat
Tragopogon porrifolius L.
nat
Tridax procumbens L.
nat
Verbesina encelioides (Cav.) Benth. & Hook.
nat
Vernonia elliptica DC.
nat
Wilkesia gymnoxiphium A.Gray
end
K
Wagner et al. 1990:375 (K)
Wilkesia hobdyi H.St.John
end
K
Wagner et al. 1990:376 (K)
Xanthium strumarium L. var. canadense (Mill.) Torr. &
A.Gray
nat
Youngia japonica (L.) DC.
nat
Zinnia palmeri A.Gray
nat
M
Wagner et al. 1990:368 (Mo, WM)
O
K
Le
Wagner et al. 1990:369 (O)
O
M
H
Wagner et al. 1990:369 (K, O, M, H); Staples & Herbst 2005:151 (KEY),
171 (DESCR)
H
Wagner et al. 1990:370 (H)
K
O Mo L
M Ka
H
Wagner et al. 1990:370 (K, O, Mo, L, M, Ka, H)
K
O Mo L
M Ka
H
Wagner et al. 1990:372 (K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:21 (Le)
O
Le? Ni
Staples & Herbst 2005:148 (KEY), 171 (DESCR); Lau & Frohlich 2012:9
(NEWNAT/O, DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:376 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:21 (Le? [observed but unvouchered])
K
O Mo L
M
H
Wagner et al. 1990:377 (K, O, Mo, L, M, H)
O
Wagner et al. 1990:379 (O [as Zinnia palmeri]); Staples et al. 2002:6
(NOM:Syn. Z. palmeri = Z. maritima); Wagner & Herbst 2003:11 (maintain
Z. palmeri as correct name for Hawaiian material) [Note: single coll. in 1980
as roadside weed, Koko Crater]
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 33
Scientific Name
Dicot
Asteraceae
Zinnia peruviana (L.) L.
Status
Kl Le Ni K
O Mo L
nat
Mo L
M Ka H
M Ka
H
Citation
Wagner et al. 1990:379 (L, M, Ka); Staples et al. 2002:6 (Mo, H)
Balsaminaceae
Impatiens sodeni Engl. & Warb. ex Engl.
nat
K
M
Wagner et al. 1990:380 (K [as Impatiens oliveri]); Grey-Wilson
1982:20/Herbst & Wagner 1999:15 (NOM:Syn. I. oliveri); Oppenheimer
2003:8 (EM); Staples & Herbst 2005:172 (KEY), 173 (DESCR)
Impatiens walleriana Hook.f
nat
K
O Mo L
M
H
Wagner et al. 1990:380 (K, O, Mo, L, M, H [as Impatiens wallerana (spelling
variant)]); Staples & Herbst 2005:172 (KEY, DESCR)
Anredera cordifolia (Ten.) Steenis
nat
K
O Mo L
M
H
Wagner et al. 1990:381 (K, O, Mo, M, H); Nagata 1995:11 (L); Staples &
Herbst 2005:174 (DESCR)
Basella alba L.
nat
Basellaceae
O
Nagata 1988:80/Wagner et al. 1990:381 (O? [cultivated, periodically
escaping]); Nagata 1995:11 (NEWNAT/O); Staples & Herbst 2005:173
(DESCR)
Bataceae
Batis maritima L.
nat
Ni
K
O Mo L
M Ka
H
Wagner et al. 1990:382 (Ni, K, O, Mo, L, M, Ka, H)
H
Lorence et al. 1995:25 (NEWNAT/K, H, DESCR)
H
Wagner et al. 1990:383 (H [as Begonia foliosa var. miniata]); Staples &
Herbst 2005:175 (KEY), 176 (DESCR); Imada 2007:35 (O, NOM: B. foliosa
var. miniata misapplied)
Begoniaceae
Begonia cucullata Willd.
nat
K
Begonia foliosa Kunth
nat
Begonia fusca Liebm.
nat
K
O
Begonia hirtella Link
nat
K
O
O
Miller 2003 (NEWNAT/K, O, DESCR); Staples & Herbst 2005:176 (KEY,
DESCR)
M
H
Wagner et al. 1990:383 (O, EM, H); Lorence et al. 1995:25 (K);
Oppenheimer 2004:9 (WM); Staples & Herbst 2005:175 (KEY), 178
(DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 34
Scientific Name
Dicot
Begoniaceae
Begonia nelumbiifolia Schltdl. & Cham.
Status
Kl Le Ni K
O Mo L
?
O?
Begonia vitifolia Schott
nat
O
Hillebrandia sandwicensis Oliv.
end
K
M Ka H
Citation
Staples & Herbst 2005:175 (KEY), 177 (DESCR); Daehler & Baker 2006:14
(O? [adventive])
H
O Mo
M
Wagner et al. 1990:384 (H [as Begonia reniformis]); Herbst & Wagner
1996:10 (O); Staples et al. 2002:6 (NOM: B. reniformis misapplied); Staples
& Herbst 2005:176 (KEY), 177 (DESCR); Wagner et al. 2012:17 (recognized
as B. reniformis in US checklist)
Wagner et al. 1990:384 (K, O, Mo, M)
Betulaceae
Alnus nepalensis D.Don
nat
Mo
Wagner et al. 1990:386 (Mo? [possibly naturalizing]); Wysong et al. 2007:3
(NEWNAT/Mo)
Bignoniaceae
Catalpa longissima (Jacq.) Dum.-Cours.
nat
O
Jacaranda mimosifolia D.Don
nat
O
L
M
H
Wagner et al. 1990:387 (O, M, H); Staples & Herbst 2005:181 (KEY), 182
(DESCR); Oppenheimer 2008:24 (L)
Macfadyena unguis-cati (L.) A.H.Gentry
nat
O
L
M
H
Wagner et al. 1990:388 (K, O, L); Imada et al. 2000:10 (H); Oppenheimer &
Bartlett 2000:2 (WM); Starr et al. 2002:18 (H); Staples & Herbst 2005:181
(KEY), 183 (DESCR)
Mansoa hymenaea (DC.) A.H.Gentry
nat
Podranea ricasoliana (Tanfani) Sprague
nat
K
Pyrostegia venusta (Ker Gawl.) Miers
nat
K
K
Staples & Herbst 2005:181 (KEY, DESCR); Lau & Frohlich 2012:9
(NEWNAT/O, DESCR)
O
Staples & Herbst 2005:181 (KEY), 184 (DESCR); Lau & Frohlich 2012:10
(NEWNAT/O, DESCR)
M
O
H
Starr et al. 2004:21 (NEWNAT/EM, DESCR); Staples & Herbst 2005:181
(KEY), 184 (DESCR); Frohlich & Lau 2012:31 (K); Parker & Parsons
2012a:57 (H)
Staples & Herbst 2005:181 (KEY), 184 (DESCR); Frohlich & Lau 2012:31
(NEWNAT/K, O, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 35
Scientific Name
Dicot
Bignoniaceae
Radermachera sinica (Hance) Hemsl.
Status
Kl Le Ni K
?
O Mo L
M Ka H
O?
Spathodea campanulata P.Beauv.
nat
K
Tabebuia aurea (Silva Manso) Benth. & Hook.f. ex
S.Moore
nat
O
Tabebuia heterophylla (DC.) Britton
nat
O
Tabebuia pallida (Lindl.) Miers
nat
Tabebuia rosea (Bertol.) DC.
K
Citation
Frohlich & Lau 2008:4 (NEWNAT/O, DESCR); Wagner et al. 2012:18 (O
record adventive)
O Mo L
M
H
Wagner et al. 1990:388 (K, O, Mo, M, H); Staples & Herbst 2005:181
(KEY), 185 (DESCR); Oppenheimer 2007:21 (L)
Staples & Herbst 2005:181, 187 (KEY), 187 (DESCR); Frohlich & Lau
2010:5 (NEWNAT/O, DESCR)
M
H
Oppenheimer 2003:8 (NEWNAT/WM); Oppenheimer 2004:10 (O); Staples
& Herbst 2005:181, 187 (KEY), 188 (DESCR); Parker & Parsons 2012b:66
(H)
O
Staples & Herbst 2005:181, 187 (KEY), 189 (DESCR); Frohlich & Lau
2012:31 (NEWNAT/K, O, DESCR)
nat
O
Staples & Herbst 2005:181, 187 (KEY), 189 (DESCR); Frohlich & Lau
2010:6 (NEWNAT/O, DESCR)
Tecoma castanifolia (D.Don) Melch.
nat
O
Herbarium Pacificum Staff 1998:9 (NEWNAT/O); Staples & Herbst
2005:181, 189 (KEY), 190 (DESCR)
Tecoma stans (L.) Juss. ex Kunth
nat
O
M
O Mo
M
H?
Wagner et al. 1990:389 (O, M, H? [reportedly naturalizing at Ocean View
Estates]); Staples & Herbst 2005:181, 189 (KEY), 190 (DESCR)
Bixaceae
Bixa orellana L.
nat
K
Wagner et al. 1990:390 (K, O, Mo, M); Staples & Herbst 2005:191 (DESCR)
Boraginaceae
Amsinckia intermedia Fisch. & C.A.Mey.
nat
Bothriospermum tenellum (Hornem.) Fisch. & C.A.Mey.
nat
O
H
Wagner et al. 1990:391 (H) [Note: single coll. in 1944 from Pu‘u ‘Ula‘ula,
Mauna Loa]
H
Wagner et al. 1990:392 (O, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 36
Scientific Name
Dicot
Boraginaceae
Status
Kl Le Ni K
O Mo L
K
nat
Cordia alliodora (Ruiz & Pav.) Oken
nat
O
Lau & Frohlich 2012:10 (NEWNAT/O, DESCR)
Cordia dichotoma G.Forst.
nat
O
Herbarium Pacificum Staff 1996:3 (NEWNAT/O, DESCR); Staples & Herbst
2005:197 (KEY), 198 (DESCR)
Cordia lutea Lam.
nat
Cordia subcordata Lam.
ind
Cynoglossum amabile Stapf & J.R.Drumm.
nat
Heliotropium amplexicaule Vahl
nat
Heliotropium anomalum Hook. & Arn. var. argenteum
A.Gray
end
Le Ni
Heliotropium curassavicum L.
ind
Kl Le Ni
Heliotropium procumbens Mill. var. depressum (Cham.)
Fosberg
nat
Myosotis azorica H.C.Watson
nat
Myosotis discolor Pers.
nat
Tournefortia argentea L.f.
nat
Ni
M
Citation
Carmona retusa (Vahl) Masam.
Ni
Mo
M Ka H
Lorence et al. 1995:25 (NEWNAT/K, WM, DESCR); Starr et al. 2003:25
(EM); Staples & Herbst 2005:197 (KEY, DESCR); Starr et al. 2006:33 (Mo)
H
Parker & Parsons 2012a:58 (NEWNAT/H)
M
H
Wagner et al. 1990:392 (Ni, K, O, L, M, H); Burney et al. 2001:633/Wagner
& Herbst 2003:12 (status change to "ind"); Staples & Herbst 2005:197
(KEY), 198 (DESCR); Oppenheimer 2007:21 (Mo)
M
H
Wagner et al. 1990:394 (H); Staples & Herbst 2005:197 (KEY), 199
(DESCR); Oppenheimer 2010:34 (EM); Starr & Starr 2011:24 (EM)
O Mo
M
H
Wagner et al. 1990:395 (O, Mo, M, H)
K
O Mo
M
H
Wagner et al. 1990:395 (Ni, K, O, Mo, M, H); Staples & Herbst 2005:197
(KEY), 199 (DESCR); Wood & LeGrande 2006:21 (Le)
K
O Mo L
M Ka
H
Wagner et al. 1990:395 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:21 (Le)
K
O
M
Wagner et al. 1990:396 (O, M); Herbst & Wagner 1999:15 (K)
M
Staples & Herbst 2005:197 (KEY), 199 (DESCR); Starr et al. 2008:44
(NEWNAT/EM, DESCR)
K
K
O Mo L
O Mo L
M
H
Wagner et al. 1990:396 (H)
H
Wagner et al. 1990:398 (Ni, K, O, Mo, L, M, H); Staples & Herbst 2005:197
(KEY), 199 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 37
Scientific Name
Dicot
Brassicaceae
Status
Kl Le Ni K
Brassica juncea (L.) Czern.
nat
Brassica nigra (L.) W.D.J.Koch
nat
Brassica rapa L.
nat
Capsella bursa-pastoris (L.) Medik.
nat
Cardamine flexuosa With.
nat
K
Cardamine hirsuta L.
nat
K
Coronopus didymus (L.) Sm.
nat
Descurainia sophia (L.) Webb ex Prantl
O Mo L
O
K
M Ka H
Citation
H
Wagner et al. 1990:400 (H); Staples & Herbst 2005:200 (KEY), 201
(DESCR)
M
H
Wagner et al. 1990:400 (O, M, H)
O
L
M
H
Wagner et al. 1990:400 (K, O, Mo, L, H [as Brassica campestris]); AlShehbaz 1985:290/Wagner & Herbst 1995:17 (EM, NOM:Syn. B.
campestris); Herbarium Pacificum Staff 1998:9 (remove Mo, spms. renamed
Sinapis arvensis); Oppenheimer & Bartlett 2002:4 (WM); Staples & Herbst
2005:200 (KEY), 202 (DESCR)
O
L
M
H
Wagner et al. 1990:402 (O, L, M, H [as Capsella rubella]); Wagner & Herbst
1995:17 (NOM:Syn. C. rubella)
O Mo L
M
H
Wagner et al. 1990:402 (K, O, M, H); Hughes 1995:2 (Mo); Oppenheimer
2003:8 (L)
H
Wagner et al. 1990:403 (K, H)
M Ka
H
Wagner et al. 1990:403 (Ni, K, O, Mo, L, M, Ka, H)
nat
M
H
Wagner et al. 1990:405 (EM, H) [Note: based on single colls. from Saddle
Road, Hawai‘i in 1963, and Haleakalä (voucher not seen)]
Hirschfeldia incana (L.) Lagr.-Foss.
nat
M
H
Lorence et al. 1995:26 (NEWNAT/EM, H, DESCR, NOM: Brassica nigra
misapplied, in part)
Lepidium africanum (Burm.f.) DC.
nat
M
H
Wagner et al. 1990:407 (H [as Lepidium hyssopifolium]); Wagner & Herbst
1995:17 (H, NOM: L. hyssopifolium misapplied); Oppenheimer 2003:8 (EM,
WM); Wood 2006:16 (K); Frohlich & Lau 2010:6 (O)
Lepidium arbuscula Hillebr.
end
Lepidium bidentatum Montin var. o-waihiense (Cham.
& Schltdl.) Fosberg
end
Ni
K
K
O Mo L
O
O
K
Wagner et al. 1990:406 (O)
O Mo L
M
H
Wagner et al. 1990:406 (K, O, Mo, L, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 38
Scientific Name
Dicot
Brassicaceae
Status
Kl Le Ni K
Lepidium bonariense L.
nat
Lepidium densiflorum Schrad.
nat
Lepidium oblongum Small
nat
Lepidium orbiculare H.St.John
end
Lepidium remyi Drake
end
Lepidium serra H.Mann
end
K
Lepidium virginicum L.
nat
K
Lobularia maritima (L.) Desv.
nat
Nasturtium microphyllum Boenn. ex Rchb.
nat
Raphanus raphanistrum L.
O Mo L
M Ka H
M
Ni
K
O Mo L
Citation
H
Wagner et al. 1990:407 (H)
H
Wagner et al. 1990:407 (H); Oppenheimer 2004:10 (WM)
M Ka
Wagner et al. 1990:407 (Ni, O, Mo, L, Ka); Lorence et al. 1995:26 (K); Starr
et al. 2006:33 (EM)
K
Wagner et al. 1990:409 (syn. of Lepidium serra); Wagner & Herbst 2003:13
(RESURRECT/K)
H
Wagner et al. 1990:406 (H [as Lepidium bidentatum var. remyi]); Wagner &
Herbst 2003:13 (NOM:Syn. L. bidentatum var. remyi)
Wagner et al. 1990:409 (K)
O Mo L
M
H
Wagner et al. 1990:409 (O, Mo, M, H); Lorence et al. 1995:27 (K);
Oppenheimer 2010:34 (L)
O Mo
M
H
Wagner et al. 1990:409 (M); Lorence et al. 1995:27 (Mo); Wagner & Herbst
1995:17 (O); Oppenheimer 2003:9 (H); Staples & Herbst 2005:200 (KEY),
203 (DESCR)
K
O Mo
M
H
Wagner et al. 1990:411 (K, O, M, H [as Nasturtium microphyllum]); Wagner
& Herbst 1995:18 (NOM: N. microphyllum = Rorippa microphylla); AlShehbaz & Price 1998:124/Herbst & Wagner 1999:15 (NOM:Syn. R.
microphylla); Staples & Herbst 2005:200 (KEY), 204 (DESCR); Wysong et
al. 2007:3 (Mo)
nat
K
O Mo
M
H
Wagner et al. 1990:412 (K?, H? [possibly sparingly naturalized]); Staples et
al. 2003:9 (NEWNAT/K, O, Mo, EM, H)
Raphanus sativus L.
nat
K
O Mo
M Ka
H
Wagner et al. 1990:412 (K, O, Mo, M, Ka, H); Staples & Herbst 2005:200
(KEY), 204 (DESCR)
Rapistrum rugosum (L.) All.
nat
K
Lorence et al. 1995:27 (NEWNAT/K, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 39
Scientific Name
Dicot
Brassicaceae
Status
Kl Le Ni K
K
O Mo L
O Mo
M Ka H
Rorippa sarmentosa (G.Forst. ex DC.) J.F.Macbr.
pol?
M
Sinapis arvensis L.
nat
Sisymbrium altissimum L.
nat
Sisymbrium irio L.
nat
Sisymbrium officinale (L.) Scop. var. leiocarpum DC.
nat
Sisymbrium officinale (L.) Scop. var. officinale
nat
K
O Mo L
Buddleja asiatica Lour.
nat
K
O Mo L
Buddleja davidii Franch.
nat
K
M
Buddleja madagascariensis Lam.
nat
K
M
Acanthocereus tetragonus (L.) Hummelinck
nat
K
Cereus uruguayanus F.Ritter ex R.Kiesling
nat
K
Harrisia bonplandii (Parm. ex Pfeiff.) Britton & Rose
nat
K
H
Mo
K
Citation
Wagner et al. 1990:411 (K, O, Mo, M, H [as Nasturtium sarmentosum]);
Wagner & Herbst 1995:18 (NOM:Syn. N. sarmentosum)
Herbarium Pacificum Staff 1998:9 (NEWNAT/Mo, DESCR, NOM: Brassica
campestris [=B. rapa] misapplied, in part) [Note: 2 colls. in 1928 from weedy
field in Ho‘olehua]
Mo
M Ka
H
Wagner et al. 1990:413 (K, M, Ka, H); Wysong et al. 2007:3 (Mo)
H
Wagner et al. 1990:413 (H)
H
Wagner et al. 1990:413 (L, H) [Note: single coll. from each island—Lalakoa,
Läna‘i, 1930; Hualalai, Hawai‘i, 1911]
M
H
Wagner et al. 1990:413 (K, Mo, L, M, H); Frohlich & Lau 2008:4 (O)
M
H
Wagner et al. 1990:415 (O, Mo, M, H); Oppenheimer & Bartlett 2002:5 (L);
Wood 2006:16 (K)
L
Buddlejaceae
Wagner et al. 1990:415 (K? [noted as cultivated at Köke‘e]); Shannon &
Wagner 1996:13 (NEWNAT/K); Starr et al. 2003:26 (EM); Staples & Herbst
2005:205 (KEY), 206 (DESCR)
H
Wagner et al. 1990:416 (H); Lorence & Flynn 1999:4 (K); Oppenheimer &
Bartlett 2000:3 (EM); Staples & Herbst 2005:205 (KEY), 206 (DESCR)
Cactaceae
Lorence et al. 1995:27 (NEWNAT/K, DESCR)
M
Wagner et al. 1990:417 (K); Staples & Herbst 2005:210 (KEY), 211
(DESCR); Oppenheimer 2008:25 (EM)
Lorence et al. 1995:28 (NEWNAT/K, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 40
Scientific Name
Dicot
Cactaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Harrisia martinii (Labour.) Britton
nat
K
Wagner et al. 1990:418 (K)
Hylocereus costaricensis (F.A.C.Weber) Britton & Rose
nat
K
Lorence et al. 1995:28 (NEWNAT/K, DESCR); Staples & Herbst 2005:214
(DESCR) [Note: BISH voucher (Flynn & Hume 3571) has since been
annotated as Hylocereus cf. trigonus (Haw.) Safford by B. Leuenberger, 16
Oct 2000]
Hylocereus undatus (Haw.) Britton & Rose
nat
Opuntia cochenillifera (L.) Mill.
nat
Opuntia ficus-indica (L.) Mill.
nat
Opuntia monacantha (Willd.) Haw.
nat
Pereskia aculeata Mill.
nat
Schlumbergera truncata (Haw.) Moran
Selenicereus macdonaldiae (Hook.) Britton & Rose
Ni
Le?
K
O Mo L
M Ka
K
O
M
K
O Mo L
M Ka
H
Wagner et al. 1990:419 (K, O); Oppenheimer 2004:10 (WM); Staples &
Herbst 2005:210 (KEY), 216 (DESCR)
H
O
O?
nat
Wagner et al. 1990:419 (K, O, M, Ka, H); Hughes 1995:2 (Mo);
Oppenheimer & Bartlett 2002:5 (L); Staples & Herbst 2005:210 (KEY), 216
(DESCR); Wood & LeGrande 2006:22 (Le? [reported but unvouchered by
Caum in 1936])
Wagner et al. 1990:420 (O [as Opuntia vulgaris]); Staples et al. 2002:6
(NOM: O. vulgaris misapplied)
Mo?
?
Wagner et al. 1990:418 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:210 (KEY), 213 (DESCR)
H
Parker & Parsons 2012b:66 (NEWNAT/Mo? [ID unconfirmed], H, DESCR)
Wagner et al. 1990:416 (O? [reportedly escaping in Honolulu, but
unvouchered], DESCR); Staples & Herbst 2005:210 (KEY), 215 (DESCR)
K
Lorence et al. 1995:28 (NEWNAT/K [as Selenicereus grandiflorus],
DESCR); Herbst & Wagner 1999:16 (NOM: S. grandiflorus misapplied);
Staples & Herbst 2005:210 (KEY), 215 (DESCR)
K
Wagner et al. 1990:423 (Ni, K); Staples & Herbst 2005:217 (KEY, DESCR)
Campanulaceae
Brighamia insignis A.Gray
end
Ni
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 41
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Brighamia rockii H.St.John
end
Mo L? M?
Wagner et al. 1990:423 (Mo, L? [based on Rock sighting in 1910 at Maunalei
Valley of inaccessible plant], M? [based on Remy 1850s voucher]); Staples &
Herbst 2005:217 (KEY, DESCR)
Clermontia arborescens (H.Mann) Hillebr. ssp.
arborescens
end
M
Wagner et al. 1990:427 (WM); Lammers 1991:30 (KEY, DESCR)
Clermontia arborescens (H.Mann) Hillebr. ssp. waihiae
(Wawra) Lammers
end
M
Wagner et al. 1990:427 (Mo, L, EM, WM); Lammers 1991:28 (remove Mo,
L to subsp. waikoluensis, KEY, DESCR)
Clermontia arborescens (H.Mann) Hillebr. ssp.
waikoluensis (H.St.John) Lammers
end
Clermontia calophylla E.Wimm.
end
H
Wagner et al. 1990:427 (H); Lammers 1991:65 (KEY, DESCR)
Clermontia clermontioides (Gaudich.) A.Heller ssp.
clermontioides
end
H
Wagner et al. 1990:427 (H); Lammers 1991:16 (KEY, DESCR)
Clermontia clermontioides (Gaudich.) A.Heller ssp.
rockiana (E.Wimm.) Lammers
end
H
Wagner et al. 1990:428 (H); Lammers 1991:18 (KEY, DESCR)
Clermontia drepanomorpha Rock
end
H
Wagner et al. 1990:428 (H); Lammers 1991:56 (KEY, DESCR)
Clermontia fauriei H.Lév.
end
Clermontia grandiflora Gaudich. ssp. grandiflora
end
M
Wagner et al. 1990:430 (WM); Lammers 1991:76 (KEY, DESCR)
Clermontia grandiflora Gaudich. ssp. maxima Lammers
end
M
Lammers 1991:77 (SSPNOV/EM, KEY, DESCR)/Herbst & Wagner
1999:16; Wood 2012:91 (REDISCOVER/EM)
Clermontia grandiflora Gaudich. ssp. munroi
(H.St.John) Lammers
end
M
Wagner et al. 1990:430 (Mo, L, EM, WM); Lammers 1991:72 (KEY,
DESCR)
Clermontia hawaiiensis (Hillebr.) Rock
end
Mo L
K
Wagner et al. 1990:427 (as Clermontia waikoluensis, syn. of C. arborescens
subsp. waihiae); Lammers 1991:25 (RESURRECT/Mo, L, COMBNOV &
STATNOV:Syn. Delissea fauriei)/Herbst & Wagner 1999:16
O
Wagner et al. 1990:428 (K, O); Lammers 1991:35 (KEY, DESCR)
Mo L
H
Wagner et al. 1990:430 (H); Lammers 1991:80 (KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 42
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
O Mo
M Ka H
M
Citation
Clermontia kakeana Meyen
end
Clermontia kohalae Rock
end
Clermontia lindseyana Rock
end
Clermontia micrantha (Hillebr.) Rock
end
Clermontia montis-loa Rock
end
Clermontia multiflora Hillebr.
end
Clermontia oblongifolia Gaudich. ssp. brevipes
(E.Wimm.) Lammers
end
Clermontia oblongifolia Gaudich. ssp. mauiensis (Rock)
Lammers
end
Clermontia oblongifolia Gaudich. ssp. oblongifolia
end
Clermontia pallida Hillebr.
end
Clermontia parviflora Gaudich. ex A.Gray
end
H
Wagner et al. 1990:433 (H); Lammers 1991:60 (KEY, DESCR)
Clermontia peleana Rock ssp. peleana
end
H
Wagner et al. 1990:435 (EM, H [as Clermontia peleana]); Lammers 1991:34
(remove EM spms. to C. peleana subsp. singuliflora, KEY, DESCR)
Clermontia peleana Rock ssp. singuliflora (Rock)
Lammers
end
H
Wagner et al. 1990:435 (as Clermontia singuliflora, syn. of C. peleana);
Lammers 1991:32 (RESURRECT/M, H, COMBNOV & STATNOV:Syn. C.
singuliflora, KEY, DESCR); Herbst & Wagner 1999:16
Clermontia persicifolia Gaudich.
end
Clermontia pyrularia Hillebr.
end
M
L
Wagner et al. 1990:430 (O, Mo, EM, WM); Lammers 1991:38 (KEY,
DESCR)
H
Wagner et al. 1990:431 (H); Lammers 1991:52 (KEY, DESCR)
H
Wagner et al. 1990:431 (EM, H); Lammers 1991:45 (KEY, DESCR)
M
Wagner et al. 1990:432 (L, WM); Lammers 1991:59 (KEY, DESCR)
H
O
M
Wagner et al. 1990:432 (H); Lammers 1991:54 (KEY, DESCR)
Wagner et al. 1990:432 (O, WM); Lammers 1991:58 (KEY, DESCR)
Mo
Wagner et al. 1990:433 (Mo); Lammers 1991:70 (KEY, DESCR)
L
M
Wagner et al. 1990:433 (L, EM, WM); Lammers 1991:69 (KEY, DESCR)
O
Wagner et al. 1990:433 (O); Lammers 1991:67 (KEY, DESCR)
Mo
Wagner et al. 1990:433 (Mo); Lammers 1991:50 (KEY, DESCR)
M
O
Wagner et al. 1990:435 (O); Lammers 1991:47 (KEY, DESCR)
H
Wagner et al. 1990:436 (H); Lammers 1991:21 (KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 43
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Clermontia samuelii C.N.Forbes ssp. hanaensis
(H.St.John) Lammers
end
M
Wagner et al. 1990:436 (EM); Lammers 1991:79 (KEY, DESCR)
Clermontia samuelii C.N.Forbes ssp. samuelii
end
M
Wagner et al. 1990:436 (EM); Lammers 1991:80 (KEY, DESCR)
Clermontia tuberculata C.N.Forbes
end
M
Wagner et al. 1990:436 (EM); Lammers 1991:30 (KEY, DESCR)
Clermontia waimeae Rock
end
Cyanea aculeatiflora Rock
end
Cyanea acuminata (Gaudich.) Hillebr.
end
O
Cyanea angustifolia (Cham.) Hillebr.
end
O Mo L
Cyanea arborea Hillebr.
end
Cyanea asarifolia H.St.John
end
Cyanea asplenifolia (H.Mann) Hillebr.
end
Cyanea calycina (Cham.) Lammers
end
Cyanea comata Hillebr.
end
Cyanea copelandii Rock ssp. copelandii
end
Cyanea copelandii Rock ssp. haleakalaensis (H.St.John)
Lammers
end
Cyanea coriacea (A.Gray) Hillebr.
end
H
M
Wagner et al. 1990:437 (H); Lammers 1991:22 (KEY, DESCR)
Wagner et al. 1990:443 (EM)
Wagner et al. 1990:444 (O)
M
Wagner et al. 1990:444 (O, Mo, L, WM); Lammers 2004:100 (KEY)
M
Wagner et al. 1990:444 (EM)
K
Wagner et al. 1990:445 (K); Lammers 2004:99 (KEY)
M
Wagner et al. 1990:445 (WM)
O
Wagner et al. 1990:483 (O [as Rollandia lanceolata subsp. calycina]);
Lammers 1993:439 (COMBNOV to Cyanea lanceolata subsp. calycina);
Lammers 1998:32 (COMBNOV & STATNOV:Syn. C. lanceolata subsp.
calycina, KEY)/Herbst & Wagner 1999:16, 17
M
Wagner et al. 1990:445 (EM)
H
M
K
Wagner et al. 1990:446 (H)
Wagner et al. 1990:446 (EM)
Wagner et al. 1990:446 (K [as Cyanea coriacea]), 448 (K [as C. fauriei);
Lammers 2004:96 (NOM:Syn. C. fauriei, KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 44
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Cyanea crispa (Gaudich.) Lammers, Givnish & Sytsma
end
Wagner et al. 1990:481 (O [as Rollandia crispa]); Lammers 1993:439
(COMBNOV:Syn. R. crispa)/Herbst & Wagner 1999:17
Cyanea cylindrocalyx (Rock) Lammers
end
Cyanea dolichopoda Lammers & Lorence
end
Cyanea dunbariae Rock
end
Cyanea duvalliorum Lammers & H.Oppenh.
end
Cyanea eleeleensis (H.St.John) Lammers
end
Cyanea elliptica (Rock) Lammers
end
Cyanea fissa (H.Mann) Hillebr. ssp. fissa
end
K
Wagner et al. 1990:449 (K); Lammers 1992:130 (KEY)
Cyanea fissa (H.Mann) Hillebr. ssp. gayana (Rock)
Lammers
end
K
Wagner et al. 1990:449 (K)
Cyanea floribunda E.Wimm.
end
H
Wagner et al. 1990:446 (H [as Cyanea degeneriana, Syn. C. floribunda);
Wagner & Herbst 2003:15 (NOM: C. degeneriana misapplied [type of C.
degeneriana = C. hirtella], replacement name is C. floribunda [based on T.
Lammers, unpublished])
Cyanea giffardii Rock
end
H
Wagner et al. 1990:449 (H)
Cyanea glabra (E.Wimm.) H.St.John
end
H
K
Wagner et al. 1990:451 (H [as Cyanea grimesiana subsp. cylindrocalyx]);
Lammers 1998:31 (COMBNOV & STATNOV:Syn. C. grimesiana subsp.
cylindrocalyx, KEY)/Herbst & Wagner 1999:16; Lammers 2004:87 (KEY)
Lammers & Lorence 1993:432 (SPNOV/K, DESCR)/Herbst & Wagner
1999:16; Lammers 2004:99 (KEY)
Mo
Wagner et al. 1990:448 (Mo [as Cyanea dunbarii]); Hughes 1995:3
(REDISCOVER/Mo); Herbst & Wagner 1999:16 (spelling change)
M
Lammers 2004:89 (SPNOV/EM, KEY, DESCR); Oppenheimer 2008:25
(status)
K
St. John 1987b:341 (SPNOV/K [as Delissea eleeleensis], DESCR); Lammers
1992:129 (COMBNOV:Syn. D. eleeleensis, KEY, DESCR)/Herbst &
Wagner 1999:16; Wood 2012:92 (EXTINCT?)
L
M
M
Wagner et al. 1990:448 (L, M); Lammers 2004:100 (KEY)
Wagner et al. 1990:449 (EM); Lammers 2004:94 (KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 45
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Cyanea grimesiana Gaudich. ssp. grimesiana
end
O Mo
Wagner et al. 1990:451 (O, Mo, L, M); Lammers 1998:31 (remove L
[=Cyanea munroi], M [=C. mauiensis], KEY); Lammers 2004:87 (KEY)
Cyanea grimesiana Gaudich. ssp. obatae (H.St.John)
Lammers
end
O
Wagner et al. 1990:451 (O); Lammers 1998:31 (KEY); Lammers 2004:87
(KEY)
Cyanea habenata (H.St.John) Lammers
end
Cyanea hamatiflora Rock ssp. carlsonii (Rock) Lammers
end
Cyanea hamatiflora Rock ssp. hamatiflora
end
Cyanea hardyi Rock
end
K
Wagner et al. 1990:452 (K); Lammers 2004:100 (KEY)
Cyanea hirtella (H.Mann) Hillebr.
end
K
Wagner et al. 1990:452 (K [as Cyanea degeneriana]), 465 (K [as C.
sylvestris]); Lammers 1992:130 (KEY [for C. hirtella, C. sylvestris]); Wagner
& Herbst 2003:15 (NOM:Syn. C. degeneriana = C. hirtella [based on T.
Lammers, unpublished]); Wagner et al. 2005– (NOM:Syn. C. sylvestris)
Cyanea horrida (Rock) O.Deg. & Hosaka
end
Cyanea humboldtiana (Gaudich.) Lammers, Givnish &
Sytsma
end
Cyanea kahiliensis (H.St.John) Lammers
end
K
St. John 1987b:343 (SPNOV/K [as Delissea kahiliensis], DESCR); Wagner et
al. 1990:464 (K [as Cyanea spathulata subsp. longipetiolata); Lammers
1998:33 (COMBNOV:Syn. D. kahiliensis, C. spathulata subsp.
longipetiolata, DESCR)/Herbst & Wagner 1999:16; Lammers 2004:100
(KEY)
Cyanea kolekoleensis (H.St.John) Lammers
end
K
St. John 1987b:344 (SPNOV/K [as Delissea kolekoleensis], DESCR);
Lammers 1992:130 (COMBNOV:Syn. D. kolekoleensis, KEY, DESCR)/
Herbst & Wagner 1999:16; Wood 2012:92 (EXTINCT?, DESCR)
K
St. John 1987b:342 (SPNOV/K [as Delissea habenata], DESCR); Lammers
1998:33 (COMBNOV:Syn. D. habenata, DESCR)/Herbst & Wagner
1999:16; Lammers 2004:100 (KEY)
H
M
M
O
Wagner et al. 1990:452 (H)
Wagner et al. 1990:452 (EM)
Wagner et al. 1990:453 (EM)
Wagner et al. 1990:483 (O [as Rollandia humboldtiana]); Lammers 1993:439
(COMBNOV:Syn. R. humboldtiana)/Herbst & Wagner 1999:17
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 46
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Cyanea koolauensis Lammers, Givnish & Sytsma
end
Wagner et al. 1990:481 (O [as Rollandia angustifolia]); Lammers 1993:439
(NOMNOV:Syn. R. angustifolia)/Herbst & Wagner 1999:17
Cyanea kuhihewa Lammers
end
Cyanea kunthiana Hillebr.
end
Cyanea lanceolata (Gaudich.) Lammers, Givnish &
Sytsma
end
Cyanea leptostegia A.Gray
end
K
Wagner et al. 1990:453 (K)
Cyanea linearifolia Rock
end
K
Wagner et al. 1990:454 (K); Lammers 2004:100 (KEY)
Cyanea lobata H.Mann ssp. baldwinii (C.N.Forbes &
G.C.Munro) Lammers
end
Cyanea lobata H.Mann ssp. lobata
end
Cyanea longiflora (Wawra) Lammers, Givnish & Sytsma
end
Cyanea longissima (Rock) H.St.John
end
Cyanea macrostegia Hillebr. ssp. gibsonii (Hillebr.)
Lammers
end
Cyanea macrostegia Hillebr. ssp. macrostegia
end
M
Wagner et al. 1990:456 (EM, WM)
Cyanea magnicalyx Lammers
end
M
Lammers 2004:85 (SPNOV/WM, KEY, DESCR)
K
Lammers 1996:238 (SPNOV/K, DESCR)/Herbst & Wagner 1999:17; Wood
2007:14 (EXTINCT?)
M
O
Wagner et al. 1990:453 (M)
Wagner et al. 1990:483 (O [as Rollandia lanceolata subsp. lanceolata]);
Lammers 1993:439 (COMBNOV:Syn. R. lanceolata/Herbst & Wagner
1999:17; Lammers 1998:32 (KEY)
L
Wagner et al. 1990:454 (as Cyanea baldwinii, syn. of C. lobata); Lammers
1999:387 (RESURRECT/L, COMBNOV & STATNOV:Syn. C. baldwinii,
KEY); Wood 2007:15 (REDISCOVER/L)
M
O
Wagner et al. 1990:454 (L, WM [as Cyanea lobata]); Lammers 1999:387
(remove L [due to resurrection of C. baldwinii], KEY)
Wagner et al. 1990:484 (O [as Rollandia longiflora]); Lammers 1993:439
(COMBNOV:Syn. R. longiflora)/Herbst & Wagner 1999:17; Lammers
1998:33 (KEY)
M
L
Wagner et al. 1990:454 (EM); Lammers 2004:94 (KEY)
Wagner et al. 1990:456 (L)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 47
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Mo
Citation
Cyanea mannii (Brigham) Hillebr.
end
Wagner et al. 1990:456 (Mo); Lammers 2004:99 (KEY)
Cyanea maritae Lammers & H.Oppenh.
end
Cyanea marksii Rock
end
Cyanea mauiensis (Rock) Lammers
end
M
Wagner et al. 1990:451 (as Cyanea grimesiana var. mauiensis, syn. of C.
grimesiana subsp. grimesiana); Lammers 1998:31 (RESURRECT/M,
COMBNOV & STATNOV:Syn. C. grimesiana var. mauiensis, KEY)/Herbst
& Wagner 1999:17; Lammers 2004:86 (KEY)
Cyanea mceldowneyi Rock
end
M
Wagner et al. 1990:457 (EM); Lammers 2004:94 (KEY)
Cyanea membranacea Rock
end
Cyanea minutiflora Lammers
end
Cyanea munroi (Hosaka) Lammers
end
Cyanea obtusa (A.Gray) Hillebr.
end
Cyanea parvifolia (C.N.Forbes) Lammers, Givnish &
Sytsma
end
Cyanea pilosa A.Gray ssp. longipedunculata (Rock)
Lammers
end
H
Wagner et al. 1990:458 (H)
Cyanea pilosa A.Gray ssp. pilosa
end
H
Wagner et al. 1990:458 (H)
Cyanea pinnatifida (Cham.) E.Wimm.
end
M
Lammers 2004:92 (SPNOV/EM, KEY, DESCR); Oppenheimer 2008:25
(status)
H
O
Wagner et al. 1990:457 (H)
Wagner et al. 1990:457 (O); Lammers 2004:99 (KEY)
K
Lammers 2004:98 (SPNOV/K, KEY, DESCR)
Mo L
Wagner et al. 1990:451 (as Cyanea grimesiana var. munroi, syn. of C.
grimesiana subsp. grimesiana); Lammers 1998:31 (RESURRECT/Mo, L,
COMBNOV & STATNOV:Syn. C. grimesiana var. munroi, KEY)/Herbst &
Wagner 1999:17; Lammers 2004:87 (KEY)
M
Wagner et al. 1990:458 (M); Lammers 2004:99 (KEY)
K
Wagner et al. 1990:484 (K [as Rollandia parvifolia]); Lammers 1993:439
(COMBNOV:Syn. R. parvifolia)/Herbst & Wagner 1999:17
O
Wagner et al. 1990:459 (O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 48
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
H
Citation
Cyanea platyphylla (A.Gray) Hillebr.
end
Cyanea pohaku Lammers
end
Cyanea procera Hillebr.
end
Mo
Wagner et al. 1990:460 (Mo); Hughes 1995:3 (REDISCOVER/Mo)
Cyanea profuga C.N.Forbes
end
Mo
Wagner et al. 1990:460 (Mo); Wood 2007:15 (REDISCOVER/Mo)
Cyanea pseudofauriei Lammers
end
Cyanea purpurellifolia (Rock) Lammers, Givnish &
Sytsma
end
Cyanea pycnocarpa (Hillebr.) E.Wimm.
end
Cyanea quercifolia (Hillebr.) E.Wimm.
end
Cyanea recta (Wawra) Hillebr.
end
K
Wagner et al. 1990:461 (K); Lammers 1992:130 (KEY); Lorence et al.
1995:29 (REDISCOVER/K)
Cyanea remyi Rock
end
K
Wagner et al. 1990:466 (not placed, possibly referable to Cyanea truncata);
Lammers & Lorence 1993:434 (RESURRECT/K, DESCR)/Herbst & Wagner
1999:17
Cyanea rivularis Rock
end
K
Wagner et al. 1990:469 (K [as Delissea rivularis]); Lorence et al. 1995:30
(REDISCOVER/K); Lammers 2005:62 (NOM:Syn. D. rivularis)
Cyanea salicina H.Lév.
end
K
Wagner et al. 1990:461 (syn. of Cyanea recta); Wagner & Herbst 1999:1868
(RESURRECT/K, based on Lammers unpubl.)
Cyanea scabra Hillebr.
end
Cyanea sessilifolia (O.Deg.) Lammers
end
M
Wagner et al. 1990:459 (H); Lammers 2004:94 (KEY)
Wagner et al. 1990:460 (EM)
K
Lammers 2004:96 (SPNOV/K, KEY, DESCR)
O
Wagner et al. 1990:484 (O [as Rollandia purpurellifolia]); Lammers
1993:439 (COMBNOV:Syn. R. purpurellifolia)/Herbst & Wagner 1999:17
H
M
M
O
Wagner et al. 1990:461 (H)
Wagner et al. 1990:461 (EM)
Wagner et al. 1990:461 (WM); Lammers 2004:94 (KEY)
Wagner et al. 1990:484 (as Rollandia sessilifolia, syn. of R. longiflora);
Lammers 1998:32 (RESURRECT/O, COMBNOV:Syn. R. sessilifolia,
KEY)/Herbst & Wagner 1999:18
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 49
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
H
Citation
Cyanea shipmanii Rock
end
Cyanea solanacea Hillebr.
end
Mo
Cyanea solenocalyx Hillebr.
end
Mo
Cyanea spathulata (Hillebr.) A.Heller
end
Cyanea st.-johnii (Hosaka) Lammers, Givnish & Sytsma
end
Cyanea stictophylla Rock
end
Cyanea superba (Cham.) A.Gray ssp. regina (Wawra)
Lammers
end
O
Wagner et al. 1990:465 (O)
Cyanea superba (Cham.) A.Gray ssp. superba
end
O
Wagner et al. 1990:465 (O)
Cyanea tritomantha A.Gray
end
Cyanea truncata (Rock) Rock
end
Cyanea undulata C.N.Forbes
end
Delissea argutidentata (E.Wimm.) H.St.John
end
H
Wagner et al. 1990:471 (syn. of Delissea undulata subsp. undulata); Lammers
2005:22 (RESURRECT/H, KEY, DESCR)
Delissea fallax Hillebr.
end
H
Wagner et al. 1990:468 (H); Lammers 2005:26 (KEY, DESCR)
M?
Wagner et al. 1990:462 (H)
Wagner et al. 1990:462 (Mo, WM? [based on a few sterile colls. from Pu‘u
Kukui])
Wagner et al. 1990:464 (Mo)
K
Wagner et al. 1990:465 (K [as Cyanea spathulata subsp. spathulata]);
Lammers 1998:33 (revert to C. spathulata, as C. spathulata subsp.
longipetiolata now = C. kahiliensis); Lammers 2004:100 (KEY)
O
Wagner et al. 1990:484 (O [as Rollandia st.-johnii]); Lammers 1993:440
(COMBNOV:Syn. R. st.-johnii)/Herbst & Wagner 1999:18
H
H
O
Wagner et al. 1990:465 (H)
Wagner et al. 1990:466 (H)
Wagner et al. 1990:466 (O)
K
Wagner et al. 1990:467 (K); Lammers 1992:130 (KEY); Lorence et al.
1995:29 (REDISCOVER/K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 50
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Mo
Citation
Delissea fauriei H.Lév.
end
Wagner et al. 1990:449 (species dubia); Lammers 1991:25 (NOM:syn. of
Clermontia arborescens subsp. waikoluensis); Lammers 2005:42
(RESURRECT/Mo, KEY, DESCR)
Delissea kauaiensis (Lammers) Lammers
end
Delissea laciniata Hillebr.
end
Delissea lanaiensis (Rock) Lammers
end
Delissea lauliiana Lammers
end
Delissea niihauensis H.St.John
end
Delissea parviflora Hillebr.
end
Delissea rhytidosperma H.Mann
end
Delissea sinuata Hillebr.
end
O
Wagner et al. 1990:471 (O [as Delissea sinuata subsp. sinuata]); Lammers
1998:34 (remove L [D. sinuata subsp. lanaiensis = D. lanaiensis], KEY);
Lammers 2005:41 (KEY, DESCR)
Delissea subcordata Gaudich. ssp. obtusifolia (Wawra)
Lammers
end
O
Wagner et al. 1990:471 (as Delissea subcordata var. obtusifolia, syn. of D.
subcordata); Lammers 2005:34 (RESURRECT/O, COMBNOV &
STATNOV:Syn. D. subcordata var. obtusifolia, KEY, DESCR)
K
Wagner et al. 1990:472 (K [as Delissea undulata subsp. kauaiensis]);
Lammers 1999:388 (COMBNOV:Syn. D. undulata subsp. kauaiensis = D.
niihauensis subsp. kauaiensis, KEY); Lammers 2005:19 (COMBNOV &
STATNOV:Syn. D. niihauensis subsp. kauaiensis, KEY, DESCR)
O
Wagner et al. 1990:468 (O); Lammers 2005:40 (KEY, DESCR)
L
Wagner et al. 1990:471 (L [as Delissea sinuata subsp. lanaiensis]); Lammers
1998:34 (COMBNOV & STATNOV:Syn. D. sinuata subsp. lanaiensis, KEY,
ELEV)/Herbst & Wagner 1999:18; Lammers 2005:42 (KEY, DESCR)
O
Wagner et al. 1990:468 (O); Lammers 2005:26 (KEY, DESCR)
Ni
Wagner et al. 1990:472 (Ni [as Delissea undulata subsp. niihauensis]);
Lammers 1999:388 (NOM:Syn. D. undulata subsp. niihauensis = D.
niihauensis subsp. niihauensis, KEY); Lammers 2005:17 (NOM: D.
niihauensis [following elevation of D. niihauensis subsp. kauaiensis to D.
kauaiensis], KEY, DESCR)
H
K
Wagner et al. 1990:469 (H); Lammers 2005:28 (KEY, DESCR)
Wagner et al. 1990:469 (K); Lammers 2005:25 (KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 51
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Delissea subcordata Gaudich. ssp. subcordata
end
O
Wagner et al. 1990:471 (O [as Delissea subcordata]); Lammers 2005:33
(KEY, DESCR)
Delissea takeuchii Lammers
end
O
Lammers 2005:38 (SPNOV/O, KEY, DESCR)
Delissea undulata Gaudich.
end
Delissea waianaeensis Lammers
end
Hippobroma longiflora (L.) G.Don
nat
K
Lobelia Xkauaiensis (A.Gray) A.Heller
end
K
Lobelia dunbariae Rock ssp. dunbariae
end
Mo
Wagner et al. 1990:474 (Mo [as Lobelia dunbarii subsp. dunbarii]); Wagner
et al. 2005–: epithet spelling change
Lobelia dunbariae Rock ssp. paniculata (Rock) Lammers
end
Mo
Wagner et al. 1990:474 (Mo [as Lobelia dunbarii subsp. paniculata]); Wagner
et al. 2005–: epithet spelling change
Lobelia erinus L.
nat
Lobelia gaudichaudii A.DC.
end
M
Wagner et al. 1990:472 (M, H [as Delissea undulata subsp. undulata], K [as
subsp. kauaiensis], Ni [as subsp. niihauensis]); Lammers 1999:387 (NOM:
revert to D. undulata [following transfer of subspp. kauaiensis & niihauensis
to D. niihauensis], KEY); Lammers 2005:21 (KEY, DESCR), 22 (remove H,
following resurrection of syn. D. argutidentata)
O
Lammers 2005:34 (SPNOV/O, KEY, DESCR)
O
M
H
Wagner et al. 1990:472 (K, O, M, H); Staples & Herbst 2005:217 (KEY),
218 (DESCR)
Wagner et al. 1990:478 (K [as Lobelia kauaensis]); Lammers 2007:801
(STATNOV (hybrid status, spelling change), KEY, DESCR)
H
O
Wagner et al. 1990:476 (H); Staples & Herbst 2005:217 (KEY, DESCR)
[Note: single naturalized coll. in 1931 from Hawai‘i Volcanoes National Park]
Wagner et al. 1990:476 (O [as Lobelia gaudichaudii subsp. gaudichaudii]);
Lammers 2007:804 (NOM: revert to L. gaudichaudii, following removal of
subsp. koolauensis = L. koolauensis, KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 52
Scientific Name
Dicot
Campanulaceae
Status
Lobelia gloria-montis Rock ssp. gloria-montis
end
Lobelia gloria-montis Rock ssp. longibracteata (Rock)
Lammers
Kl Le Ni K
O Mo L
Mo
M Ka H
Citation
M
Wagner et al. 1990:476 (Mo? [sterile colls. from Päpa‘alä Pali], M [as
Lobelia gloria-montis]); Lammers 2007:805 (confirmation of ID of Mo
records; NOM: L. gloria-montis subsp. gloria-montis, following addition of
subsp. longibracteata, KEY, DESCR)
end
M
Wagner et al. 1990:476 (as Lobelia gloria-montis var. longibracteata, syn. of
L. gloria-montis); Lammers 2007:806 (RESURRECT/EM, WM, COMBNOV
& STATNOV, Syn. L. gloria-montis var. longibracteata, KEY, DESCR)
Lobelia grayana E.Wimm.
end
M
Wagner et al. 1990:477 (EM)
Lobelia hillebrandii Rock
end
Mo
M
Wagner et al. 1990:477 (M); Lorence et al. 1995:30 (Mo)
Lobelia hypoleuca Hillebr.
end
O Mo L
M
Lobelia koolauensis (Hosaka & Fosberg) Lammers
end
O
Wagner et al. 1990:476 (O [as Lobelia gaudichaudii subsp. koolauensis]);
Lammers 2007:803 (COMBNOV & STATNOV:Syn. L. gaudichaudii subsp.
koolauensis, KEY, DESCR)
Lobelia monostachya (Rock) Lammers
end
O
Wagner et al. 1990:478 (O)
Lobelia niihauensis H.St.John
end
O
Wagner et al. 1990:478 (Ni, K, O); Wagner & Herbst 2003:16 (NOM:Syn.
Neowimmera niihauensis var. forbesii)
Lobelia oahuensis Rock
end
O
Wagner et al. 1990:479 (O)
Lobelia remyi Rock
end
O
Wagner et al. 1990:479 (O)
Lobelia villosa (Rock) H.St.John & Hosaka
end
K
Wagner et al. 1990:479 (K); Lammers 2007:800 (KEY, DESCR)
Lobelia wahiawa Lammers
end
K
Lammers 2007:802 (SPNOV/K, KEY, DESCR)
Lobelia yuccoides Hillebr.
end
K
K
Ni
K
O
H
Wagner et al. 1990:477 (K, O, Mo, L, M, H)
Wagner et al. 1990:480 (K, O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 53
Scientific Name
Dicot
Campanulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Platycodon grandiflorus (Jacq.) A.DC.
nat
Staples & Herbst 2005:217 (KEY), 218 (DESCR); Lau & Frohlich 2012:12
(NEWNAT/O, DESCR)
Trematolobelia auriculata H.St.John
end
Trematolobelia grandifolia (Rock) O.Deg.
end
Trematolobelia kaalae (O.Deg.) Lammers
end
Trematolobelia kauaiensis (Rock) Skottsb.
end
Trematolobelia macrostachys (Hook. & Arn.) Zahlbr.
end
O Mo
Trematolobelia rockii H.St.John
end
Mo
Trematolobelia singularis H.St.John
end
Trematolobelia wimmeri O.Deg. & I.Deg.
end
H
Wagner et al. 1990:486 (syn. of Trematolobelia grandifolia); Lammers
2009:140 (RESURRECT/H, Syn. T. lustrialis [former syn. of T.
macrostachys], KEY, DESCR)
Triodanis biflora (Ruiz & Pav.) Greene
nat
H
Wagner et al. 1990:488 (H) [Note: only collected twice in 1945 in Hawai‘i
Volcanoes National Park]
Wahlenbergia gracilis (G.Forst.) A.DC.
nat
M
H
Wagner et al. 1990:489 (Mo, L, M, H); Herbst et al. 2004:4, 5 (KEY,
DESCR)
Wahlenbergia marginata (Thunb.) A.DC.
nat
M
H
Herbst et al. 2004:4, 5 (NEWNAT/H, KEY, DESCR); Oppenheimer 2008:26
(EM)
L
Wagner et al. 1990:486 (syn. of Trematolobelia macrostachys); Lammers
2009:138 (RESURRECT/L, KEY, DESCR)
H
O
Wagner et al. 1990:486 (H); Lammers 2009:139 (KEY, DESCR)
Wagner et al. 1990:486 (as Trematolobelia macrostachys var. kaalae, syn. of
T. macrostachys); Lammers 2009:134 (RESURRECT/O & STATNOV, Syn.
T. macrostachys var. kaalae, KEY, DESCR)
K
Wagner et al. 1990:486 (K); Lammers 2009:132 (KEY, DESCR)
M
Wagner et al. 1990:486 (O, Mo, L, EM, WM, H); Lammers 2009:134 (KEY,
DESCR, remove L [following resurrection of Trematolobelia auriculata], H
[following transfer of T. lustrialis to synonymy of T. wimmeri)
Wagner et al. 1990:486 (syn. of Trematolobelia macrostachys); Lammers
2009:138 (RESURRECT/Mo, KEY, DESCR)
O
Wagner et al. 1990:488 (O); Lammers 2009:137 (KEY, DESCR)
Mo L
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 54
Scientific Name
Dicot
Cannabaceae
Cannabis sativa L. ssp. indica (Lam.) E.Small &
Cronquist
Status
Kl Le Ni K
?
O Mo L
M Ka H
Ni? K? O? Mo? L? M? Ka? H?
Citation
Wagner et al. 1990:490 (Ni?, K?, O?, Mo?, L?, M?, Ka?, H? [widely
cultivated, but no naturalized vouchers], DESCR); Staples & Herbst
2005:219 (DESCR)
Capparaceae
Capparis mollicella Standl.
nat
Capparis sandwichiana DC.
end
Cleome gynandra L.
nat
Cleome spinosa Jacq.
ind?
O
Kl
Ni
Lau & Frohlich 2012:12 (NEWNAT/O, DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:492 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:219 (DESCR)
K
O Mo L
M
H
Wagner et al. 1990:493 (K, O, Mo, L, M, H)
O
M
Ni
Wagner et al. 1990:494 (Ni, O, M) [Note: not collected since 1864–1865]
Caprifoliaceae
Lonicera japonica Thunb.
nat
K
O
M
H
Wagner et al. 1990:495 (K, H); Starr et al. 2002:19 (EM); Staples & Herbst
2005:220 (KEY), 221 (DESCR); Frohlich & Lau 2008:5 (O)
Sambucus mexicana C.Presl ex DC.
nat
K
O
M
H
Wagner et al. 1990:495 (K, O, EM, H); Oppenheimer & Bartlett 2002:5
(WM); Staples & Herbst 2005:220 (KEY), 222 (DESCR)
nat
K
O Mo L
M
H
Wagner et al. 1990:497 (K, Mo, H); Oppenheimer & Bartlett 2000:3 (EM,
WM); Staples & Herbst 2005:223 (DESCR); Oppenheimer 2010:34 (L);
Oppenheimer 2011:6 (O)
M
H
Wagner et al. 1990:502 (M, H)
M
H
Wagner et al. 1990:503 (K, O, Mo, L, M, H)
Caricaceae
Carica papaya L.
Caryophyllaceae
Arenaria serpyllifolia L.
nat
Cerastium fontanum Baumg. ssp. triviale (Link) Jalas
nat
Cerastium glomeratum Thuill.
nat
Dianthus armeria L. ssp. armeria
nat
K
O Mo L
M
Starr et al. 2004:21 (NEWNAT/EM)
H
Wagner et al. 1990:503 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 55
Scientific Name
Dicot
Caryophyllaceae
Status
Drymaria cordata (L.) Willd. ex Roem. & Schult. var.
pacifica M.Mizush.
nat
Petrorhagia velutina (Guss.) P.W.Ball & Heywood
nat
Polycarpon tetraphyllum (L.) L.
nat
Sagina decumbens (Elliott) Torr. & A.Gray ssp.
occidentalis (S.Watson) G.E.Crow
nat
Sagina japonica (Sw.) Ohwi
nat
Sagina procumbens L.
nat
Kl Le Ni K
K
O Mo L
M Ka H
Citation
O Mo L
M
H
Wagner et al. 1990:505 (K, O, Mo, M, H); Oppenheimer 2007:21 (L)
O
M
H
Wagner et al. 1990:505 (M, H); Frohlich & Lau 2012:32 (O)
M Ka
H
Wagner et al. 1990:506 (K, Mo, L, M, Ka, H)
H
Wagner et al. 1990:506 (H)
K
Mo L
O?
Wagner et al. 1990:506 (O? [single coll. in 1985 at Honolulu International
Airport])
L
M
Oppenheimer 2003:9 (NEWNAT/L, EM, WM)
M?
Wagner et al. 1990:506 (EM? [reported by Hillebrand (1888) from
‘Ulupalakua, not collected since], DESCR)
Sagina subulata (Sw.) K.Presl
?
Schiedea adamantis H.St.John
end
Schiedea amplexicaulis H.Mann
end
Ni? K
Schiedea apokremnos H.St.John
end
K
Wagner et al. 1990:511 (K); Wagner et al. 2005:63 (KEY, DESCR)
Schiedea attenuata W.L.Wagner, Weller & A.K.Sakai
end
K
Wagner et al. 1994:187 (SPNOV/K, DESCR); Wagner et al. 2005:45 (KEY,
DESCR)
Schiedea diffusa A.Gray ssp. diffusa
end
Schiedea diffusa A.Gray ssp. macraei (Sherff)
W.L.Wagner & Weller
end
O
Wagner et al. 1990:510 (O); Wagner et al. 2005:144 (KEY, DESCR)
Wagner et al. 1990:511 (Ni? [Remy coll. from 1851–1855 cites either Ni‘ihau
or Kaua‘i], K); Wagner et al. 2005:71 (KEY, DESCR)
Mo
M
Wagner et al. 1990:511 (Mo, EM, H [as Schiedea diffusa]); Wagner et al.
2005:103 (KEY, DESCR)
H
Wagner et al. 1990:511 (as Schiedea diffusa var. macraei, syn. of S. diffusa);
Wagner et al. 2005:106 (RESURRECT/H, COMBNOV & STATNOV:Syn.
S. diffusa var. macraei, KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 56
Scientific Name
Dicot
Caryophyllaceae
Status
Schiedea globosa H.Mann
end
Schiedea haleakalensis O.Deg. & Sherff
end
Schiedea hawaiiensis Hillebr.
end
Schiedea helleri Sherff
end
Schiedea hookeri A.Gray
end
Schiedea implexa (Hillebr.) Sherff
Kl Le Ni K
O Mo L
O Mo L
M Ka H
M
H
M
Wagner et al. 1990:512 (O, Mo, EM, WM); Herbst & Wagner 1999:18 (H);
Wagner et al. 2005:122 (L, KEY, DESCR)
Wagner et al. 1990:512 (EM); Wagner et al. 2005:119 (KEY, DESCR)
H
K
Citation
Wagner et al. 1990:511 (syn. of Schiedea diffusa); Herbst & Wagner 1999:18
(RESURRECT/H, DESCR); Wagner et al. 2005:92 (KEY, DESCR)
Wagner et al. 1990:514 (K); Lorence et al. 1995:30 (REDISCOVER/K);
Wagner et al. 2005:39 (KEY, DESCR)
O
M?
Wagner et al. 1990:514 (O, EM? [a single pre-1920 coll. from Haleakalä]);
Wagner et al. 2005:111 (KEY, DESCR)
end
M
Wagner et al. 1990:514 (EM); Wagner et al. 2005:87 (KEY, DESCR)
Schiedea jacobii W.L.Wagner, Weller & A.C.Medeiros
end
M
Wagner, Weller et al. 1999:284 (SPNOV/EM, DESCR); Wagner et al.
2005:90 (KEY, DESCR)
Schiedea kaalae Wawra
end
Schiedea kauaiensis H.St.John
end
Schiedea kealiae Caum & Hosaka
end
Schiedea laui W.L.Wagner & Weller
end
Schiedea ligustrina Cham. & Schltdl.
end
Schiedea lychnoides Hillebr.
end
Schiedea lydgatei Hillebr.
end
O
Wagner et al. 1990:515 (O); Wagner et al. 2005:84 (KEY, DESCR)
K
St. John 1988a:177 (SPNOV/K, DESCR); Russell & Bruegmann
1996a:53115; Wagner et al. 2005:74 (KEY, DESCR)
O
Wagner et al. 1990:515 (O); Wagner et al. 2005:107 (KEY, DESCR)
Mo
O
Wagner et al. 2005:82 (SPNOV/Mo, KEY, DESCR)
Wagner et al. 1990:515 (O); Wagner et al. 2005:140 (KEY, DESCR)
K
Wagner et al. 1990:500 (K [as Alsinidendron lychnoides]); Wagner et al.
2005:51 (NOM:Syn. A. lychnoides, KEY, DESCR)
Mo
Wagner et al. 1990:516 (Mo); Wagner et al. 2005:134 (KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 57
Scientific Name
Dicot
Caryophyllaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Schiedea mannii H.St.John
end
Wagner et al. 1990:516 (O); Wagner et al. 2005:130 (KEY, DESCR)
Schiedea membranacea H.St.John
end
Schiedea menziesii Hook.
end
Schiedea nuttallii Hook.
end
O Mo
Schiedea obovata (Sherff) W.L.Wagner & Weller
end
O
Wagner et al. 1990:501 (O [as Alsinidendron obovatum]); Wagner et al.
2005:57 (COMBNOV:Syn. A. obovatum, KEY, DESCR)
Schiedea pentandra W.L.Wagner & E.M.Harris
end
O
Wagner et al. 1990:519 (O [as Schiedea pubescens var. purpurascens]);
Wagner & Harris 2000:159 (SPNOV/O [S. pubescens var. purpurascens, an
invalid name], DESCR); Wagner et al. 2005:96 (KEY, DESCR)
Schiedea perlmanii W.L.Wagner & Weller
end
Schiedea pubescens Hillebr.
end
Schiedea salicaria Hillebr.
end
Schiedea sarmentosa O.Deg. & Sherff
end
Schiedea spergulina A.Gray
end
K
Wagner et al. 1990:520 (K, consisting of vars. spergulina & leiopoda);
Lorence et al. 1995:31 (REDISCOVER/K [var. leiopoda]); Wagner et al.
2005:127 (NOM:Syn. S. spergulina var. leiopoda, KEY, DESCR)
Schiedea stellarioides H.Mann
end
K
Wagner et al. 1990:520 (K); Lorence et al. 1995:31 (REDISCOVER/K);
Wagner et al. 2005:67 (KEY, DESCR)
K
Wagner et al. 1990:517 (K); Wagner et al. 2005:41 (KEY, DESCR)
L
M
Wagner et al. 1990:517 (Mo, L, WM); Wagner et al. 2005:114 (remove Mo
[due to resurrection of Schiedea sarmentosa from synonymy], KEY, DESCR)
M
Wagner et al. 1990:517 (K, O); Wagner et al. 2005:77 (add Mo, M(WM?),
remove K [population newly named Schiedea perlmanii], KEY, DESCR)
K
Wagner et al. 2005:71 (SPNOV/K, KEY, DESCR)
Mo L
Mo
M
Wagner et al. 1990:519 (Mo, L, M [as Schiedea pubescens var. pubescens], O
[as var. purpurascens]); Wagner et al. 2005:99 (remove O [var. purpurascens
an invalid name, renamed S. pentandra], KEY, DESCR)
M
Wagner et al. 1990:519 (WM); Wagner et al. 2005:137 (KEY, DESCR)
Wagner et al. 1990:517 (syn. of Schiedea menziesii); Bruegmann
1996:53131/Wagner et al. 2005:116 (RESURRECT/Mo, KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 58
Scientific Name
Dicot
Caryophyllaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Schiedea trinervis (H.Mann) Pax & K.Hoffm.
end
Schiedea verticillata F.Br.
end
Schiedea viscosa H.Mann
end
Silene alexandri Hillebr.
end
Silene armeria L.
nat
M
Starr & Starr 2012:88 (NEWNAT/EM, DESCR)
Silene cryptopetala Hillebr.
end
M
Wagner et al. 1990:522 (EM)
Silene degeneri Sherff
end
M
Wagner et al. 1990:522 (EM)
Silene gallica L.
nat
Silene hawaiiensis Sherff
end
Silene lanceolata A.Gray
end
Silene perlmanii W.L.Wagner, D.R.Herbst & Sohmer
end
Silene struthioloides A.Gray
end
Spergula arvensis L.
nat
K
Spergularia marina (L.) Griseb.
nat
Stellaria media (L.) Vill.
nat
Vaccaria hispanica (Mill.) Rauschert
?
O
Citation
Wagner et al. 1990:501 (O [as Alsinidendron trinerve]); Wagner et al.
2005:60 (NOM:Syn. A. trinerve, KEY, DESCR)
Only in Northwestern Hawaiian Islands; see NWHI checklist
K
Wagner et al. 1990:501 (K [as Alsinidendron viscosum]); Lorence et al.
1995:30 (REDISCOVER/K); Wagner et al. 2005:54 (NOM:Syn. A.
viscosum, KEY, DESCR)
Mo
K
K
O Mo L
Wagner et al. 1990:522 (Mo)
M
O Mo L
H
Wagner et al. 1990:522 (K, O, Mo, L, M, H)
H
Wagner et al. 1990:523 (H)
H
Wagner et al. 1990:523 (K, Mo, L, H); Hawley & Pyle 1995:1 (O)
O
Wagner et al. 1990:523 (O)
M
H
Wagner et al. 1990:525 (EM, H)
O
M
H
Wagner et al. 1990:525 (K, O, M, H)
K
O Mo
M
K
O Mo L
M
Mo?
Wagner et al. 1990:526 (K, O, Mo, M)
H
Wagner et al. 1990:526 (K, O, L, M, H); Oppenheimer 2010:34 (Mo)
Hughes 1995:3 (Mo? [single spm. seen was collected, Kamiloloa Heights])
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 59
Scientific Name
Dicot
Casuarinaceae
Status
{Casuarina cunninghamiana x Casuarina glauca}
nat
Casuarina equisetifolia L.
nat
Casuarina glauca Sieber ex Spreng.
Kl Le Ni K
O Mo L
K
Ni
M Ka H
M
Citation
Lorence et al. 1995:31 (NEWNAT/K); Oppenheimer & Bartlett 2002:5
(WM); Staples & Herbst 2005:228 (KEY [as to Casuarina cunninghamiana]),
230 (DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:528 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:228 (KEY), 229 (DESCR)
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:529 (K, O, Mo, L, M, Ka, H); Oppenheimer & Bartlett
2000:3 (WM, H); Staples & Herbst 2005:228 (KEY), 230 (DESCR);
Oppenheimer 2008:26 (Mo); Starr & Starr 2011:24 (Ka); Oppenheimer
2011:7 (K)
nat
K
O
H
Wagner et al. 1990:530 (K, O, H); Staples & Herbst 2005:230 (DESCR)
Cecropiaceae
Cecropia obtusifolia Bertol.
Celastraceae
Catha edulis (Vahl) Forssk. ex Endl.
nat
Perrottetia sandwicensis A.Gray
end
O
K
Frohlich & Lau 2012:32 (NEWNAT/O, DESCR)
O Mo L
M
H
Wagner et al. 1990:531 (K, O, Mo, L, M, H)
O
M
H
Wagner et al. 1990:533 (M, H); Staples et al. 2002:7 (O); Oppenheimer &
Bartlett 2002:5 (WM); Staples & Herbst 2005:233 (DESCR)
Ceratophyllaceae
Ceratophyllum demersum L.
nat
Chenopodiaceae
Atriplex canescens (Pursh) Nutt.
nat
H
Staples et al. 2003:10 (NEWNAT/H, DESCR)
Atriplex eardleyae Aellen
nat
H
Wagner et al. 1990:534 (H) [Note: single coll. in 1984 from Pu‘uhonua o
Hönaunau National Historical Park]
Atriplex lentiformis (Torr.) S.Watson
nat
Atriplex maximowicziana Makino
nat
Mo
Wagner et al. 1990:535 (Mo)
H
Wagner et al. 1997:55 (NEWNAT/H, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 60
Scientific Name
Dicot
Chenopodiaceae
Status
Atriplex semibaccata R.Br.
nat
Atriplex suberecta I.Verd.
nat
Bassia hyssopifolia (Pall.) Kuntze
nat
Chenopodium album L.
nat
Chenopodium berlandieri Moq.
?
Chenopodium hircinum Schrad.
nat
Chenopodium murale L.
nat
Chenopodium oahuense (Meyen) Aellen
end
Dysphania ambrosioides (L.) Mosyakin & Clemants
nat
Dysphania carinata (R.Br.) Mosyakin & Clemants
nat
Enchylaena tomentosa R.Br. var. tomentosa
nat
Salicornia virginica L.
nat
Kl Le Ni K
Kl
Ni
O Mo L
M Ka H
K
O Mo L
M Ka
H
Wagner et al. 1990:535 (Ni, K, O, Mo, L, M, Ka, H)
K
O Mo L
M
H
Wagner et al. 1990:535 (O, Mo, L, M, H); Lorence et al. 1995:31 (K)
O Mo
M
M
Wagner et al. 1990:536 (O, M); Staples et al. 2002:7 (Mo); Starr et al.
2002:19 (EM)
H
O?
Kl
Citation
Wagner et al. 1990:537 (EM); Staples et al. 2002:7 (H); Oppenheimer 2003:9
(WM)
Wagner et al. 1990:536 (O? [single coll. in 1914 from Kaimuki vegetable
garden])
M
H
Wagner et al. 1990:538 (EM, H) [Note: collected twice—Haleakalä Crater,
Maui in 1909; Parker Ranch, Hawai‘i in 1982]
Le Ni
K
O Mo L
M Ka
H
Wagner et al. 1990:538 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:22 (Le)
Ni
K
O Mo L
M Ka
H
Wagner et al. 1990:538 (Ni, K, O, Mo, L, M, H); Starr et al. 2006:34 (Ka)
K
O
M
H
Wagner et al. 1990:537 (O, L, M, H [as Chenopodium ambrosioides]);
Staples et al. 2002:7 (K); Staples & Herbst 2005:234 (DESCR); Clemants &
Mosyakin 2003:269 (KEY), 270 (NOM:Syn. C. ambrosioides,
DESCR)/Wagner et al. 2012:28
K
O Mo L
M Ka
H
Wagner et al. 1990:537 (Ni, Mo, L, M, Ka, H [as Chenopodium carinatum]);
Lorence & Flynn 1997:10 (K); Staples et al. 2003:10 (O); Wood & LeGrande
2006:22 (Le); Clemants & Mosyakin 2003:268 (KEY), 274 (NOM:Syn. C.
carinatum, DESCR)/Wagner et al. 2012:28
Le Ni
L
M
Imada et al. 2000:11 (NEWNAT/M, DESCR)
Only in Northwestern Hawaiian Islands; see NWHI checklist
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 61
Scientific Name
Dicot
Chenopodiaceae
Salsola tragus L.
Status
Kl Le Ni K
O Mo L
nat
O Mo
nat
O
M Ka H
M Ka
H
Citation
Wagner et al. 1990:540 (H [as Salsola kali]); Wagner et al. 1997:55 (NOM:
S. kali misapplied); Herbst & Wagner 1999:19 (Ka); Oppenheimer & Bartlett
2002:5 (EM); Herbst et al. 2004:5 (O); Starr et al. 2006:34 (Mo); Starr &
Starr 2012:88 (WM)
Chrysobalanaceae
Chrysobalanus icaco L.
Staples & Herbst 2005:236 (DESCR); Lau & Frohlich 2012:13
(NEWNAT/O, DESCR)
Clethraceae
Clethra lanata M.Martens & Galeotti
nat
K
Frohlich & Lau 2012:33 (NEWNAT/K, DESCR)
Calophyllum inophyllum L.
pol
K
O Mo
M
H
Wagner et al. 1990:542 (K, O, Mo, M, H); Staples & Herbst 2005:237 (KEY,
DESCR)
Clusia rosea Jacq.
nat
K
O Mo
M
H
Wagner et al. 1990:542 (K, O, H); Oppenheimer & Bartlett 2000:3 (EM,
WM); Staples & Herbst 2005:237 (KEY), 238 (DESCR); Daehler & Baker
2006:12 (O); Wysong et al. 2007:3 (Mo)
Hypericum canariense L.
nat
Hypericum gramineum G.Forst.
nat
H
Wagner et al. 1990:544 (H)
Hypericum kouytchense H.Lév.
?
H?
Wagner et al. 1990:543 (H? [sometimes escaping cultivation, but not
naturalized])
H
Wagner et al. 1990:544 (Mo, M, H [as Hypericum mutilum]); Lorence et al.
1995:32 (K); Robson 1990:116/Wagner & Herbst 1995:18 (NOM: Hawaiian
material is subsp. mutilum); Starr & Starr 2012:88 (WM)
Clusiaceae
M
K
Mo
M
Wagner et al. 1990:544 (EM)
Hypericum mutilum L. ssp. mutilum
nat
Hypericum parvulum Greene
nat
H
Wagner et al. 1990:544 (H)
Hypericum perforatum L.
nat
H
Wagner et al. 1990:546 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 62
Scientific Name
Dicot
Combretaceae
Conocarpus erectus L.
Terminalia bellirica (Gaertn.) Roxb.
Status
Kl Le Ni K
nat
K
?
O Mo L
O
L
M Ka H
M
H
O?
K
Citation
Wagner et al. 1990:547 (O, L, M); Lorence & Flynn 1997:10 (K); Staples &
Herbst 2005:239 (KEY), 242 (DESCR); Parker & Parsons 2012a:58 (H)
Staples & Herbst 2005:239, 243 (KEY), 244 (DESCR); Daehler & Baker
2006:14 (O? [adventive], DESCR)
Terminalia catappa L.
nat
O Mo L
M
Terminalia melanocarpa F.Muell.
nat
Terminalia myriocarpa Van Heurck & Müll.Arg.
nat
K
O
M
Argyreia nervosa (Burm.f.) Bojer
nat
K
O
M
Bonamia menziesii A.Gray
end
K
O Mo L
M
Convolvulus arvensis L.
nat
O Mo
M
Convolvulus erubescens Sims
nat
O
Cressa truxillensis Kunth
ind
O Mo
Dichondra micrantha Urb.
nat
O
Evolvulus alsinoides (L.) L.
nat
H
Wagner et al. 1990:547 (K, O, M, H); Oppenheimer 2003:9 (Mo); Staples &
Herbst 2005:239, 243 (KEY), 244 (DESCR); Oppenheimer 2008:26 (L)
H
Staples & Herbst 2005:239, 243 (KEY), 244 (DESCR); Parker & Parsons
2012a:58 (NEWNAT/H)
H
Wagner et al. 1990:548 (K, M, H); Herbarium Pacificum Staff 1998:9 (O);
Staples & Herbst 2005:239, 243 (KEY), 245 (DESCR)
Convolvulaceae
Ipomoea Xleucantha Jacq.
?
Lorence et al. 1995:32 (NEWNAT/K, DESCR); Oppenheimer 2004:10 (EM);
Staples & Herbst 2005:245 (KEY), 246 (DESCR); Lau & Frohlich 2012:13
(O)
H
Wagner et al. 1990:550 (K, O, Mo, L, M, H); Staples 2009:649 (name
conservation proposal); Brummitt 2011 (recommendation for conservation)
Wagner et al. 1990:552 (O, M); Imada et al. 2008:12 (Mo)
Herbst 1998:2 (NEWNAT/O, DESCR)
Ka
L
O?
Wagner et al. 1990:552 (O, Mo, Ka)
M
Oppenheimer & Bartlett 2002:6 (NEWNAT/O, EM, WM, DESCR); Staples
& Herbst 2005:245 (KEY), 246 (DESCR); Oppenheimer 2010:34 (L)
M
Oppenheimer 2004:10 (NEWNAT/WM)
Wagner et al. 1990:560 (O? [single coll. ca. 1950, not naturalized])
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 63
Scientific Name
Dicot
Convolvulaceae
Status
Ipomoea alba L.
nat
Ipomoea aquatica Forssk.
nat
Ipomoea batatas (L.) Lam.
pol
Ipomoea cairica (L.) Sweet
nat?
Ipomoea carnea Jacq. ssp. fistulosa (Mart. ex Choisy)
D.F.Austin
nat
Ipomoea hederifolia L.
nat
Ipomoea imperati (Vahl) Griseb.
ind
Ipomoea indica (Burm.) Merr.
ind
Ipomoea littoralis Blume
ind?
Ipomoea obscura (L.) Ker Gawl.
nat
Ipomoea ochracea (Lindl.) G.Don
nat
Ipomoea pes-caprae (L.) R.Br. ssp. brasiliensis (L.)
Ooststr.
ind
Kl Le Ni K
K
Kl
Ni
O Mo L
M Ka H
O Mo L
M
H
Wagner et al. 1990:554 (K, O, Mo, L, M, H)
O
M
H
Wagner et al. 1990:555 (O, M); Imada et al. 2000:11 (H); Staples & Herbst
2005:245, 247 (KEY), 247 (DESCR)
K
O
M
H
Wagner et al. 1990:555 (K, O, H); Oppenheimer 2003:10 (WM); Staples &
Herbst 2005:245, 247 (KEY), 248 (DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:555 (Ni, K, O, Mo, L, Ka, H); Oppenheimer & Bartlett
2000:4 (WM); Starr et al. 2006:34 (EM)
O
Staples & Herbst 2005:245, 247 (KEY), 248 (DESCR); Lau & Frohlich
2012:13 (NEWNAT/O, DESCR)
H
Kl
Citation
Wagner et al. 1990:556 (H); Staples & Herbst 2005:245, 247 (KEY), 248
(DESCR)
Ni
K
O Mo
M
Ni
K
O Mo L
M Ka
H
Wagner et al. 1990:556 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:245, 247 (KEY), 249 (DESCR)
O Mo
M
H?
Wagner et al. 1990:557 (O, Mo, M, H? [reported by Hillebrand (1888) from
Hilo, but unvouchered])
K
O Mo L
M
H
Wagner et al. 1990:557 (K, O, M, H); Oppenheimer 2003:10 (Mo); Staples &
Herbst 2005:245, 247 (KEY), 250 (DESCR); Oppenheimer 2007:22 (L)
K
O
M
H
Wagner et al. 1990:559 (K, O, M, H); Staples & Herbst 2005:245, 247
(KEY), 250 (DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:559 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:245, 247 (KEY), 249 (DESCR); Wood & LeGrande 2006:22 (Le?
[reported by Caum in 1936 but unvouchered])
Kl Le? Ni
Wagner et al. 1990:556 (Ni, K, O, Mo, M)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 64
Scientific Name
Dicot
Convolvulaceae
Status
Ipomoea triloba L.
nat
Ipomoea tuboides O.Deg. & Ooststr.
end
Ipomoea violacea L.
Kl Le Ni K
O Mo L
M Ka H
Citation
K
O Mo L
M
H
Wagner et al. 1990:559 (K, O, M); Oppenheimer & Bartlett 2002:6 (H);
Oppenheimer 2006:11 (Mo); Oppenheimer 2007:22 (L)
K
O Mo L
M Ka
H
Wagner et al. 1990:560 (Ni, K, O, Mo, L, M, Ka, H)
nat
O Mo
M
H
Wagner et al. 1990:560 (O, Mo, M, H)
Jacquemontia pentantha (Jacq.) G.Don
nat
O
Jacquemontia sandwicensis A.Gray
end
Merremia aegyptia (L.) Urb.
Ni
Kl Le Ni
Wagner et al. 1990:562 (O); Staples & Herbst 2005:245 (KEY), 251
(DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:562 (Ni, K, O, Mo, L, M, Ka, H [as Jacquemontia
ovalifolia subsp. sandwicensis]); Staples & Herbst 2005:245 (KEY), 251
(DESCR [as J. ovalifolia subsp. sandwicensis]); Wood & LeGrande 2006:22
(Le); Namoff et al. 2010 (NOM:Syn. Jacquemontia ovalifolia subsp.
sandwicensis)
nat?
K
O Mo L
M Ka
H
Wagner et al. 1990:563 (K, O, Mo, L, M, Ka, H); Wood 2012:93 (KEY)
Merremia peltata (L.) Merr.
nat
K
Merremia tuberosa (L.) Rendle
nat
K
Merremia umbellata (L.) Hallier f.
nat
Poranopsis paniculata (Roxb.) Roberty
nat
Stictocardia tiliifolia (Desr.) Hallier f.
nat
Wood 2012:93 (NEWNAT/K, KEY)
O Mo L
M
H
O
K
Wagner et al. 1990:563 (K, O, M, H); Staples & Herbst 2005:245 (KEY),
251 (DESCR); Oppenheimer 2006:11 (Mo); Oppenheimer 2007:22 (L);
Wood 2012:93 (KEY)
Wagner et al. 1990:563 (O? [cultivated and perhaps escaping]); Imada et al.
2000:11 (NEWNAT/O); Staples & Herbst 2005:245 (KEY), 251 (DESCR);
Wood 2012:93 (KEY)
O Mo
M
H
Starr et al. 2004:21 (NEWNAT/EM, DESCR); Staples & Herbst 2005:245
(KEY), 251 (DESCR); Parker & Parsons 2012a:58 (H)
M
H
Wagner et al. 1990:564 (K, O, Mo, M, H); Staples & Herbst 2005:245
(KEY), 251 (DESCR)
Cornaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 65
Scientific Name
Dicot
Cornaceae
Cornus kousa Buerg. ex Hance
Status
Kl Le Ni K
O Mo L
M Ka H
nat
Citation
H
Pratt & Bio 2012:75 (NEWNAT/H)
H
Wagner et al. 1990:566 (K, O, Mo, H)
Corynocarpaceae
Corynocarpus laevigatus J.R.Forst. & G.Forst.
nat
K
O Mo
Crassula multicava Lem.
nat
K
O
Crassula ovata (Mill.) Druce
nat
K
Crassula sieberiana (Schult.) Druce
nat
H
Wagner et al. 1990:567 (H)
Kalanchoe beauverdii Raym.-Hamet
?
H?
Wagner et al. 1990:567 (H? [as Kalanchoe beauverdii; single coll. in 1987 as
escape on Miloli‘i-Honomalino Trail], DESCR); Staples et al. 2002:8, 9
(NOM:Syn. K. beauverdii = Bryophyllum beauverdii, KEY); Mort et al.
2001:83/Wagner & Herbst 2003:21 (taxa segregated as Bryophyllum are
nested within Kalanchoe, = K. beauverdii); Staples & Herbst 2005:253
(KEY), 255 (DESCR [as Bryophyllum beauverdii])
Crassulaceae
Lorence et al. 1995:33 (NEWNAT/K, DESCR); Staples & Herbst 2005:253
(KEY), 255 (DESCR); Lau & Frohlich 2012:13 (O)
Lorence et al. 1995:32 (NEWNAT/K, DESCR); Staples & Herbst 2005:253
(KEY), 256 (DESCR)
Kalanchoe beharensis Drake
nat
O
Staples & Herbst 2005:253, 258 (KEY), 258 (DESCR); Lau & Frohlich
2012:14 (NEWNAT/O, DESCR)
Kalanchoe crenata (Andrews) Haw.
nat
O
Herbst et al. 2004:6 (NEWNAT/O, DESCR); Staples & Herbst 2005:253,
258 (KEY), 259 (DESCR)
Kalanchoe daigremontiana Raym.-Hamet & H.Perrier
nat
K
O
L
M
H
Lorence et al. 1995:33 (NEWNAT/K [as Kalanchoe daigremontiana],
DESCR); Oppenheimer & Bartlett 2002:6 (L); Staples et al. 2002:8, 9 (O,
KEY, NOM:Syn. K. daigremontiana = Bryophyllum daigremontianum); Mort
et al. 2001:83/Wagner & Herbst 2003:21 (taxa segregated as Bryophyllum
are nested within Kalanchoe, = K. daigremontiana); Staples & Herbst
2005:253 (KEY), 254 (DESCR [as Bryophyllum daigremontianum]); Starr et
al. 2006:34 (EM); Parker & Parsons 2010:41 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 66
Scientific Name
Dicot
Crassulaceae
Status
Kl Le Ni K
O Mo L
K
M Ka H
M
Citation
Kalanchoe fedtschenkoi Raym.-Hamet & H.Perrier
nat
Lorence et al. 1995:34 (NEWNAT/K [as Kalanchoe fedtschenkoi], DESCR);
Staples et al. 2002:8, 9 (KEY, NOM:Syn. K. fedtschenkoi = Bryophyllum
fedtschenkoi); Mort et al. 2001:83/Wagner & Herbst 2003:21 (taxa
segregated as Bryophyllum are nested within Kalanchoe, = K. fedtschenkoi);
Staples & Herbst 2005:253 (KEY), 254 (DESCR [as Bryophyllum
fedtschenkoi]); Oppenheimer 2006:11 (WM)
Kalanchoe lateritia Engl.
nat
O
Kraus 2003:76 (NEWNAT/O, DESCR); Staples & Herbst 2005:253, 258
(KEY), 260 (DESCR)
Kalanchoe laxiflora Baker
nat
O
Mort et al. 2001:83/Wagner & Herbst 2003:21 (taxa segregated as
Bryophyllum are nested within Kalanchoe); Staples & Herbst 2005:253
(KEY), 255 (DESCR [as Bryophyllum laxiflorum]); Frohlich & Lau 2012:34
(NEWNAT/O, DESCR [as B. laxiflorum])
Kalanchoe pinnata (Lam.) Pers.
nat
K
O Mo L
M
H
Wagner et al. 1990:568 (K, O, Mo, L, M, H [as Kalanchoe pinnata]); Staples
et al. 2002:8, 9 (KEY, NOM:Syn. K. pinnata = Bryophyllum pinnatum);
Mort et al. 2001:83/Wagner & Herbst 2003:21 (taxa segregated as
Bryophyllum are nested within Kalanchoe, = K. pinnata); Staples & Herbst
2005:253 (KEY), 254 (DESCR [as Bryophyllum pinnatum])
Kalanchoe tubiflora (Harv.) Raym.-Hamet
nat
K
O
L
M
H
Wagner et al. 1990:568 (K, O, L, M, H [as Kalanchoe tubiflora]); Staples et
al. 2002:8, 9 (KEY, NOM:Syn. K. tubiflora = Bryophyllum tubiflorum);
Mort et al. 2001:83/Wagner & Herbst 2003:21 (taxa segregated as
Bryophyllum are nested within Kalanchoe, = K. tubiflora); Staples & Herbst
2005:253 (KEY), 254 (DESCR [as Bryophyllum tubiflorum])
O
L
M
H
Wagner et al. 1990:569 (O, H); Starr et al. 1999:11 (EM); Oppenheimer &
Bartlett 2000:4 (WM); Staples & Herbst 2005:262 (KEY), 265 (DESCR);
Oppenheimer 2007:22 (L)
M Ka
H
Wagner et al. 1990:570 (Ni, K, O, L, M, Ka, H); Wysong et al. 2007:4 (Mo)
Cucurbitaceae
Coccinia grandis (L.) Voigt
nat
Cucumis dipsaceus Ehrenb. ex Spach
nat
Ni
K
O Mo L
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 67
Scientific Name
Dicot
Cucurbitaceae
Status
Kl Le Ni K
O Mo L
Lagenaria siceraria (Molina) Standl.
pol
Momordica charantia L.
nat
Sechium edule (Jacq.) Sw.
nat
Sicyos albus (H.St.John) I.Telford
end
Sicyos anunu (H.St.John) I.Telford
end
Sicyos cucumerinus A.Gray
end
Mo
Sicyos erostratus H.St.John
end
O Mo
Sicyos herbstii (H.St.John) I.Telford
end
Sicyos hillebrandii H.St.John
end
Sicyos hispidus Hillebr.
end
Sicyos lanceoloideus (H.St.John) W.L.Wagner &
D.R.Herbst
end
Sicyos lasiocephalus Skottsb.
end
Sicyos macrophyllus A.Gray
end
M Ka H
Citation
Non-naturalized Polynesian introduction; Staples & Herbst 2005:262 (KEY),
267 (DESCR)
Ni
K
O Mo L
M Ka
H
Wagner et al. 1990:572 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:262 (KEY), 268 (DESCR)
K
O
M
H
Wagner et al. 1990:572 (K, O, M, H); Staples & Herbst 2005:262 (KEY),
269 (DESCR)
H
Wagner et al. 1990:574 (H); Wagner & Shannon 1999:442
H
Wagner et al. 1990:575 (L, H); Wagner & Shannon 1999:442
H
Wagner et al. 1990:575 (Mo, M, H); Wagner & Shannon 1999:442
L
K
M
Wagner et al. 1990:575 (O, Mo); Wagner & Shannon 1999:443
Mo?
Wagner et al. 1990:575 (K, Mo? [single disjunct coll.]); Wagner & Shannon
1999:443
M
Mo L
K
M
Wagner et al. 1990:577 (EM); Wagner & Shannon 1999:443
H
O
Wagner et al. 1990:577 (Mo, L, EM, H); Wagner & Shannon 1999:443;
Oppenheimer & Bartlett 2002:6 (WM)
St. John 1987h:52 (SPNOV/K [as Sicyocarya lanceoloidea], DESCR);
Wagner et al. 1990:581 (K, O [as Sicyos sp. A); Wagner & Shannon
1999:444 (O, COMBNOV:Syn. Sicyocarya lanceoloidea)
Mo
M
H
Wagner et al. 1990:577 (H); Herbst 1998:2 (Mo); Wagner & Shannon
1999:444
H
St. John 1987h:52 (SPNOV/EM [as Sicyocarya kipahuluensis], DESCR);
Wagner et al. 1990:578 (H [as S. macrophyllus]); Wagner & Shannon
1999:444 (NOM:Syn. S. kipahuluensis; add EM to distribution)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 68
Scientific Name
Dicot
Cucurbitaceae
Status
Kl Le Ni K
Sicyos maximowiczii Cogn.
end
Le Ni
Sicyos pachycarpus Hook. & Arn.
end
Ni
Sicyos semitonsus H.St.John
end
Sicyos waimanaloensis H.St.John
end
O Mo L
M Ka H
O
K
Citation
Wagner et al. 1990:578 (Ni, O); Wagner & Shannon 1999:444; Wood &
LeGrande 2006:22 (Le)
O Mo L
M Ka
H
Wagner et al. 1990:579 (Ni, K, O, Mo, L, M, Ka, H); Wagner & Shannon
1999:445
Only in Northwestern Hawaiian Islands; see NWHI checklist
K
O Mo
Wagner et al. 1990:581 (K, O, Mo); Wagner & Shannon 1999:446
Cuscutaceae
Cuscuta campestris Yunck.
nat
Cuscuta sandwichiana Choisy
end
O
Ni
K
L
M
H
Wagner et al. 1990:582 (O, H); Oppenheimer 2003:10 (WM); Starr et al.
2004:22 (EM); Oppenheimer 2011:7 (L)
O Mo L
M
H
Wagner et al. 1990:582 (Ni, O, Mo, L, M, H); Herbarium Pacificum Staff
1997:18 (K)
Dilleniaceae
Dillenia suffruticosa (Griff.) Martelli
nat
O
Sato 1998 (NEWNAT/O, DESCR); Staples & Herbst 2005:270 (DESCR)
Dipsacaceae
Scabiosa palaestrina L.
nat
H
Herbst & Wagner 1996:10 (NEWNAT/H, DESCR)
Droseraceae
Drosera anglica Huds.
ind
K
Wagner et al. 1990:583 (K)
Ebenaceae
Diospyros blancoi A.DC.
?
O?
Wagner et al. 1990:584 (O? [forestry planting, sometimes escaping in
Roundtop–Tantalus area]); Staples & Herbst 2005:271 (DESCR)
Wagner et al. 1990:585 (K, O)
Diospyros hillebrandii (Seem.) Fosberg
end
K
O
Diospyros sandwicensis (A.DC.) Fosberg
end
K
O Mo L
M
H
Wagner et al. 1990:585 (K, O, Mo, L, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 69
Scientific Name
Dicot
Elaeagnaceae
Elaeagnus umbellata Thunb.
Status
Kl Le Ni K
O Mo L
M Ka H
nat
Citation
H
Wagner et al. 1990:588 (H)
H
Staples & Herbst 2005:274 (DESCR); Daehler & Baker 2006:7
(NEWNAT/O, DESCR); Parker & Parsons 2012b:67 (H)
Elaeocarpaceae
Elaeocarpus angustifolius Blume
nat
Elaeocarpus bifidus Hook. & Arn.
end
Elaeocarpus serratus L.
O
K
?
O
Wagner et al. 1990:589 (K, O)
O?
Daehler & Baker 2006:14 (O? [adventive], DESCR)
Epacridaceae
Leptecophylla tameiameiae (Cham. & Schltdl.)
C.M.Weiller
ind
K
O Mo L
M
H
Wagner et al. 1990:590 (K, O, Mo, L, M, H [as Styphelia tameiameiae]);
Weiller 1999/Staples et al. 2003:11 (COMBNOV:Syn. S. tameiameiae)
Ericaceae
Erica lusitanica Rudolph
nat
M
Starr et al. 2008:45 (NEWNAT/EM, DESCR)
Vaccinium calycinum Sm.
end
K
O Mo L
M
H
Wagner et al. 1990:593 (K, O, Mo, L, M, H)
Vaccinium dentatum Sm.
end
K
O Mo L
M
H
Wagner et al. 1990:593 (K, O, Mo, L, M, H)
Vaccinium reticulatum Sm.
end
K
O Mo
M
H
Wagner et al. 1990:593 (K, O, Mo, M, H)
K
O Mo L
M
H
Wagner et al. 1990:598 (Ni, K, O, Mo, L, M, H); Herbst & Wagner 1999:20
(author correction for var. katoi [a syn.]); Staples & Herbst 2005:279 (KEY),
281 (DESCR)
H
Herbst 1998:2 (NEWNAT/H)
Euphorbiaceae
Aleurites moluccana (L.) Willd.
pol
Aleurites montana (Lour.) E.H.Wilson
nat
Antidesma platyphyllum H.Mann var. hillebrandii Pax
& K.Hoffm.
end
Ni
K
Wagner et al. 1990:600 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 70
Scientific Name
Dicot
Euphorbiaceae
Status
Antidesma platyphyllum H.Mann var. platyphyllum
end
Antidesma pulvinatum Hillebr.
Kl Le Ni K
K
O Mo L
M Ka H
Citation
O Mo L
M
H
Wagner et al. 1990:600 (O, Mo, L, M, H); Herbarium Pacificum Staff 1996:4
(K)
end
O Mo L
M
H
Wagner et al. 1990:601 (O, Mo, M, H); Oppenheimer 2010:35 (L)
Bischofia javanica Blume
nat
O
Breynia disticha J.R.Forst. & G.Forst. 'Roseopicta'
nat
Claoxylon sandwicense Müll.Arg.
end
Croton guatemalensis Lotsy
nat
Euphorbia albomarginata Torr. & A.Gray
nat
O
Wagner et al. 1990:604 (K, O [as Chamaesyce albomarginata]); Wagner et al.
1997:56 (remove K [spms. reidentified as C. serpens], KEY); Govaerts et al.
2000:667 (NOM:Syn. C. albomarginata); Steinmann & Porter 2002/Yang &
Berry 2011 (molecular support for nesting of Chamaesyce within Euphorbia)
Euphorbia arnottiana Endl.
end
O
Wagner et al. 1990:604 (O [as Chamaesyce arnottiana]); Govaerts et al.
2000:674 (NOM:Syn. C. arnottiana); Steinmann & Porter 2002/Yang &
Berry 2011 (molecular support for nesting of Chamaesyce within Euphorbia)
Euphorbia atrococca A.Heller
end
Euphorbia celastroides Boiss. var. amplectens Sherff
end
K
Wagner et al. 1990:601 (O); Staples & Herbst 2005:278 (KEY), 295
(DESCR)
O Mo L
M
H
Lorence et al. 1995:35 (NEWNAT/EM, DESCR); Staples & Herbst 2005:278
(KEY), 282 (DESCR); Parker & Parsons 2012b:67 (H)
M
H
Wagner et al. 1990:617 (K, O, Mo, L, EM, WM, H)
M
Starr et al. 2004:22 (NEWNAT/EM)
K
Ni
K
Wagner et al. 1990:604 (K [as Chamaesyce atrococca]); Govaerts et al.
2000:676 (NOM:Syn. C. atrococca); Steinmann & Porter 2002/Yang & Berry
2011 (molecular support for nesting of Chamaesyce within Euphorbia)
O Mo L
M Ka
H
Wagner et al. 1990:606 (Ni, K, O, Mo, L, M, Ka, H [as Chamaesyce
celastroides var. amplectens]); Govaerts et al. 2000:692 (NOM:Syn. C.
celastroides var. amplectens); Steinmann & Porter 2002/Yang & Berry 2011
(molecular support for nesting of Chamaesyce within Euphorbia)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 71
Scientific Name
Dicot
Euphorbiaceae
Status
Kl Le Ni K
Ni
O Mo L
M Ka H
Citation
Euphorbia celastroides Boiss. var. celastroides
end
K
Wagner et al. 1990:606 (Ni, K [as Chamaesyce celastroides var.
celastroides]); Govaerts et al. 2000:692 (NOM:Syn. C. celastroides var.
celastroides); Steinmann & Porter 2002/Yang & Berry 2011 (molecular
support for nesting of Chamaesyce within Euphorbia)
Euphorbia celastroides Boiss. var. hanapepensis Sherff
end
K
Wagner et al. 1990:606 (K [as Chamaesyce celastroides var. hanapepensis]);
Govaerts et al. 2000:692 (NOM:Syn. C. celastroides var. hanapepensis);
Steinmann & Porter 2002/Yang & Berry 2011 (molecular support for nesting
of Chamaesyce within Euphorbia)
Euphorbia celastroides Boiss. var. kaenana Sherff
end
Euphorbia celastroides Boiss. var. laehiensis O.Deg.,
I.Deg. & Sherff
end
L
M
Wagner et al. 1990:606 (L, EM [as Chamaesyce celastroides var. laehiensis]);
Govaerts et al. 2000:692 (NOM:Syn. C. celastroides var. laehiensis);
Steinmann & Porter 2002/Yang & Berry 2011 (molecular support for nesting
of Chamaesyce within Euphorbia)
Euphorbia celastroides Boiss. var. lorifolia (A.Gray)
Sherff
end
L
M
Wagner et al. 1990:606 (L, M [as Chamaesyce celastroides var. lorifolia]);
Govaerts et al. 2000:692 (NOM:Syn. C. celastroides var. lorifolia);
Steinmann & Porter 2002/Yang & Berry 2011 (molecular support for nesting
of Chamaesyce within Euphorbia)
Euphorbia celastroides Boiss. var. stokesii (C.N.Forbes)
Sherff
end
Euphorbia celastroides Boiss. var. tomentella (Boiss.)
Oudejans
end
O
Kl
Ni
K
Wagner et al. 1990:606 (O [as Chamaesyce celastroides var. kaenana]);
Govaerts et al. 2000:692 (NOM:Syn. C. celastroides var. kaenana);
Steinmann & Porter 2002/Yang & Berry 2011 (molecular support for nesting
of Chamaesyce within Euphorbia)
Mo
O
Ka
Wagner et al. 1990:606 (Ni, K, Mo, Ka [as Chamaesyce celastroides var.
stokesii]); Govaerts et al. 2000:692 (NOM:Syn. C. celastroides var. stokesii);
Steinmann & Porter 2002/Yang & Berry 2011 (molecular support for nesting
of Chamaesyce within Euphorbia)
Wagner et al. 1990:606 (O [as Chamaesyce celastroides var. tomentella]);
Oudejans 1989:44/Govaerts et al. 2000:692 (COMBNOV:Syn. C.
celastroides var. tomentella); Steinmann & Porter 2002/Yang & Berry 2011
(molecular support for nesting of Chamaesyce within Euphorbia)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 72
Scientific Name
Dicot
Euphorbiaceae
Status
Kl Le Ni K
Euphorbia clusiifolia Hook. & Arn.
end
Euphorbia cyathophora Murray
nat
Euphorbia degeneri Sherff
end
Euphorbia deppeana Boiss.
end
Euphorbia eleanoriae (Lorence & W.L.Wagner)
Govaerts
end
Euphorbia graminea Jacq.
nat
Euphorbia haeleeleana D.R.Herbst
end
K
Euphorbia halemanui Sherff
end
K
Euphorbia herbstii (W.L.Wagner) Oudejans
end
Euphorbia heterophylla L.
nat
O Mo L
M Ka H
O
Ni
Wagner et al. 1990:606 (O [as Chamaesyce clusiifolia]); Govaerts et al.
2000:698 (NOM:Syn. C. clusiifolia); Steinmann & Porter 2002/Yang &
Berry 2011 (molecular support for nesting of Chamaesyce within Euphorbia)
K
O Mo
M
H
Wagner et al. 1990:618 (K, O, Mo, M); Staples & Herbst 2005:278, 285
(KEY), 288 (DESCR); Parker & Parsons 2010:41 (H)
K
O Mo
M
H
Wagner et al. 1990:607 (Ni, K, O, Mo, M, H [as Chamaesyce degeneri]);
Govaerts et al. 2000:709 (NOM:Syn. C. degeneri); Steinmann & Porter
2002/Yang & Berry 2011 (molecular support for nesting of Chamaesyce
within Euphorbia)
O
Wagner et al. 1990:607 (O [as Chamaesyce deppeana]); Govaerts et al.
2000:712 (NOM:Syn. C. deppeana); Steinmann & Porter 2002/Yang & Berry
2011 (molecular support for nesting of Chamaesyce within Euphorbia)
K
Lorence & Wagner 1996:68 (SPNOV/K, DESCR)/Herbst & Wagner 1999:20
[as Chamaesyce eleanoriae]; Govaerts et al. 2000:717 (COMBNOV:Syn. C.
eleanoriae); Steinmann & Porter 2002/Yang & Berry 2011 (molecular
support for nesting of Chamaesyce within Euphorbia)
O
Wagner et al. 1990:618 (O)
O
Wagner et al. 1990:619 (K, O)
Wagner et al. 1990:607 (K [as Chamaesyce halemanui]); Govaerts et al.
2000:742 (NOM:Syn. C. halemanui); Steinmann & Porter 2002/Yang &
Berry 2011 (molecular support for nesting of Chamaesyce within Euphorbia)
O
Ni
Citation
K
O
Wagner et al. 1990:609 (O [as Chamaesyce herbstii]); Oudejans
1989:46/Govaerts et al. 2000:745 (COMBNOV:Syn. C. herbstii); Steinmann
& Porter 2002/Yang & Berry 2011 (molecular support for nesting of
Chamaesyce within Euphorbia)
L
M Ka
H
Wagner et al. 1990:619 (Ni, K, O, L, M, Ka, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 73
Scientific Name
Dicot
Euphorbiaceae
Status
Euphorbia hirta L.
nat
Euphorbia hypericifolia L.
Kl Le Ni K
Le Ni
O Mo L
M Ka H
Citation
K
O Mo L
M Ka
H
Wagner et al. 1990:609 (Ni, K, O, Mo, L, M, Ka, H [as Chamaesyce hirta]);
Govaerts et al. 2000:320 (NOM:Syn. C. hirta); Wood & LeGrande 2006:23
(Le); Steinmann & Porter 2002/Yang & Berry 2011 (molecular support for
nesting of Chamaesyce within Euphorbia)
nat
K
O Mo L
M
H
Wagner et al. 1990:609 (K, O, M, H [as Chamaesyce hypericifolia]);
Govaerts et al. 2000:320 (NOM:Syn. C. hypericifolia); Oppenheimer 2003:11
(Mo, L); Steinmann & Porter 2002/Yang & Berry 2011 (molecular support
for nesting of Chamaesyce within Euphorbia)
Euphorbia hyssopifolia L.
nat
K
O Mo L
M
H
Wagner et al. 1990:610 (K, O, M, H [as Chamaesyce hyssopifolia]);
Herbarium Pacificum Staff 1999:3 (L); Govaerts et al. 2000:320 (NOM:Syn.
C. hyssopifolia); Oppenheimer 2003:11 (Mo); Steinmann & Porter
2002/Yang & Berry 2011 (molecular support for nesting of Chamaesyce
within Euphorbia)
Euphorbia kuwaleana O.Deg. & Sherff
end
Euphorbia lactea Haw.
nat
K
Euphorbia leucocephala Lotsy
nat
K
Euphorbia maculata L.
nat
Euphorbia multiformis Gaudich. ex Hook. & Arn. var.
microphylla Boiss.
end
O
Wagner et al. 1990:610 (O [as Chamaesyce kuwaleana]); Govaerts et al.
2000:764 (NOM:Syn. C. kuwaleana); Steinmann & Porter 2002/Yang &
Berry 2011 (molecular support for nesting of Chamaesyce within Euphorbia)
O
Staples & Herbst 2005:278, 284 (KEY), 285 (DESCR); Frohlich & Lau
2012:35 (NEWNAT/K, O, DESCR)
Staples & Herbst 2005:278, 285 (KEY), 286 (DESCR); Frohlich & Lau
2012:35 (NEWNAT/K, DESCR)
Only in Northwestern Hawaiian Islands; see NWHI checklist
O Mo
M
H
Wagner et al. 1990:611 (O, Mo, WM, H [as Chamaesyce multiformis var.
microphylla]); Govaerts et al. 2000:783 (NOM:Syn. C. multiformis var.
microphylla); Steinmann & Porter 2002/Yang & Berry 2011 (molecular
support for nesting of Chamaesyce within Euphorbia)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 74
Scientific Name
Dicot
Euphorbiaceae
Status
Kl Le Ni K
O Mo L
O
M Ka H
M
Citation
Euphorbia multiformis Gaudich. ex Hook. & Arn. var.
multiformis
end
Euphorbia olowaluana Sherff
end
Euphorbia peplus L.
nat
Euphorbia prostrata Aiton
nat
Euphorbia remyi A.Gray ex Boiss. var. hanaleiensis
Sherff
end
K
Wagner et al. 1990:614 (K [as Chamaesyce remyi var. hanaleiensis]);
Govaerts et al. 2000:818 (NOM:Syn. C. remyi var. hanaleiensis); Steinmann
& Porter 2002/Yang & Berry 2011 (molecular support for nesting of
Chamaesyce within Euphorbia)
Euphorbia remyi A.Gray ex Boiss. var. kauaiensis
O.Deg. & Sherff
end
K
Wagner et al. 1990:614 (K [as Chamaesyce remyi var. kauaiensis]); Govaerts
et al. 2000:818 (NOM:Syn. C. remyi var. kauaiensis); Steinmann & Porter
2002/Yang & Berry 2011 (molecular support for nesting of Chamaesyce
within Euphorbia)
Euphorbia remyi A.Gray ex Boiss. var. remyi
end
K
Wagner et al. 1990:614 (K [as Chamaesyce remyi var. remyi]); Govaerts et
al. 2000:818 (NOM:Syn. C. remyi var. remyi); Steinmann & Porter
2002/Yang & Berry 2011 (molecular support for nesting of Chamaesyce
within Euphorbia)
Euphorbia rockii C.N.Forbes
end
K
Le
K
O Mo L
O
Wagner et al. 1990:611 (O, M [as Chamaesyce multiformis var.
multiformis]); Govaerts et al. 2000:783 (NOM:Syn. C. multiformis var.
multiformis); Steinmann & Porter 2002/Yang & Berry 2011 (molecular
support for nesting of Chamaesyce within Euphorbia)
M
H
Wagner et al. 1990:611 (WM, H [as Chamaesyce olowaluana]); Govaerts et
al. 2000:793 (NOM:Syn. C. olowaluana); Steinmann & Porter 2002/Yang &
Berry 2011 (molecular support for nesting of Chamaesyce within Euphorbia)
M
H
Wagner et al. 1990:619 (M, H); Lorence et al. 1995:35 (K)
M Ka
H
Wagner et al. 1990:613 (K, O, L, M, Ka, H [as Chamaesyce prostrata]);
Hughes 1995:4 (Mo); Govaerts et al. 2000:324 (NOM:Syn. C. prostrata);
Wood 2012:94 (Le); Steinmann & Porter 2002/Yang & Berry 2011
(molecular support for nesting of Chamaesyce within Euphorbia)
Wagner et al. 1990:614 (O [as Chamaesyce rockii]); Govaerts et al. 2000:821
(NOM:Syn. C. rockii); Steinmann & Porter 2002/Yang & Berry 2011
(molecular support for nesting of Chamaesyce within Euphorbia)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 75
Scientific Name
Dicot
Euphorbiaceae
Status
Kl Le Ni K
O Mo L
Euphorbia serpens Kunth
nat
K
Euphorbia skottsbergii Sherff var. skottsbergii
end
O Mo
Euphorbia skottsbergii Sherff var. vaccinioides Sherff
end
Mo
Euphorbia sparsiflora A.Heller
end
K
Euphorbia thymifolia L.
nat
K
Euphorbia tirucalli L.
nat
K
Flueggea neowawraea W.J.Hayden
end
K
Flueggea virosa (Roxb. ex Willd.) Voigt
nat
M Ka H
M
Citation
Wagner et al. 1997:56 (NEWNAT/K [as Chamaesyce serpens], KEY, NOM:
C. albomarginata misapplied, in part); Oppenheimer 2003:11 (WM);
Govaerts et al. 2000:832 (NOM:Syn. C. serpens); Steinmann & Porter
2002/Yang & Berry 2011 (molecular support for nesting of Chamaesyce
within Euphorbia)
Wagner et al. 1990:615 (O, Mo [as Chamaesyce skottsbergii var.
skottsbergii]); Govaerts et al. 2000:835 (NOM:Syn. C. skottsbergii var.
skottsbergii); Morden & Gregoritza 2005 (molecular evidence presented for
resurrecting C. skottsbergii var. audens as accepted name for var. skottsbergii
populations on Moloka‘i); Steinmann & Porter 2002/Yang & Berry 2011
(molecular support for nesting of Chamaesyce within Euphorbia)
M Ka
Wagner et al. 1990:615 (Mo, EM, Ka [as Chamaesyce skottsbergii var.
vaccinioides]); Govaerts et al. 2000:835 (NOM:Syn. C. skottsbergii var.
vaccinioides); Steinmann & Porter 2002/Yang & Berry 2011 (molecular
support for nesting of Chamaesyce within Euphorbia)
Wagner et al. 1990:615 (K, Mo [as Chamaesyce sparsiflora]); Govaerts et al.
2000:837 (NOM:Syn. C. sparsiflora); Steinmann & Porter 2002/Yang &
Berry 2011 (molecular support for nesting of Chamaesyce within Euphorbia)
O Mo L
O Mo
M
M
M
H
Wagner et al. 1990:615 (O, H [as Chamaesyce thymifolia]); Lorence et al.
1995:35 (K, EM); Oppenheimer & Bartlett 2000:5 (WM); Oppenheimer
2003:11 (Mo, L); Govaerts et al. 2000:848 (NOM:Syn. C. thymifolia);
Steinmann & Porter 2002/Yang & Berry 2011 (molecular support for nesting
of Chamaesyce within Euphorbia)
H
Lorence et al. 1995:35 (NEWNAT/K, DESCR); Staples & Herbst 2005:278,
284 (KEY), 288 (DESCR); Parker & Parsons 2012a:59 (H)
H
Wagner et al. 1990:620 (K, O, Mo, EM, H)
Starr et al. 2004:22 (NEWNAT/EM, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 76
Scientific Name
Dicot
Euphorbiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
M
H
Citation
Homalanthus populifolius Graham
nat
Homalanthus populneus (Geiseler) Pax
nat
Jatropha curcas L.
nat
Jatropha gossypiifolia L.
nat
Jatropha multifida L.
nat
H
Staples & Herbst 2005:279, 289 (KEY), 290 (DESCR); Parker & Parsons
2012a:59 (NEWNAT/H)
Jatropha podagrica Hook.
nat
H
Wagner et al. 1990:621; Staples & Herbst 2005:278, 279, 289 (KEY), 290
(DESCR); Parker & Parsons 2012a:59 (NEWNAT/H)
Macaranga mappa (L.) Müll.Arg.
nat
H
Wagner et al. 1990:624 (O, H); Staples & Herbst 2005:279 (KEY), 291
(DESCR)
Macaranga tanarius (L.) Müll.Arg.
nat
H
Wagner et al. 1990:624 (K, O); Oppenheimer et al. 1999:7 (WM); Parker &
Parsons 2010:41 (H)
Mallotus philippensis (Lam.) Müll.Arg.
nat
O
Wagner et al. 1990:624 (O); Daehler & Baker 2006:15 (O)
Manihot esculenta Crantz
nat
O
Wagner et al. 1990:625 [adventive]; Staples & Herbst 2005:278, 279 (KEY),
291 (DESCR); Frohlich & Lau 2012:36 (NEWNAT/O, DESCR)
Manihot glaziovii Müll.Arg.
nat
Pedilanthus tithymaloides (L.) Poit. ssp. padifolius (L.)
Dressler
nat
O
Frohlich & Lau 2012:35 (NEWNAT/O, DESCR)
M
H
O
K
Wagner et al. 1990:623 (EM); Staples & Herbst 2005:279, 289 (KEY), 289
(DESCR); Parker & Parsons 2012a:59 (H)
Wagner et al. 1990:623 (O)
O
K
Oppenheimer 2004:11 (NEWNAT/EM, H [as Omalanthus populifolius],
DESCR)
O
M
O
M
O
H
Wagner et al. 1990:625 (K, O, M, H); Staples & Herbst 2005:279 (KEY),
291 (DESCR)
Wagner et al. 1990:596 [as Pedilanthus tithymaloides; persisting after
cultivation, but not naturalized]; Staples & Herbst 2005:278 (KEY), 292
(DESCR, for subsp. padifolius); Frohlich & Lau 2012:37 (NEWNAT/O,
DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 77
Scientific Name
Dicot
Euphorbiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Phyllanthus acidus (L.) Skeels
nat
Phyllanthus debilis Klein ex Willd.
nat
K
O Mo L
M
Phyllanthus distichus Hook. & Arn.
end
K
O Mo L
M
Phyllanthus reticulatus Poir.
nat
Phyllanthus tenellus Roxb.
nat
Ricinus communis L.
nat
Sauropus androgynus (L.) Merr.
Ni
Citation
H
Staples & Herbst 2005:279, 293 (KEY), 294 (DESCR); Parker & Parsons
2012a:59 (NEWNAT/H)
H
Wagner et al. 1990:627 (K, O, M, H); Oppenheimer 2003:12 (Mo, L)
Wagner et al. 1990:627 (K, O, Mo, EM, WM); Govaerts & Radcliffe-Smith
1996:176 (COMBNOV: Phyllanthus distichus vars. degeneri & ellipticus
[replacing P. sandwicensis vars. degeneri & ellipticus, already treated as syns.
of P. distichus); Herbst & Wagner 1999:20 (vars. degeneri & ellipticus to be
treated as synonyms of P. distichus)
H
Parker & Parsons 2012b:67 (NEWNAT/H, DESCR)
K
O Mo L
M
H
Wagner et al. 1990:628 (K, O); Oppenheimer et al. 1999:8 (EM, WM);
Oppenheimer & Bartlett 2000:5 (H); Oppenheimer & Bartlett 2002:7 (L);
Oppenheimer 2007:23 (Mo)
K
O Mo L
M Ka
H
Wagner et al. 1990:629 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:278, 279 (KEY), 296 (DESCR)
?
O?
Staples & Herbst 2005:279 (KEY), 296 (DESCR); Frohlich & Lau 2008:5
(NEWNAT/O, DESCR); Wagner et al. 2012:35 (O record adventive)
Fabaceae
Abrus precatorius L.
nat
Acacia aneura F.Muell. ex Benth.
nat
Acacia auriculiformis A.Cunn. ex Benth.
Acacia catechu (L.f.) Willd.
Ni
K
Mo
M
L
H
Wagner et al. 1990:638 (Ni, K, Mo, M, H); Staples & Herbst 2005:299
(KEY), 330 (DESCR)
Wagner & Herbst 1995:19 (NEWNAT/L, DESCR)
?
O?
Frohlich & Lau 2007:9 (NEWNAT/O, DESCR); Wagner et al. 2012:36 (O
record adventive)
nat
O
Staples & Herbst 2005:298, 299, 301 (KEY), 302 (DESCR); Frohlich & Lau
2010:10 (NEWNAT/O, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 78
Scientific Name
Dicot
Fabaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Acacia confusa Merr.
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:641 (K, O, Mo, L, M, Ka, H); Staples & Herbst 2005:298,
299, 301 (KEY), 301 (DESCR)
Acacia koa A.Gray
end
K
O Mo L
M
H
Wagner et al. 1990:641 (K, O, Mo, L, M, H); Staples & Herbst 2005:298,
299, 301 (KEY), 302 (DESCR); Rico-Arce 2007:106 (DESCR)
Acacia koaia Hillebr.
end
K
O? Mo L
M
H
Wagner et al. 1990:641 (Mo, L, M, H [syn. of Acacia koa]); Wagner &
Herbst 1999:1875 (RESURRECT, also K, perhaps O); Rico-Arce 2007:108
(DESCR)
Acacia mangium Willd.
nat
K
O
H
Frohlich & Lau 2008:6 (NEWNAT/O, DESCR); Frohlich & Lau 2012:37
(K); Parker & Parsons 2012b:67 (H)
Acacia mearnsii De Wild.
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:642 (K, O, L, M, H); Hughes 1995:5 (Mo); Starr et al.
2006:35 (Ka)
Acacia melanoxylon R.Br. ex Aiton
nat
K
O? Mo
M
Acacia parramattensis Tindale
nat
Acacia retinodes Schltdl.
?
Adenanthera pavonina L.
nat
Aeschynomene paniculata Willd. ex Vogel
nat
Albizia chinensis (Osbeck) Merr.
nat
Wagner et al. 1990:640 (K?, O?, Mo?, M? [forestry planting, but definitely
naturalizing on EM]); Lorence et al. 1995:36 (NEWNAT/K, DESCR);
Meidell et al. 1998:7 (WM); Staples & Herbst 2005:298, 299, 301 (KEY),
301 (DESCR); Oppenheimer 2010:35 (Mo)
O Mo
K
Wagner et al. 1990:643 (O, Mo)
M?
Starr & Starr 2011:25 (NEWNAT/EM, DESCR); Wagner et al. 2012:36 (EM
record adventive)
M
Wagner & Herbst 1995:19 (NEWNAT/K, DESCR); Staples & Herbst
2005:298 (KEY), 302 (DESCR); Starr & Starr 2011:25 (EM)
Mo
O
Hughes 1995:5 (NEWNAT/Mo)
H
Wagner et al. 1990:644 (O); Herbst 1998:3 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 79
Scientific Name
Dicot
Fabaceae
Status
Kl Le Ni K
Albizia lebbeck (L.) Benth.
nat
Ni
K
Albizia niopoides (Benth.) Burkart
nat
Albizia saponaria (Lour.) Blume ex Miq.
nat
Alysicarpus vaginalis (L.) DC.
nat
Bauhinia glauca (Benth.) Benth. ssp. tenuiflora
(C.B.Clarke) K.Larsen & S.S.Larsen
nat
Bauhinia monandra Kurz
nat
Caesalpinia bonduc (L.) Roxb.
ind
Caesalpinia crista L.
nat
Caesalpinia decapetala (Roth) Alston
nat
Ni
Caesalpinia major (Medik.) Dandy & Exell
nat?
Ni
Cajanus cajan (L.) Huth
nat
K
Calliandra houstoniana (Mill.) Standl. var. calothyrsa
(Meisn.) Barneby
nat
K
Calliandra surinamensis Benth.
nat
O Mo L
O Mo
M Ka H
M
H
Wagner et al. 1990:644 (Ni, K, O, H); Oppenheimer & Bartlett 2002:7
(WM); Staples & Herbst 2005:298 (KEY), 330 (DESCR); Oppenheimer
2007:23 (EM); Oppenheimer 2008:28 (Mo)
O
Lau & Frohlich 2012:14 (NEWNAT/O, DESCR)
K
O
Lorence et al. 1995:36 (NEWNAT/K, DESCR); Lau & Frohlich 2012:15 (O)
K
O Mo
M
H?
O
Ni
Citation
K
Frohlich & Lau 2012:37 (NEWNAT/O, DESCR)
O Mo
M
H
Lorence & Flynn 1999:4 (NEWNAT/H); Staples & Herbst 2005:297, 304
(KEY), 306 (DESCR); Starr & Starr 2011:26 (EM)
M
H
Wagner et al. 1990:647 (Ni, K, O, Mo, EM, H)
O
K
O
Wagner et al. 1990:644 (K, O, M, H? [reported but unvouchered]);
Oppenheimer 2007:23 (Mo)
Frohlich & Lau 2008:6 (NEWNAT/O, DESCR)
L
M
H
Wagner et al. 1990:647 (Ni, K, O, EM, H); Starr et al. 2006:35 (L)
O Mo L
M
H
Wagner et al. 1990:648 (Ni, O, Mo, L, H); Herbst & Wagner 1996:11 (WM)
O
M
H
Wagner et al. 1990:649 (K, O, M, H); Staples & Herbst 2005:300 (KEY),
308 (DESCR)
M
H
Staples & Herbst 2005:299, 308 (KEY), 309 (DESCR); Imada et al. 2008:13
(NEWNAT/L); Starr et al. 2010:64 (EM); Frohlich & Lau 2012:38 (K);
Parker & Parsons 2012b:68 (H)
L
M
Staples & Herbst 2005:299, 308 (KEY), 309 (DESCR); Starr & Starr 2011:26
(NEWNAT/WM, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 80
Scientific Name
Dicot
Fabaceae
Calopogonium caeruleum (Benth.) Hemsl.
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
?
M?
Wagner et al. 1990:630 (M? [introduced in 1946 by Pineapple Research
Institute, no evidence of naturalization], DESCR)
Calopogonium galactioides (Kunth) Benth. ex Hemsl.
nat
M
Starr et al. 2003:27 (NEWNAT/EM, DESCR)
Canavalia cathartica Thouars
nat
Canavalia galeata (Gaudich.) Vogel
end
Canavalia hawaiiensis O.Deg., I.Deg. & J.D.Sauer
end
Canavalia kauaiensis J.D.Sauer
end
Canavalia molokaiensis O.Deg., I.Deg. & J.D.Sauer
end
Canavalia napaliensis H.St.John
end
Canavalia pubescens Hook. & Arn.
end
Canavalia sericea A.Gray
nat
Cassia javanica L. var. indochinensis Gagnep.
nat
Centrosema molle Mart. ex Benth.
nat
K
O Mo
M
H
O
Wagner et al. 1990:651 (O); Wysong et al. 2007:4 (Mo); Wagner et al.
2012:37 (remove Mo, spm. reidentified as Canavalia molokaiensis)
L
M
H
K
Wagner et al. 1990:653 (Mo); Wagner et al. 2012:37 (NOM: Canavalia
galeata misapplied, in part)
K
Wagner et al. 1990:654 (K)
K
K
K
Wagner et al. 1990:651 (L, M, H)
Wagner et al. 1990:653 (K)
Mo
Ni
Wagner et al. 1990:651 (K, O, Mo, M, H); Staples & Herbst 2005:299
(KEY), 310 (DESCR)
L
O
M
M
O
Wagner et al. 1990:654 (Ni, K, L, EM)
H
Wagner et al. 1990:654 (O, WM); Wagner & Herbst 1995:19 (K);
Oppenheimer & Bartlett 2002:7 (EM); Oppenheimer 2003:12 (H); Staples &
Herbst 2005:299 (KEY), 310 (DESCR)
M
Staples & Herbst 2005:298, 311 (KEY), 311 (DESCR); Starr & Starr 2011:26
(NEWNAT/EM, DESCR)
M
Flynn & Lorence 1998:5 (NEWNAT/K [as Centrosema pubescens],
DESCR); Starr et al. 2003:27 (EM); Imada et al. 2008:13 (O); Fantz
1996:330/Wagner & Herbst 2003:24 (NOM:C. pubescens misapplied)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 81
Scientific Name
Dicot
Fabaceae
Status
Chamaecrista nictitans (L.) Moench ssp. patellaria (DC.
ex Collad.) H.S.Irwin & Barneby var. glabrata (Vogel)
H.S.Irwin & Barneby
nat
Clitoria heterophylla Lam.
nat
Clitoria ternatea L.
nat
Coronilla varia L.
?
Kl Le Ni K
Ni
K
O Mo L
O Mo L
O
L
M Ka H
M Ka
Citation
H
Wagner et al. 1990:655 (Ni, K, O, L, M, Ka, H); Hughes 1995:5 (Mo)
H
Staples et al. 2002:9, 10 (NEWNAT/H, KEY, DESCR)
M
Wagner et al. 1990:656 (O [as Clitoria ternata]); Oppenheimer & Bartlett
2000:5 (WM); Staples et al. 2002:10 (KEY); Staples & Herbst 2005:299
(KEY), 313 (DESCR); Starr et al. 2010:64 (L)
M?
Wagner et al. 1990:630 (EM? [single coll. in 1983 below Haleakalä National
Park], DESCR)
Crotalaria assamica Benth.
nat
O Mo
M
Crotalaria berteroana DC.
nat
O
M
Wagner et al. 1990:658 (O, M)
Crotalaria brevidens Benth.
nat
M
Wagner et al. 1990:658 (EM); Oppenheimer 2003:12 (WM)
Crotalaria incana L.
nat
Crotalaria juncea L.
Ni
Wagner et al. 1990:657 (O, H); Starr et al. 2002:20 (EM); Starr et al. 2006:35
(Mo)
O Mo L
M Ka
H
Wagner et al. 1990:660 (Ni, K, O, L, M, Ka, H); Hughes 1995:5 (Mo)
nat
O?
M
H
Wagner et al. 1997:56 (NEWNAT/O? [single coll. in 1962 from Kaläkaua
Ave., likely cultivated], H, DESCR); Staples et al. 2003:11 (EM); Staples &
Herbst 2005:299 (KEY), 331 (DESCR)
Crotalaria lanceolata E.Mey.
nat
O
M
H
Wagner et al. 1990:660 (H); Oppenheimer 2004:12 (EM); Imada 2007:37 (O)
Crotalaria longirostrata Hook. & Arn.
nat
O Mo
M
H
Wagner et al. 1990:660 (O, Mo, EM, H)
Crotalaria micans Link
nat
O
M
H
Wagner et al. 1990:661 (O, M, H)
Crotalaria pallida Aiton
nat
O Mo L
M
H
Wagner et al. 1990:661 (K, O, Mo, L, M, H); Wood 2007:15 (Le)
Crotalaria pumila Ortega
nat
O
M
Le
K
H
K
Wagner et al. 1990:662 (O, EM); Oppenheimer & Bartlett 2000:5 (WM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 82
Scientific Name
Dicot
Fabaceae
Status
Crotalaria retusa L.
nat
Crotalaria sagittalis L.
nat
Crotalaria spectabilis Roth
nat
Crotalaria trichotoma Bojer
nat
Crotalaria verrucosa L.
nat
Cytisus palmensis (H.Christ) Hutch.
nat
Cytisus scoparius (L.) Link
nat
Dalea emarginata (Torr. & A.Gray) Shinners
nat
Delonix regia (Bojer ex Hook.) Raf.
nat
Derris elliptica (Wall.) Benth.
nat
Desmanthus pernambucanus (L.) Thell.
nat
Desmodium barbatum (L.) Benth. & Oerst.
nat
Desmodium cajanifolium (Kunth) DC.
nat
Desmodium heterocarpon (L.) DC. var. strigosum
Meeuwen
nat
Kl Le Ni K
K
Ni
K
O Mo L
O Mo
M Ka H
M
O Mo
H
Wagner et al. 1990:662 (K, O, Mo, M, H)
H
Pratt & Bio 2012:76 (NEWNAT/H, DESCR)
H
Wagner et al. 1990:663 (Ni, K, O, Mo, H)
M
Starr et al. 2006:35 (NEWNAT/EM, DESCR)
O
Wagner & Herbst 1995:19 (NEWNAT/O, DESCR)
M
H
Wagner et al. 1990:663 (M); Staples et al. 2002:10 (H)
H
Wagner et al. 1990:664 (H)
O
Herbarium Pacificum Staff 1996:4 (NEWNAT/O)
Mo?
K
Citation
O Mo L
Wagner et al. 1990:664 (Mo? [undocumented by vouchers]); Staples &
Herbst 2005:298 (KEY), 314 (DESCR)
M Ka
H
Staples & Herbst 2005:299 (KEY), 331 (DESCR); Parker & Parsons
2012a:60 (NEWNAT/H)
H
Wagner et al. 1990:665 (K, O, Mo, M, Ka, H [as Desmanthus virgatus]);
Wagner & Herbst 1995:20 (NOM:Syn. D. virgatus); Oppenheimer 2003:12
(L)
Mo
K
Oppenheimer 2010:35 (NEWNAT/Mo, DESCR)
H
O
Wagner et al. 1990:666 (H); Frohlich & Lau 2012:38 (K)
Wagner et al. 1990:667 (O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 83
Scientific Name
Dicot
Fabaceae
Status
Desmodium heterophyllum (Willd.) DC.
nat
Desmodium incanum DC.
nat
Desmodium intortum (Mill.) Urb.
nat
Desmodium sandwicense E.Mey.
nat
Desmodium tortuosum (Sw.) DC.
Kl Le Ni K
O Mo L
M Ka H
Citation
Mo?
M
H
Wagner & Herbst 1995:20 (NEWNAT/Mo? [cultivated at Plant Materials
Center, Ho‘olehua], EM, DESCR); Herbst & Wagner 1999:20 (H);
Oppenheimer 2003:12 (WM)
K
O Mo L
M
H
Wagner et al. 1990:667 (K, O, M, H); Oppenheimer 2003:12 (Mo, L)
K
O
H
Wagner et al. 1990:667 (H); Herbarium Pacificum Staff 1998:9 (O); Imada et
al. 2000:12 (O); Staples et al. 2003:11 (K)
K
O Mo L
M Ka
H
Wagner et al. 1990:669 (Ni, K, O, Mo, L, M, Ka, H)
nat
K
O Mo
M Ka
H
Wagner et al. 1990:669 (K, O, M, Ka, H); Hughes 1995:5 (Mo)
Desmodium triflorum (L.) DC.
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:670 (K, O, Mo, L, M, Ka, H)
Dioclea wilsonii Standl.
nat?
K
H
Wagner et al. 1990:670 (K, H)
Entada phaseoloides (L.) Merr.
ind?
K
Enterolobium cyclocarpum (Jacq.) Griseb.
nat
Erythrina crista-galli L.
nat
O
Erythrina poeppigiana (Walp.) O.F.Cook
nat
O
Erythrina sandwicensis O.Deg.
end
Ni
Ni
K
Wagner et al. 1990:671 (K)
H
Wagner et al. 1990:630 [cultivated, possibly adventive]; Staples & Herbst
2005:298 (KEY), 314 (DESCR); Parker & Parsons 2012b:68 (NEWNAT/H,
DESCR)
H
Staples & Herbst 2005:300, 315 (KEY), 315 (DESCR); Lau & Frohlich
2012:15 (NEWNAT/O, DESCR); Parker & Parsons 2012b:68 (NEWNAT/H,
DESCR)
Daehler & Baker 2006:7 (NEWNAT/O, DESCR)
O Mo L
M Ka
H
Wagner et al. 1990:672 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:300, 315 (KEY), 316 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 84
Scientific Name
Dicot
Fabaceae
Status
Kl Le Ni K
Falcataria moluccana (Miq.) Barneby & J.W.Grimes
nat
Flemingia macrophylla (Willd.) Merr.
nat
Flemingia strobilifera (L.) Aiton & W.T.Aiton
nat
Gliricidia sepium (Jacq.) Walp.
nat
Glycine microphylla Tindale
nat
O
Haematoxylum campechianum L.
nat
O
Indigofera spicata Forssk.
nat
Indigofera suffruticosa Mill.
nat
Indigofera tinctoria L.
K
O Mo L
nat
Inga sertulifera DC. ssp. leptopus (Benth.) T.D.Penn.
nat
Kanaloa kahoolawensis Lorence & K.R.Wood
end
M
Wagner et al. 1990:690 (K, O, Mo, H [as Paraserianthes falcataria]);
Herbarium Pacificum Staff 1998:10 (NOM:Syn. P. falcataria); Staples &
Herbst 2005:299 (KEY), 331 (DESCR); Oppenheimer & Bartlett 2002:8
(WM); Starr et al. 2003:27 (EM); Oppenheimer 2007:23 (L)
H
Parker & Parsons 2012b:69 (NEWNAT/H, DESCR)
Lorence et al. 1995:37 (NEWNAT/EM, DESCR)
H
Ni
Citation
H
M
Staples & Herbst 2005:300 (KEY), 331 (DESCR); Parker & Parsons
2012b:69 (NEWNAT/H)
Lau & Frohlich 2012:15 (NEWNAT/O, DESCR)
H
Wagner et al. 1990:674 (O, H); Staples & Herbst 2005:298 (KEY), 332
(DESCR)
K
O Mo L
M
H
Wagner et al. 1990:675 (K, O, M [as Indigofera spicata]); Du Puy et al.
1993:727 (KEY), 731/Herbst & Wagner 1999:21 (NOM: I. spicata
misapplied = I. hendecaphylla); Starr et al. 2002:20 (H); Oppenheimer
2003:13 (Mo); Herbst et al. 2004:7 (L); Wagner et al. 2012:39 (NOM: I.
hendecaphylla misapplied)
K
O Mo L
M Ka
H
Wagner et al. 1990:676 (Ni, K, O, L, M, Ka, H); Hughes 1995:6 (Mo)
?
Inga feuillei DC.
O Mo L
M Ka H
O?
Wagner et al. 1990:675 (O? [reported in 1800s from Honolulu, not vouchered
since], DESCR)
H
K
Staples & Herbst 2005:298 (KEY), 332 (DESCR); Parker & Parsons
2012b:69 (NEWNAT/H, DESCR)
Lorence & Flynn 2006:2 (NEWNAT/K, DESCR)
Ka
Lorence & Wood 1994:137 (SPNOV/Ka, DESCR)/Herbst & Wagner 1999:21
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 85
Scientific Name
Dicot
Fabaceae
Status
Kummerowia striata (Thunb.) Schindl.
nat
Lablab purpureus (L.) Sweet
nat?
Lathyrus latifolius L.
Lathyrus tingitanus L.
Lespedeza cuneata (Dum.Cours.) G.Don
Kl Le Ni K
O Mo L
M Ka H
Citation
O
M
H?
Wagner et al. 1990:679 (H? [as Lespedeza striata; single coll. in 1932 as
pasture legume on Parker Ranch land]); Staples et al. 2002:10 (NOM:Syn. L.
striata); Oppenheimer 2003:13 (NEWNAT/WM, DESCR); Lau & Frohlich
2012:16 (O)
O Mo
M
H
Wagner et al. 1990:676 (K, O, Mo, M, H); Staples & Herbst 2005:299
(KEY), 318 (DESCR)
nat
M
H
Wagner et al. 1990:678 (EM); Starr et al. 2002:20 (EM); Staples & Herbst
2005:299 (KEY), 332 (DESCR); Pratt & Bio 2012:76 (H)
nat
M
K
?
Wagner et al. 1990:678 (EM)
O?
Wagner et al. 1990:679 (O? [apparently sparingly naturalized, but represented
by a single coll. in 1940 from a Honolulu experimental station garden],
DESCR) [Note: the single spm. (Hosaka 2528) redetermined as Lespedeza
juncea (L.f.) Pers. var. sericea (Thunb.) Lace & Hemsley by D. Herbst & G.
Staples, Oct 1995 (BISH)]
Leucaena Xspontanea C.E.Hughes & S.A.Harris
nat
M
Oppenheimer 2004:12 (NEWNAT/EM, DESCR)
Leucaena diversifolia (Schltdl.) Benth.
nat
M
Starr et al. 2008:45 (NEWNAT/EM, DESCR)
Leucaena leucocephala (Lam.) de Wit
nat
Lotus subbiflorus Lag.
nat
Lotus uliginosus Schkuhr
nat
Lupinus angustifolius L.
Lupinus arboreus Sims
Kl
Ni
K
M Ka
H
Wagner et al. 1990:679 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:298 (KEY), 333 (DESCR)
M
H
Wagner et al. 1990:681 (M, H)
M
H
Wagner et al. 1990:681 (Mo, M, H); Lorence et al. 1995:37 (K)
nat
H
Staples & Herbst 2005:299 (KEY), 333 (DESCR); Pratt & Bio 2012:76
(NEWNAT/H)
nat
H
Staples et al. 2002:10 (NEWNAT/H, DESCR)
K
O Mo L
Mo
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 86
Scientific Name
Dicot
Fabaceae
Status
Kl Le Ni K
Lupinus hybridus Lem.
nat
Macroptilium atropurpureum (DC.) Urb.
nat
Macroptilium lathyroides (L.) Urb.
nat
Macrotyloma axillare (E.Mey.) Verdc. var. glabrum
(E.Mey.) Verdc.
nat
Medicago lupulina L.
nat
K
Medicago minima (L.) Bartal.
nat
K
Medicago polymorpha L.
nat
Medicago rugosa Desr.
Ni
O Mo L
M Ka H
Citation
H
Wagner et al. 1990:682 (H)
K
O Mo L
M Ka
H
Wagner et al. 1990:682 (O, M, H); Oppenheimer 2003:13 (Mo); Lorence &
Flynn 2006:3 (K); Oppenheimer 2008:28 (L); Starr & Starr 2011:27 (Ka)
K
O Mo L
M Ka
H
Wagner et al. 1990:683 (Ni, K, O, L, M, Ka, H); Shannon & Wagner
1996:13/Herbarium Pacificum Staff 1996:4 (Mo)
O
Staples et al. 2003:11 (NEWNAT/O, DESCR)
M
H
Wagner et al. 1990:684 (O, M, H); Lorence et al. 1995:37 (K)
L
M
H
Wagner et al. 1990:684 (L, M, H); Wagner & Herbst 1995:20 (K)
K
O Mo L
M
H
Wagner et al. 1990:684 (K, O, L, M, H); Shannon & Wagner 1996:14 (Mo)
nat
K
O Mo L
M
H
Wagner et al. 1990:684 (O, H); Lorence & Flynn 1997:10 (K); Oppenheimer
& Bartlett 2002:8 (WM); Oppenheimer 2007:24 (Mo, L)
Medicago sativa L.
nat
K
O Mo L
M
H
Wagner et al. 1990:686 (K, O, L, H); Wagner & Herbst 1995:20 (EM);
Wagner et al. 1997:57 (EM); Oppenheimer et al. 1999:8 (WM); Oppenheimer
2003:13 (Mo)
Melilotus alba Medik.
nat
Mo
M
H
Wagner et al. 1990:686 (H); Wagner & Herbst 1995:20 (EM); Shannon &
Wagner 1996:14 (Mo); Oppenheimer 2004:12 (WM)
Melilotus indica (L.) All.
nat
K
O
M
H
Wagner et al. 1990:687 (K, O, M, H)
Mezoneuron kavaiense (H.Mann) Hillebr.
end
K
O
M
H
Wagner et al. 1990:647 (K, O, WM, H [as Caesalpinia kavaiensis]); Bruneau
et al. 2008:711/Wagner et al. 2012:37 (NOM:Syn. C. kavaiensis)
Mimosa pudica L. var. unijuga (Duchass. & Walp.)
Griseb.
nat
K
O Mo L
M
H
Wagner et al. 1990:687 (K, O, L, M, H); Oppenheimer 2003:13 (Mo); Imada
2007:37 (Ni)
Ni
O
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 87
Scientific Name
Dicot
Fabaceae
Status
Kl Le Ni K
K
O Mo L
O Mo
M Ka H
M
H
Citation
Mucuna gigantea (Willd.) DC. ssp. gigantea
ind
Mucuna sloanei Fawc. & Rendle var. persericea WilmotDear
end
Mucuna sloanei Fawc. & Rendle var. sloanei
ind
K
O
M
H
Wagner et al. 1990:689 (O, M, H [as Mucuna urens]); Wilmot-Dear 1990:25
(K, O, EM, WM, H, NOM: M. urens misapplied, KEY, DESCR)/Wagner &
Herbst 1995:20
Neonotonia wightii (Wight & Arn.) Lackey
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:674 (O, M, Ka, H [as Glycine wightii]); Hughes 1995:6
(Mo); Herbarium Pacificum Staff 1996:4 (K, NOM:Syn. G. wightii);
Oppenheimer 2007:24 (L)
Pachyrhizus erosus (L.) Urb.
nat
H
Wagner et al. 1990:689 (H); Staples & Herbst 2005:299 (KEY), 319
(DESCR)
Paraserianthes lophantha (Willd.) I.Nielsen ssp.
montana (Jungh.) I.Nielsen
nat
K? O? Mo?
M
H?
Wagner et al. 1990:690 (K?, O?, Mo?, M?, H? [forestry planting, especially
on Maui, where it is reproducing but not spreading]); Oppenheimer & Bartlett
2002:8 (NEWNAT/EM)
Parkinsonia aculeata L.
nat
K
M
Peltophorum pterocarpum (DC.) K.Heyne
nat
K
Piscidia piscipula (L.) Sargent
nat
Pithecellobium dulce (Roxb.) Benth.
nat
M
O
Wagner et al. 1990:689 (K, O, Mo, M, H [as Mucuna gigantea]); WilmotDear 1990:27 (VARNOV/EM, KEY, DESCR)/Wagner & Herbst 1995:20
Wagner et al. 1990:691 (K, O, M); Staples & Herbst 2005:298 (KEY), 334
(DESCR)
Staples & Herbst 2005:298 (KEY), 319 (DESCR); Frohlich & Lau 2012:38
(NEWNAT/K, DESCR)
O
Ni
K
Wagner et al. 1990:688 (K, O, Mo, M, H [as Mucuna gigantea]); WilmotDear 1990:5 (KEY)/Wagner & Herbst 1995:21 (NOM: Haw'n material
recognized as subsp. gigantea)
Staples & Herbst 2005:300 (KEY), 334 (DESCR); Lau & Frohlich 2012:16
(NEWNAT/O, DESCR)
O Mo L
M
H
Wagner et al. 1990:691 (Ni, K, O, Mo, M, H); Oppenheimer & Bartlett
2002:8 (L); Staples & Herbst 2005:298 (KEY), 321 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 88
Scientific Name
Dicot
Fabaceae
Platymiscium stipulare Benth.
Status
Kl Le Ni K
?
O Mo L
M Ka H
O?
Citation
Staples & Herbst 2005:300 (KEY), 322 (DESCR); Frohlich & Lau 2008:6
(NEWNAT/O, DESCR); Wagner et al. 2012:41 (O record adventive)
Prosopis juliflora (Sw.) DC.
nat
Prosopis pallida (Humb. & Bonpl. ex Willd.) Kunth
nat
Pueraria montana (Lour.) Merr. var. chinensis (Ohwi)
Maesen & S.M.Almeida ex Sanjappa & Pradeep
nat
K
Pueraria montana (Lour.) Merr. var. lobata (Willd.)
Maesen & S.M.Almeida ex Sanjappa & Pradeep
nat
K
Pueraria phaseoloides (Roxb.) Benth.
nat
Rhynchosia minima (L.) DC.
nat
K
Samanea saman (Jacq.) Merr.
nat
K
O Mo
M
H
Wagner et al. 1990:696 (O, H); Lorence et al. 1995:37 (K); Oppenheimer &
Bartlett 2002:8 (WM); Staples & Herbst 2005:298 (KEY), 324 (DESCR);
Oppenheimer 2007:24 (Mo)
Senna alata (L.) Roxb.
nat
K
O Mo L
M
H
Wagner et al. 1990:698 (K, O); Oppenheimer & Bartlett 2000:5 (WM); Starr
et al. 2003:28 (EM); Staples & Herbst 2005:298, 325 (KEY), 325 (DESCR);
Wysong et al. 2007:4 (Mo); Oppenheimer 2007:24 (L); Parker & Parsons
2012b:69 (H)
Senna didymobotrya (Fresen.) H.S.Irwin & Barneby
nat
K
Senna gaudichaudii (Hook. & Arn.) H.S.Irwin &
Barneby
ind
K
Le
K
O
L
K
O Mo L
Wagner et al. 1990:692 (O); Imada et al. 2000:12 (K); Staples & Herbst
2005:298 (KEY), 322 (DESCR); Oppenheimer 2007:24 (L)
M Ka
H
Wagner et al. 1990:692 (K, O, L, M, Ka, H); Hughes 1995:6 (Mo); Staples &
Herbst 2005:298 (KEY), 322 (DESCR); Wood & LeGrande 2006:23 (Le)
Wagner et al. 1990:695 (K [as Pueraria lobata var. thomsoni]); Staples et al.
2002:11 (NOM:Syn. P. lobata var. thomsoni)
O
M
H
O
Wagner et al. 1990:695 (O, M, H [as Pueraria lobata var. lobata]); Herbst &
Wagner 1999:21 (COMBNOV:Syn. P. lobata var. lobata); Frohlich & Lau
2012:39 (K)
Wagner et al. 1990:693 (cultivated, with tendency to naturalize); Frohlich &
Lau 2008:7 (NEWNAT/O, DESCR)
Wagner et al. 1990:695 (K)
M
O Mo L
M Ka
Wagner et al. 1990:698 (K, EM); Staples & Herbst 2005:298, 325 (KEY),
326 (DESCR)
H
Wagner et al. 1990:699 (K, O, Mo, L, M, H); Wagner et al. 1997:57 (Ka)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 89
Scientific Name
Dicot
Fabaceae
Status
Senna multijuga (Rich.) H.S.Irwin & Barneby var.
multijuga
nat
Senna obtusifolia (L.) H.S.Irwin & Barneby
nat
Senna occidentalis (L.) Link
nat
Senna pendula (Humb. & Bonpl. ex Willd.) H.S.Irwin
& Barneby var. advena (Vogel) H.S.Irwin & Barneby
Kl Le Ni K
O Mo L
M Ka H
K
Herbarium Pacificum Staff 1998:10 (NEWNAT/K); Staples & Herbst
2005:298, 325 (KEY), 326 (DESCR [as Senna multijuga])
O
Ni
Citation
L
H
Wagner et al. 1990:700 (H); Staples et al. 2003:12 (O); Oppenheimer 2011:7
(L)
K
O Mo
M
H
Wagner et al. 1990:700 (Ni, K, O, EM, H); Oppenheimer et al. 1999:8 (WM);
Oppenheimer 2003:14 (Mo)
nat
K
O Mo L
M
H
Wagner et al. 1990:701 (K, O, Mo, L, M, H)
Senna septemtrionalis (Viv.) H.S.Irwin & Barneby
nat
K
O Mo L
M
H
Wagner et al. 1990:701 (K, O, Mo, M); Wagner & Herbst 1995:21 (H);
Herbarium Pacificum Staff 1996:5 (L)
Senna siamea (Lam.) H.S.Irwin & Barneby
nat
O
Staples & Herbst 2005:298, 325 (KEY), 326 (DESCR); Lau & Frohlich
2012:16 (NEWNAT/O, DESCR)
Senna spectabilis (DC.) H.S.Irwin & Barneby
nat
O
Staples & Herbst 2005:298, 325 (KEY), 326 (DESCR); Lau & Frohlich
2012:17 (NEWNAT/O, DESCR)
Senna surattensis (Burm.f.) H.S.Irwin & Barneby
nat
K
O Mo L
Sesbania sesban (L.) Merr.
nat
K
O
Sesbania tomentosa Hook. & Arn.
end
K
O Mo L
M Ka
H
Wagner et al. 1990:704 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:300 (KEY), 327 (DESCR)
Sophora chrysophylla (Salisb.) Seem.
end
K
O Mo L
M
H
Wagner et al. 1990:706 (K, O, Mo, L, M, H); Staples & Herbst 2005:300
(KEY), 334 (DESCR)
Sophora tomentosa L.
nat
H
Pratt & Bio 2012:77 (NEWNAT/H)
Ni
M
Wagner et al. 1990:702 (K, O, M); Staples et al. 2002:11 (Mo); Staples &
Herbst 2005:298, 325 (KEY), 325 (DESCR [as subsp. surattensis]);
Oppenheimer 2007:25 (L)
Wagner et al. 1990:704 (K, O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 90
Scientific Name
Dicot
Fabaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Spartium junceum L.
nat
K
Strongylodon ruber Vogel
end
K
O Mo
Stylosanthes guianensis (Aubl.) Sw. var. guianensis
nat
K
Mo?
Herbst et al. 2004:7, 8 (NEWNAT/K, Mo? [based on cultivated colls. in
USDA Plant Materials Center field plot, Ho‘olehua], KEY)
Stylosanthes guianensis (Aubl.) Sw. var. intermedia
(Vogel) Hassler
?
Mo?
Herbst et al. 2004:7, 8 (NEWNAT/Mo? [based on cultivated coll. in USDA
Plant Materials Center field plot, Ho‘olehua], KEY)
Stylosanthes scabra Vogel
nat
Stylosanthes viscosa Sw.
K
Wagner et al. 1990:707 (K)
M
H
Wagner et al. 1990:707 (K, O, Mo, M, H)
O Mo L
M
H
Wagner et al. 1990:708 (K, O, Mo, L [as Stylosanthes fruticosa]); Herbst et
al. 2004:7 (NOM: S. fruticosa misapplied, remove K [spms. reidentified as S.
guianensis var. guianensis], add EM, KEY); Kennedy et al. 2010:21 (H);
Frohlich & Lau 2012:39 (K)
nat
O
M
H
Herbst et al. 2004:7 (NEWNAT/O, WM, KEY); Pratt & Bio 2012:77 (H)
Tamarindus indica L.
nat
O Mo
Tephrosia candida Roxb. ex DC.
nat
Tephrosia purpurea (L.) Pers. var. purpurea
pol
Trifolium arvense L. var. arvense
Staples & Herbst 2005:298 (KEY), 328 (DESCR); Wysong et al. 2007:4
(NEWNAT/Mo); Lau & Frohlich 2012:17 (O)
M
Ni
K
O Mo L
Oppenheimer & Bartlett 2000:5 (NEWNAT/WM, DESCR)
M Ka
H
Wagner et al. 1990:710 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:300 (KEY), 334 (DESCR [as Tephrosia purpurea])
nat
M
H
Wagner et al. 1990:711 (M, H)
Trifolium campestre Schreb.
nat
M
Trifolium dubium Sibth.
nat
Trifolium glomeratum L.
nat
Trifolium hybridum L. var. hybridum
nat
K
M
Starr & Starr 2011:27 (NEWNAT/EM, DESCR)
H
M
Wagner et al. 1990:712 (EM, H); Lorence et al. 1995:37 (K)
Wagner et al. 1990:712 (EM)
H
Wagner et al. 1990:712 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 91
Scientific Name
Dicot
Fabaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Trifolium incarnatum L. var. incarnatum
nat
M
Trifolium pratense L. var. sativum Schreb.
nat
M
H
Wagner et al. 1990:714 (M, H)
Trifolium repens L. var. repens
nat
M
H
Wagner et al. 1990:714 (K, M, H); Oppenheimer 2008:28 (L)
Trifolium resupinatum L.
nat
H
Wagner et al. 1990:715 (H)
Trifolium subterraneum L.
nat
H
Wagner et al. 1990:715 (H)
Ulex europaeus L.
nat
M
H
Wagner et al. 1990:715 (M, H); Conant 1996:1 (Mo)
Vachellia farnesiana (L.) Wight & Arn.
nat
M Ka
H
Wagner et al. 1990:641 (K, O, Mo, M, Ka, H [as Acacia farnesiana]);
Oppenheimer & Bartlett 2002:7 (L); Mabberley 2008:890/Wagner et al.
2012:36 (NOM:Syn. A. farnesiana)
Vicia menziesii Spreng.
end
H
Wagner et al. 1990:716 (H)
Vicia sativa L. ssp. nigra (L.) Ehrh.
nat
M
H
Wagner et al. 1990:717 (EM, H); Wood 2007:15 (K); Starr & Starr 2011:32
(ELEV)
Vicia tenuifolia Roth
nat
M
H
Wagner et al. 1990:717 (M, H)
Vicia villosa Roth
nat
H
Wagner et al. 1990:719 (H)
Vigna adenantha (G.Mey.) Maréchal, Mascherpa &
Stainier
ind
O
H
Wagner et al. 1990:720 (O, H)
Vigna hosei (Craib) Backer
nat
O
H
Staples et al. 2006:7 (NEWNAT/O, DESCR); Pratt & Bio 2012:77 (H)
Vigna luteola (Jacq.) Benth.
nat
K
O
Vigna marina (Burm.) Merr.
ind
K
O Mo
K
L
Mo
K
O Mo L
K
Wagner et al. 1990:712 (EM)
Staples et al. 2003:12 (NEWNAT/O, KEY); Frohlich & Lau 2012:39 (K)
M
H
Wagner et al. 1990:720 (K, O, Mo, M, H); Staples et al. 2003:12 (KEY);
Staples & Herbst 2005:299, 300 (KEY), 330 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 92
Scientific Name
Dicot
Fabaceae
Status
Kl Le Ni K
Vigna o-wahuensis Vogel
end
Ni
Vigna speciosa (Kunth) Verdc.
nat
K
Vigna vexillata (L.) A.Rich.
nat
K
Zornia cf.reticulata Sm.
nat
Zornia gemella (Willd.) Vogel
nat
O Mo L
M Ka H
O Mo L
M Ka
O
M
H
Citation
Wagner et al. 1990:720 (Ni, O, Mo, L, M, Ka, H)
Herbst 1998:3 (NEWNAT/O, DESCR); Staples & Herbst 2005:299, 300
(KEY), 330 (DESCR); Kennedy et al. 2010:22 (K, EM)
Wagner et al. 2012:43 (NEWNAT/K)
M
Starr et al. 2006:35 (NEWNAT/EM, DESCR)
O Mo
Staples et al. 2006:7 (NEWNAT/O); Oppenheimer 2008:28 (Mo)
Fagaceae
Quercus suber L.
nat
H
Herbst & Wagner 1996:11 (NEWNAT/H); Staples & Herbst 2005:335
(DESCR)
Flacourtiaceae
Caloncoba echinata (Oliver) Gilg
?
O?
Dovyalis hebecarpa (Gardner) Warb.
nat
Flacourtia indica (N.L.Burm.) Merr.
nat
Xylosma crenatum (H.St.John) H.St.John
end
K
Xylosma hawaiiense Seem.
end
K
Daehler & Baker 2006:15 (O? [adventive], DESCR)
H
O
Herbst 1998:3 (NEWNAT/H, DESCR); Staples & Herbst 2005:336 (KEY,
DESCR)
Staples & Herbst 2005:336 (KEY), 337 (DESCR); Lau & Frohlich 2012:17
(NEWNAT/O, DESCR)
Wagner et al. 1990:722 (K)
O Mo L
M
H
Wagner et al. 1990:722 (K, O, Mo, L, M, H); Govaerts & Radcliffe-Smith
1996:175 (NOMNOV: Drypetes sherffii, replacement name for D. forbesii
Sherff, a name previously published); Herbst & Wagner 1999:21 (NOM:Syn.
D. sherffii)
Frankeniaceae
Frankenia salina (Molina) I.M.Johnst.
nat
Only in Northwestern Hawaiian Islands; see NWHI checklist
Fumariaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 93
Scientific Name
Dicot
Fumariaceae
Fumaria officinalis L.
Status
Kl Le Ni K
O Mo L
nat
M Ka H
M
Citation
Starr & Starr 2011:27 (NEWNAT/EM, DESCR)
Gentianaceae
Centaurium erythraea Raf. ssp. erythraea
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:725 (K, O, Mo, L, M, Ka, H)
Schenkia sebaeoides Griseb.
end
K
O Mo L
M
Erodium cicutarium (L.) L'Hér. ex Aiton
nat
K
O Mo L
M
Erodium sp.
nat
Geranium arboreum A.Gray
end
Geranium cuneatum Hook. ssp. cuneatum
end
H
Wagner et al. 1990:730 (H)
Geranium cuneatum Hook. ssp. hololeucum (A.Gray)
Carlquist & Bissing
end
H
Wagner et al. 1990:730 (H)
Geranium cuneatum Hook. ssp. hypoleucum (A.Gray)
Carlquist & Bissing
end
H
Wagner et al. 1990:730 (H)
Geranium cuneatum Hook. ssp. tridens (Hillebr.)
Carlquist & Bissing
end
Geranium dissectum L.
nat
Geranium hanaense A.C.Medeiros & H.St.John
end
M
Wagner et al. 1990:730 (EM)
Geranium hillebrandii Aedo & Muñoz Garm.
end
M
Wagner et al. 1990:732 (EM [as Geranium humile]); Aedo & Garmendia
1997:725/Herbst & Wagner 1999:21 (NOMNOV: the combination G. humile
was previously published, thus a new epithet was chosen)
Wagner et al. 1990:725 (K, O, Mo, WM [as Centaurium sebaeoides]);
Wagner & Herbst 1995:21 (L); Mansion 2004:725 (NOM:Syn. C. sebaeoides)
Geraniaceae
H
Mo
Hughes 1995:6 (NEWNAT/Mo)
M
Wagner et al. 1990:729 (EM)
M
K
Wagner et al. 1990:727 (K, O, Mo, L, M, H)
M
Wagner et al. 1990:730 (EM)
H
Wagner et al. 1990:730 (K, EM, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 94
Scientific Name
Dicot
Geraniaceae
Status
Kl Le Ni K
Geranium homeanum Turcz.
nat
K
Geranium kauaiense (Rock) H.St.John
end
K
Geranium molle L.
nat
Geranium multiflorum A.Gray
end
Geranium pusillum L.
nat
Geranium retrorsum L'Hér. ex DC.
nat
Pelargonium capitatum (L.) L'Hér. ex Aiton
nat
Pelargonium spp.
?
O Mo L
M Ka H
M
H
Citation
Wagner et al. 1990:732 (K, M, H)
Wagner et al. 1990:733 (K)
H
M
L
M
M
K?
Wagner et al. 1990:733 (H)
Wagner et al. 1990:733 (EM)
H
Wagner et al. 1990:734 (H)
H
Wagner et al. 1990:734 (L, M, H)
Starr et al. 2008:46 (NEWNAT/EM, DESCR)
Wagner et al. 1990:727 (K? [cultivated and persisting or escaping in Köke‘e
State Park]); Staples & Herbst 2005:340 (KEY)
Gesneriaceae
Cyrtandra biserrata Hillebr.
end
Mo
Cyrtandra calpidicarpa (Rock) H.St.John & Storey
end
Cyrtandra confertiflora (Wawra) C.B.Clarke
end
Cyrtandra cordifolia Gaudich.
end
O
Wagner et al. 1990:751 (O)
Cyrtandra crenata H.St.John & Storey
end
O
Wagner et al. 1990:752 (O)
Cyrtandra cyaneoides Rock
end
O
M
Wagner et al. 1990:749 (Mo); St. John 1987d:479 (SPNOV/EM [as
Cyrtandra higashinoi], DESCR), 483 (SPNOV/EM [as C. ustulata], DESCR);
St. John 1988b:200 (NOMNOV: C. hemisphaerica, to replace nom. illeg. C.
ustulata); Herbst & Wagner 1999:22 (EM, NOM:Syns. C. hemisphaerica, C.
higashinoi, C. ustulata)
Wagner et al. 1990:749 (O)
K
Wagner et al. 1990:751 (K [as Cyrtandra confertiflora var. confertiflora]);
Wagner & Herbst 2003:29 (C. confertiflora var. obovata = C. heinrichii)
K
Wagner et al. 1990:752 (K); Lorence et al. 1995:37 (REDISCOVER/K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 95
Scientific Name
Dicot
Gesneriaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Cyrtandra dentata H.St.John & Storey
end
Wagner et al. 1990:753 (O)
Cyrtandra ferripilosa H.St.John
end
Cyrtandra filipes Hillebr.
end
Cyrtandra garnotiana Gaudich.
end
Cyrtandra giffardii Rock
end
Cyrtandra gracilis Hillebr. ex C.B.Clarke
end
O
Wagner et al. 1990:755 (O); Wagner et al. 2012:44 (REDISCOVER/O)
Cyrtandra grandiflora Gaudich.
end
O
Wagner et al. 1990:755 (O)
Cyrtandra grayana Hillebr.
end
Mo L
M
Wagner et al. 1990:756 (Mo, L, WM)
Cyrtandra grayi C.B.Clarke
end
Mo
M
Wagner et al. 1990:758 (Mo, EM, WM)
Cyrtandra halawensis Rock
end
Mo
Cyrtandra hashimotoi Rock
end
Cyrtandra hawaiensis C.B.Clarke
end
Cyrtandra heinrichii H.St.John
end
Cyrtandra hematos H.St.John
end
Cyrtandra kalihii Wawra
end
Cyrtandra kamoolaensis H.St.John
end
Mo
M
St. John 1987f:497 (SPNOV/EM, DESCR); Wagner & Herbst 2003:29
(acceptance of species)
M
Wagner et al. 1990:753 (Mo, WM)
O
Wagner et al. 1990:754 (O)
H
Wagner et al. 1990:758 (Mo)
M
O Mo
K
Wagner et al. 1990:754 (H)
M
Wagner et al. 1990:759 (M)
H
Wagner et al. 1990:759 (O, Mo, M, H)
Wagner et al. 1990:751 (K [as Cyrtandra confertiflora var. obovata]); St. John
1987f:498 (NOMNOV/K [as C. heinrichii], DESCR); Wagner & Herbst
2003:29 (acceptance of C. heinrichii, NOM:Syn. C. confertiflora var. obovata)
Mo
O
Wagner et al. 1990:760 (Mo)
Wagner et al. 1990:762 (O)
K
St. John 1987f:499 (SPNOV/K, DESCR); Wagner et al. 1990:739 (accepted
as a valid species, but not given full treatment due to publication deadline)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 96
Scientific Name
Dicot
Gesneriaceae
Status
Kl Le Ni K
O Mo L
M Ka H
K
Citation
Cyrtandra kauaiensis Wawra
end
Wagner et al. 1990:762 (K)
Cyrtandra kaulantha H.St.John & Storey
end
Cyrtandra kealiae Wawra ssp. kealiae
end
K
Wagner et al. 1990:763 (K [as Cyrtandra kealiae]), 766 (K [as C.
limahuliensis]); Wagner & Lorence 2000:17 (NOM:Syn. C. limahuliensis,
creation of subsp. kealiae with addition of subsp. urceolata, DESCR)
Cyrtandra kealiae Wawra ssp. urceolata W.L.Wagner &
Lorence
end
K
Wagner & Lorence 2000:19 (SSPNOV/K, DESCR)
Cyrtandra kohalae H.St.John
end
Cyrtandra laxiflora H.Mann
end
O
Wagner et al. 1990:764 (O)
Cyrtandra lessoniana Gaudich.
end
O
Wagner et al. 1990:765 (O)
Cyrtandra longifolia (Wawra) Hillebr. ex C.B.Clarke
end
Cyrtandra lydgatei Hillebr.
end
Cyrtandra lysiosepala (A.Gray) C.B.Clarke
end
Cyrtandra macraei A.Gray
end
Cyrtandra macrocalyx Hillebr.
end
Cyrtandra menziesii Hook. & Arn.
end
Cyrtandra munroi C.N.Forbes
end
Cyrtandra nanawaleensis H.St.John
end
O
Wagner et al. 1990:763 (O)
H
K
Wagner et al. 1990:764 (H)
Wagner et al. 1990:766 (K)
Mo L
M
Wagner et al. 1990:767 (Mo, L. WM)
H
O
Wagner et al. 1990:767 (H)
Wagner et al. 1990:767 (O)
Mo
M
Wagner et al. 1990:769 (Mo); Oppenheimer 2010:36 (EM, WM)
H
L
M
Wagner et al. 1990:769 (H)
Wagner et al. 1990:770 (L); St. John 1987e:487 (SPNOV/WM [as Cyrtandra
acmule], DESCR), 489 (SPNOV/WM [as C. imparis], DESCR); Herbst &
Wagner 1999:21 (WM, NOM:Syns. C. acmule, C. imparis)
H
St. John 1987f:500 (SPNOV/H, DESCR); Wagner & Herbst 2003:29
(acceptance of species)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 97
Scientific Name
Dicot
Gesneriaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Cyrtandra oenobarba H.Mann
end
K
Wagner et al. 1990:770 (K)
Cyrtandra olona C.N.Forbes
end
K
Wagner et al. 1990:771 (K)
Cyrtandra oxybapha W.L.Wagner & D.R.Herbst
end
Cyrtandra paliku W.L.Wagner, K.R.Wood & Lorence
end
K
Wagner et al. 2001:146 (SPNOV/K, DESCR)
Cyrtandra paludosa Gaudich. var. microcarpa Wawra
end
K
Wagner et al. 1990:773 (K)
Cyrtandra paludosa Gaudich. var. paludosa
end
K
Cyrtandra pickeringii A.Gray
end
K
Cyrtandra platyphylla A.Gray
end
Cyrtandra polyantha C.B.Clarke
end
Cyrtandra procera Hillebr.
end
Cyrtandra propinqua C.N.Forbes
end
O
Wagner et al. 1990:775 (O)
Cyrtandra pruinosa H.St.John & Storey
end
O
Wagner et al. 1990:777 (O)
Cyrtandra rivularis H.St.John & Storey
end
O
Wagner et al. 1990:777 (O)
Cyrtandra sandwicensis (H.Lév.) H.St.John & Storey
end
O
Wagner et al. 1990:777 (O)
Cyrtandra sessilis H.St.John & Storey
end
O
Wagner et al. 1990:778 (O)
Cyrtandra spathulata H.St.John
end
Cyrtandra subumbellata (Hillebr.) H.St.John & Storey
end
Cyrtandra tintinnabula Rock
end
M
O
M
Wagner et al. 1990:771 (WM)
H
Wagner et al. 1990:773 (K, O, M, H)
Wagner et al. 1990:773 (K)
M
H
O
Wagner et al. 1990:773 (M, H)
Wagner et al. 1990:774 (O)
Mo
Wagner et al. 1990:775 (Mo)
M
Wagner et al. 1990:778 (M)
O
Wagner et al. 1990:779 (O)
H
Wagner et al. 1990:779 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 98
Scientific Name
Dicot
Gesneriaceae
Status
Cyrtandra viridiflora H.St.John & Storey
end
Cyrtandra wagneri Lorence & Perlman
end
Cyrtandra waianaeensis H.St.John & Storey
end
Cyrtandra wainihaensis H.Lév.
end
Cyrtandra waiolani Wawra
end
Cyrtandra wawrae C.B.Clarke
end
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Wagner et al. 1990:780 (O)
H
O
Lorence & Perlman 2007:357 (SPNOV/H, DESCR)
Wagner et al. 1990:780 (O)
K
Wagner et al. 1990:762 (syn. of Cyrtandra kauaiensis); Wagner & Herbst
2003:29 (RESURRECT/K)
O
Wagner et al. 1990:781 (O)
K
Wagner et al. 1990:781 (K)
Goodeniaceae
Scaevola chamissoniana Gaudich.
end
Mo L
M
H
Wagner et al. 1990:783 (Mo, L. M, H)
Scaevola coriacea Nutt.
end
K
O Mo L
M
H
Wagner et al. 1990:784 (Ni, K, O, Mo, L. M, H)
Scaevola gaudichaudiana Cham.
end
K
O
Scaevola gaudichaudii Hook. & Arn.
end
K
O Mo L
Scaevola glabra Hook. & Arn.
end
K
O
Scaevola hobdyi W.L.Wagner
end
Scaevola kilaueae O.Deg.
end
Scaevola mollis Hook. & Arn.
end
K
O Mo
Scaevola procera Hillebr.
end
K
Mo
Ni
Wagner et al. 1990:784 (K, O)
M
H
Wagner et al. 1990:784 (K, O, Mo, L. M, H)
Wagner et al. 1990:786 (K, O)
M
Wagner 1996:225 (SPNOV/WM, DESCR)
H
Wagner et al. 1990:786 (H)
Wagner et al. 1990:786 (K, O, Mo)
Wagner et al. 1990:788 (K, Mo)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 99
Scientific Name
Dicot
Goodeniaceae
Scaevola taccada (Gaertn.) Roxb.
Status
Kl Le Ni K
ind
Ni
K
O Mo L
M Ka H
O Mo L
M Ka
O
M
H
Citation
Wagner et al. 1990:788 (Ni, K, O, Mo, L. M, Ka, H [as Scaevola sericea]);
Green 1991:121/Shannon et al. 1997:801 (NOM:Syn. S. sericea); Staples &
Herbst 2005:348 (DESCR)
Grossulariaceae
Brexia madagascariensis (Lam.) Thouars ex Ker Gawl.
nat
Escallonia rubra (Ruiz & Pav.) Pers. var. macrantha
(Hook. & Arn.) Reiche
nat
Starr et al. 2003:28 ( (NEWNAT/EM, DESCR); Staples & Herbst 2005:350
(DESCR); Frohlich & Lau 2010:11 (O); Starr & Starr 2011:28 (WM)
H
Staples & Herbst 2005:350 (DESCR); Parker & Parsons 2012a:60
(NEWNAT/H, DESCR)
Gunneraceae
Gunnera kauaiensis Rock
end
K
Wagner et al. 1990:790 (K)
Gunnera petaloïdea Gaudich.
end
K
Gonocarpus chinensis (Lour.) Orchard ssp. verrucosus
(Maiden & Betche) Orchard
nat
K
Myriophyllum aquaticum (Vell.) Verdc.
nat
K
O
Broussaisia arguta Gaudich.
end
K
O Mo L
Philadelphus karwinskyanus Koehne
nat
K
O Mo
M
H
Wagner et al. 1990:790 (O, Mo, M, H); Wood 2006:17 (K)
H
Wagner et al. 1990:791 (H); Lorence et al. 1995:38 (K)
H
Wagner et al. 1990:792 (K, O, H); Staples & Herbst 2005:351 (DESCR)
H
Wagner et al. 1990:794 (K, O, Mo, L, M, H)
Haloragaceae
Hydrangeaceae
M
M
Lorence et al. 1995:38 (NEWNAT/K, EM, DESCR); Starr et al. 2002:20
(EM); Staples & Herbst 2005:354 (DESCR)
Hydrophyllaceae
Nama sandwicensis A.Gray
end
Le Ni
K
O Mo L
M
H
Wagner et al. 1990:795 (Ni, K, O, Mo, L, M, H)
Lamiaceae
Haplostachys bryanii Sherff
end
Mo
Wagner et al. 1990:799 (Mo)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 100
Scientific Name
Dicot
Lamiaceae
Status
Haplostachys haplostachya (A.Gray) H.St.John
end
Haplostachys linearifolia (Drake) Sherff
end
Haplostachys munroi C.N.Forbes
end
Haplostachys truncata (A.Gray) Hillebr.
end
Hyptis capitata Jacq.
nat
Hyptis pectinata (L.) Poit.
nat
Hyptis suaveolens (L.) Poit.
nat
Hyptis verticillata Jacq.
nat
Lamium amplexicaule L.
nat
Leonotis leonurus (L.) R.Br.
Kl Le Ni K
O Mo L
M Ka H
K
M
Mo
H
M
Wagner et al. 1990:799 (L)
M
Wagner et al. 1990:801 (M)
O
Wagner & Herbst 1995:21 (NEWNAT/O, DESCR)
O Mo L
M
H
Wagner et al. 1990:802 (K, O, M, H); Wysong et al. 2007:5 (Mo);
Oppenheimer 2007:25 (L)
O Mo
M
H
Wagner et al. 1990:802 (H); Wagner & Herbst 1995:22 (O); Starr et al.
2006:36 (EM); Oppenheimer 2008:29 (Mo)
Mo
?
Wagner et al. 1990:799 (K, M, H)
Wagner et al. 1990:799 (Mo, M)
L
K
Citation
M
Starr & Starr 2011:28 (NEWNAT/EM, DESCR)
M
Wagner et al. 1990:802 (EM); Wysong et al. 2007:5 (Mo)
O?
Wagner et al. 1990:803 (O? [single coll. in 1929 from Nu‘uanu Valley])
Leonotis nepetifolia (L.) R.Br.
nat
K
O Mo L
M
H
Wagner et al. 1990:803 (K, O, M, H); Oppenheimer 2003:14 (Mo); Staples &
Herbst 2005:356 (KEY), 365 (DESCR); Oppenheimer 2010:36 (L)
Leonurus japonicus Houtt.
nat
K
O Mo
M
H
Wagner et al. 1990:803 (K, O, M, H [as Leonurus sibiricus]); Harley & Paton
2001:244/Staples et al. 2003:14 (NOM: L. sibiricus misapplied); Wysong et
al. 2007:5 (Mo)
Lepechinia hastata (A.Gray) Epling
ind?
Marrubium vulgare L.
nat
Mentha Xspicata L.
nat
M
L
O
Wagner et al. 1990:804 (EM)
H
Wagner et al. 1990:806 (L, H)
H
Wagner et al. 1990:807 (O, H [as Mentha spicata]); Staples & Herbst
2005:356, 357 (KEY), 358 (DESCR [as M. xspicata])
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 101
Scientific Name
Dicot
Lamiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Mentha pulegium L.
nat
M
Mentha suaveolens Ehrh.
nat
Ocimum basilicum L.
nat
Ni
K
O Mo L
Ocimum gratissimum L.
nat
Ni
K
Phyllostegia ambigua (A.Gray) Hillebr.
end
Phyllostegia bracteata Sherff
end
M
Phyllostegia brevidens A.Gray
end
M
Phyllostegia electra C.N.Forbes
end
Phyllostegia floribunda Benth.
end
Phyllostegia glabra (Gaudich.) Benth. var. glabra
end
O Mo L
Phyllostegia glabra (Gaudich.) Benth. var. lanaiensis
Sherff
end
L
Phyllostegia grandiflora (Gaudich.) Benth.
end
O
Citation
Wagner et al. 1990:807 (M); Staples & Herbst 2005:356, 357 (KEY), 357
(DESCR)
H
Wagner et al. 1990:807 (O, H [as Mentha xvillosa]); Staples et al. 2003:14
(NOM: M. xvillosa, M. xrotundifolia misapplied); Staples & Herbst
2005:356, 357 (KEY), 358 (DESCR)
M
H
Wagner et al. 1990:808 (Ni, O, Mo, M, H); Staples & Herbst 2005:356, 359
(KEY), 359 (DESCR); Oppenheimer 2010:36 (L); Frohlich & Lau 2012:40
(K)
O Mo
M
H
Wagner et al. 1990:808 (Ni, O, Mo, M, H); Lorence et al. 1995:39 (K);
Staples & Herbst 2005:356, 359 (KEY), 360 (DESCR)
Mo
M
H
Wagner et al. 1990:813 (M, H); Wagner 1999b:266 (EM, WM); Wood
2006:17 (Mo)
Wagner et al. 1990:814 (M); Wagner 1999b:267 (EM, WM); Oppenheimer
2004:13 (REDISCOVER/WM)
H
K
Wagner et al. 1990:814 (H); Wagner 1999b:267 (EM)
Wagner et al. 1990:815 (K); Wagner 1999b:267
H
O
M
Wagner et al. 1990:815 (H); Wagner 1999b:268
Wagner et al. 1990:816 (O, Mo, L, M); Wagner 1999b:268; Oppenheimer et
al. 1999:8 (WM)
Wagner et al. 1990:816 (L); Wagner 1999b:269
Wagner et al. 1990:816 (O); Wagner 1999b:269
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 102
Scientific Name
Dicot
Lamiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Mo L
M
Citation
Phyllostegia haliakalae Wawra
end
Wagner et al. 1990:819 (L, EM [as Phyllostegia imminuta]), 821 [as syn. of
P. mollis]; Wagner 1999b:269 (RESURRECT/Mo, L, EM, NOM:Syn. P.
imminuta); Oppenheimer 2011:7 (REDISCOVER/EM)
Phyllostegia helleri Sherff
end
Phyllostegia hillebrandii H.Mann ex Hillebr.
end
Phyllostegia hirsuta Benth.
end
Phyllostegia hispida Hillebr.
end
Phyllostegia kaalaensis H.St.John
end
Phyllostegia kahiliensis H.St.John
end
K
St. John 1987a:176 (SPNOV/K, DESCR); Wagner 1999b:270
Phyllostegia knudsenii Hillebr.
end
K
Wagner et al. 1990:819 (K); Lorence et al. 1995:39 (REDISCOVER/K);
Wagner 1999b:271; Wood 2012:95 (EXTINCT?)
Phyllostegia lantanoides Sherff
end
Phyllostegia macrophylla (Gaudich.) Benth.
end
Phyllostegia mannii Sherff
end
Phyllostegia micrantha H.St.John
end
O
St. John 1987a:178 (SPNOV/O, DESCR); Wagner 1999b:272
Phyllostegia mollis Benth.
end
O
Wagner et al. 1990:821 (O, Mo, EM); Wagner 1999b:272 (removal of Mo,
EM after resurrection of Phyllostegia haliakalae, DESCR)
Phyllostegia parviflora (Gaudich.) Benth. var.
glabriuscula A.Gray
end
K
Wagner et al. 1990:816 (K); Wagner 1999b:269; Wood 2006:18
(REDISCOVER/K)
M
Wagner et al. 1990:817 (EM); Wagner 1999b:269
O
Wagner et al. 1990:817 (O); Wagner 1999b:270
Mo
Wagner et al. 1990:817 (Mo); Wagner 1999b:270
O
St. John 1987a:176 (SPNOV/O, DESCR); Russell & Bruegmann
1996b:53097; Wagner 1999b:270
O
Wagner et al. 1990:820 (O); Wagner 1999b:271
M
Mo
H
M
Wagner et al. 1990:820 (EM, H); Wagner 1999b:271 (WM)
Wagner et al. 1990:820 (Mo, EM); Wagner 1999b:272
H
Wagner et al. 1990:822 (H); Wagner 1999b:273
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 103
Scientific Name
Dicot
Lamiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Phyllostegia parviflora (Gaudich.) Benth. var. lydgatei
(Sherff) W.L.Wagner
end
O
Wagner et al. 1990:822 (as Phyllostegia mollis var. lydgatei, syn. of P.
parviflora var. parviflora); Wagner 1999a:282 (RESURRECT/O,
COMBNOV:Syn. P. mollis var. lydgatei, DESCR); Wagner 1999b:274
Phyllostegia parviflora (Gaudich.) Benth. var. parviflora
end
O
Phyllostegia pilosa H.St.John
end
Phyllostegia racemosa Benth.
end
Phyllostegia renovans W.L.Wagner
end
Phyllostegia rockii Sherff
end
Phyllostegia stachyoides A.Gray
end
Phyllostegia variabilis Bitter
end
Phyllostegia velutina (Sherff) H.St.John
end
H
Wagner et al. 1990:825 (H); Wagner 1999b:276
Phyllostegia vestita Benth.
end
H
Wagner et al. 1990:825 (H); Wagner 1999b:276
Phyllostegia waimeae Wawra
end
Phyllostegia warshaueri H.St.John
end
Phyllostegia wawrana Sherff
end
Plectranthus cylindraceus Hochst. ex Benth.
nat
Plectranthus neochilus Schltr.
nat
Mo
M
Wagner et al. 1990:822 (O, WM); Wagner 1999b:273
M
St. John 1987a:179 (SPNOV/EM, DESCR); Wagner 1999b:274 (Mo, EM,
WM, DESCR)
H
K
Wagner et al. 1990:822 (H); Wagner 1999b:275
Wagner 1999a:280 (SPNOV/K, DESCR); Wagner 1999b:275 (DESCR)
M
Mo
M
Wagner et al. 1990:823 (EM); Wagner 1999b:275
H
Wagner et al. 1990:823 (Mo, WM, H); Wagner 1999b:275
Only in Northwestern Hawaiian Islands; see NWHI checklist
K
Wagner et al. 1990:825 (K); Wagner 1999b:277; Wood 2007:16
(REDISCOVER/K)
H
K
Wagner et al. 1990:826 (H); Wagner 1999b:277
Wagner et al. 1990:826 (K); Lorence et al. 1995:39 (REDISCOVER/K);
Wagner 1999b:277
H
O
Staples & Herbst 2005:356, 360 (KEY), 361 (DESCR); Parker & Parsons
2010:42 (NEWNAT/H, DESCR)
Lau & Frohlich 2012:18 (NEWNAT/O, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 104
Scientific Name
Dicot
Lamiaceae
Status
Kl Le Ni K
Ni
K
O Mo L
Plectranthus parviflorus Willd.
ind
Plectranthus prostratus Gürke
nat
Plectranthus scutellarioides (L.) R.Br.
nat
Plectranthus verticillatus (L.f.) Druce
nat
Prunella vulgaris L.
nat
Salvia coccinea Etl.
nat
Salvia occidentalis Sw.
nat
Ni
Stachys arvensis L.
nat
Stenogyne angustifolia A.Gray
end
Mo
Stenogyne bifida Hillebr.
end
Mo
Stenogyne calaminthoides A.Gray
end
Stenogyne calycosa Sherff
end
Stenogyne campanulata Weller & A.K.Sakai
end
Stenogyne cinerea Hillebr.
end
M Ka H
O Mo L
M
H
O
K
Citation
Wagner et al. 1990:827 (Ni, K, O, Mo, L, M, H)
Staples et al. 2002:11 (NEWNAT/O, DESCR); Staples & Herbst 2005:356,
360 (KEY), 361 (DESCR)
O
M
O
M
H
Wagner et al. 1990:828 (K, O, M, H [as Plectranthus scutellarioides]);
Wagner & Herbst 1999:1883 (NOM:Syn. P. scutellarioides = Solenostemon
scutellarioides); Staples & Herbst 2005:356 (KEY), 364 (DESCR); Grayer et
al. 2010/Wagner et al. 2012:48 (NOM:Syn. S. scutellarioides)
Starr et al. 2004:23 (NEWNAT/O, EM, DESCR); Staples & Herbst
2005:356, 360 (KEY), 361 (DESCR)
Mo
M
H
Wagner et al. 1990:828 (Mo, M, H)
K
O Mo L
M
H
Wagner et al. 1990:829 (O, Mo, L, M, H); Lorence et al. 1995:40 (K);
Staples & Herbst 2005:355, 362 (KEY), 363 (DESCR)
K
O Mo L
M
H
Wagner et al. 1990:829 (Ni, K, O, Mo, L, M, H)
Ni? K
O Mo L
M Ka? H
Wagner et al. 1990:831 (Ni?, K, O, Mo, L, M, Ka?, H) [Note: assumed
naturalized on Ni, Ka, but unvouchered]
M
Wagner et al. 1990:835 (Mo, M, H); Wood & Oppenheimer 2008:548 (KEY)
H
Wagner et al. 1990:835 (Mo)
H
M
K
Wagner et al. 1990:835 (H)
Wagner et al. 1990:837 (EM, WM); Wood & Oppenheimer 2008:548 (KEY)
Wagner et al. 1990:837 (K); Wood & Oppenheimer 2008:548 (KEY)
M
Wagner et al. 1990:837 (EM); Wood & Oppenheimer 2008:548 (KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 105
Scientific Name
Dicot
Lamiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
H
Citation
Stenogyne cranwelliae Sherff
end
Stenogyne haliakalae Wawra
end
Stenogyne kaalae Wawra ssp. kaalae
end
O
Wagner et al. 1990:838 (O); Wagner & Weller 1999:449 (now referred to as
subsp. kaalae following recognition of subsp. sherffii, KEY)
Stenogyne kaalae Wawra ssp. sherffii (O.Deg.)
W.L.Wagner & Weller
end
O
Wagner et al. 1990:838 (as Stenogyne sherffii, syn. of S. kaalae); Wagner &
Weller 1999:449 (RESURRECT/O, COMBNOV & STATNOV:Syn. S.
sherffii, KEY, DESCR)
Stenogyne kamehamehae Wawra
end
Stenogyne kanehoana O.Deg. & Sherff
end
Stenogyne kauaulaensis K.R.Wood & H.Oppenheimer
end
Stenogyne kealiae Wawra
end
Stenogyne macrantha Benth.
end
Stenogyne microphylla Benth.
end
Stenogyne oxygona O.Deg. & Sherff
end
Stenogyne purpurea H.Mann
end
Stenogyne rotundifolia A.Gray
end
M
Stenogyne rugosa Benth.
end
M
Stenogyne scrophularioides Benth.
end
M
Mo
Wagner et al. 1990:837 (H)
Wagner et al. 1990:837 (EM); Wood & Oppenheimer 2008:548 (KEY)
M
Wagner et al. 1990:838 (Mo, EM, WM); Wood & Oppenheimer 2008:548
(KEY)
O
Wagner et al. 1990:838 (O)
M
Wood & Oppenheimer 2008:544 (SPNOV/WM, KEY, DESCR)
K
Wagner et al. 1990:841 (syn. of Stenogyne purpurea); Wagner & Weller
1991:50 (RESURRECT/K, KEY, DESCR)
M
H
Wagner et al. 1990:839 (H)
H
Wagner et al. 1990:839 (EM, H); Wood & Oppenheimer 2008:548 (KEY)
H
Wagner et al. 1990:839 (H)
K
Wagner et al. 1990:841 (K); Wagner & Weller 1991:51 (Stenogyne kealiae
resurrected from synonymy, KEY, DESCR)
Wagner et al. 1990:841 (EM); Wood & Oppenheimer 2008:548 (KEY)
H
Wagner et al. 1990:841 (EM, H); Wood & Oppenheimer 2008:548 (KEY)
H
Wagner et al. 1990:842 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 106
Scientific Name
Dicot
Lamiaceae
Status
Stenogyne sessilis Benth.
end
Stenogyne viridis Hillebr.
end
Kl Le Ni K
O Mo L
M Ka H
L
M
H
M
Citation
Wagner et al. 1990:842 (L, M, H); Wood & Oppenheimer 2008:548 (KEY)
Wagner et al. 1990:843 (WM); Wood & Oppenheimer 2008:548 (KEY)
Lauraceae
Cassytha filiformis L.
ind
Cinnamomum burmanni (Nees) Blume
Ni
K
O Mo L
M
H
Wagner et al. 1990:844 (Ni, K, O, Mo, L, M, H)
nat
K
O
L
M
H
Wagner et al. 1990:846 (O [as "burmannii"]); Wagner & Herbst 1995:22
(EM); Wagner et al. 1997:57 (H); Meidell et al. 1997:18 (WM); Wagner et al.
2003: 32 (corrected spelling); Starr et al. 2004:23 (K); Staples & Herbst
2005:366 (DESCR); Oppenheimer 2007:25 (L)
Cinnamomum camphora (L.) J.Presl
nat
K
O
L
M
Cinnamomum verum J.Presl
nat
K
O
Cryptocarya mannii Hillebr.
end
K
O
Persea americana Mill.
nat
K
O
M
Wagner et al. 1990:846 (K, O, L, M); Staples & Herbst 2005:366 (DESCR)
H
Wagner et al. 1990:846 (K, O, M); Imada et al. 2000:12 (H); Staples &
Herbst 2005:366 (DESCR)
Wagner et al. 1990:847 (K, O)
L
M
H
Wagner et al. 1990:847 (K, O, M, H); Staples & Herbst 2005:368 (DESCR);
Oppenheimer 2010:36 (L)
Leeaceae
Leea indica (Burm.f.) Merr.
?
O?
Staples & Herbst 2005:371 (DESCR); Daehler & Baker 2006:15 (O?
[adventive])
Lentibulariaceae
Utricularia gibba L.
nat
H
Wagner et al. 1990:849 (H)
H
Wagner et al. 1990:850 (EM); Oppenheimer 2004:13 (K); Starr et al. 2006:36
(H)
Linaceae
Linum bienne Mill.
nat
Linum trigynum L.
nat
K
M
O
Wagner et al. 1990:850 (O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 107
Scientific Name
Dicot
Loganiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Labordia cyrtandrae (Baill.) H.St.John
end
Wagner et al. 1990:854 (O)
Labordia degeneri Sherff
end
Labordia fagraeoidea Gaudich.
end
Labordia hedyosmifolia Baill.
end
Labordia helleri Sherff
end
K
Labordia hirtella H.Mann
end
K
Labordia hosakana (Sherff) W.L.Wagner, D.R.Herbst &
Sohmer
end
O
Wagner et al. 1990:859 (O)
Labordia kaalae C.N.Forbes
end
O
Wagner et al. 1990:859 (O); Wood et al. 2007:198 (KEY)
Labordia lorenciana K.R.Wood, W.L.Wagner &
T.Motley
end
K
Wood et al. 2007:195 (SPNOV/K, KEY, DESCR)
Labordia lydgatei C.N.Forbes
end
K
Wagner et al. 1990:860 (K)
Labordia pumila (Hillebr.) Skottsb.
end
K
Wagner et al. 1990:860 (K)
Labordia sessilis A.Gray
end
Labordia tinifolia A.Gray var. lanaiensis Sherff
end
Labordia tinifolia A.Gray var. tinifolia
end
K
Labordia tinifolia A.Gray var. wahiawaensis H.St.John
end
K
K
Wagner et al. 1990:855 (K)
O
Wagner et al. 1990:855 (O)
Mo L
M
H
Wagner et al. 1990:856 (Mo, L, M, H)
Wagner et al. 1990:856 (K); Wood et al. 2007:197 (KEY)
O Mo L
M
H
O
Wagner et al. 1990:857 (K, O, Mo, L, M, H)
Wagner et al. 1990:860 (O)
L
O Mo
Wagner et al. 1990:861 (Mo, L [incl. syn. Labordia triflora]); Motley
1995:223, 224 (remove Mo, following resurrection of L. triflora from
synonymy, KEY)
M
H
Wagner et al. 1990:861 (K, O, Mo, M, H); Motley 1995:223, (KEY); Wood
et al. 2007:197 (KEY)
Wagner et al. 1990:862 (K); Motley 1995:223, 225 (KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 108
Scientific Name
Dicot
Loganiaceae
Status
Labordia triflora Hillebr.
end
Labordia venosa Sherff
end
Labordia waialealae Wawra
end
Labordia waiolani Wawra
end
Polypremum procumbens L.
Kl Le Ni K
O Mo L
M Ka H
Mo
Wagner et al. 1990:861 (syn. of Labordia tinifolia var. lanaiensis); Motley
1995:221 (RESURRECT/Mo, KEY, DESCR)/Herbst & Wagner 1999:23;
Wood et al. 2007:197 (KEY)
M
K
Citation
Wagner et al. 1990:862 (EM)
Mo?
Wagner et al. 1990:862 (K); Staples et al. 2003:14 (Mo? [previously
overlooked Faurie coll. from 1910, possibly mislabled])
O Mo L
Wagner et al. 1990:863 (O, Mo, L)
?
H?
Wagner et al. 1990:850 (H? [reportedly naturalized in Hawai‘i Volcanoes
National Park, but no vouchers seen], DESCR)
Lythraceae
Ammannia coccinea Rottb.
nat
K
O
M
Cuphea carthagenensis (Jacq.) J.F.Macbr.
nat
K
O Mo L
M
H
Wagner et al. 1990:866 (K, O, Mo, M, H); Oppenheimer & Bartlett 2002:9
(L)
Cuphea hyssopifolia Kunth
nat
O
M
H
Wagner et al. 1990:866 (H); Staples & Herbst 2005:373 (KEY), 374
(DESCR); Imada 2007:37 (EM); Oppenheimer 2011:8 (O)
Cuphea ignea A.DC.
nat
H
Wagner et al. 1990:866 (H); Staples & Herbst 2005:373 (KEY), 374
(DESCR)
Cuphea subuligera Koehne
nat
H
Staples & Herbst 2005:373 (KEY), 375 (DESCR); Parker & Parsons
2012a:60 (NEWNAT/H, DESCR)
Lythrum maritimum Kunth
nat
H
Wagner et al. 1990:868 (K, O, Mo, L, M, H); Herbst & Wagner 1999:23
(status change from "ind?" to "nat")
K
O Mo L
M
Wagner et al. 1990:865 (O, M [as Ammannia auriculata], K [as A. coccinea]);
Herbst & Wagner 1999:23 (NOM: A. auriculata misapplied, O/M records
transferred to A. coccinea)
Magnoliaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 109
Scientific Name
Dicot
Magnoliaceae
Liriodendron tulipifera L.
Michelia champaca L.
Status
Kl Le Ni K
O Mo L
M Ka H
nat
H
?
Citation
Staples & Herbst 2005:378 (DESCR); Parker & Parsons 2012b:70
(NEWNAT/H)
O?
Staples & Herbst 2005:377 (DESCR); Daehler & Baker 2006:15 (O?
[adventive])
O
Staples & Herbst 2005:379 (KEY), 380 (DESCR); Daehler & Baker 2006:8
(NEWNAT/O); Lorence & Flynn 2006:3 (K)
Malpighiaceae
Hiptage benghalensis (L.) Kurz
nat
K
Abelmoschus moschatus Medik.
nat
K
Abutilon auritum (Wall. ex Link) Sweet
nat
Abutilon eremitopetalum Caum
end
Abutilon grandifolium (Willd.) Sweet
nat
Le Ni
K
O Mo L
M Ka
Abutilon incanum (Link) Sweet
ind?
Ni
K
O Mo L
M Ka
Abutilon indicum (L.) Sweet
nat
O
Abutilon menziesii Seem.
end
O?
Abutilon sandwicense (O.Deg.) Christoph.
end
O
Bastardia viscosa (L.) Kunth
nat
Malvaceae
Wagner et al. 1990:870 (K); Staples & Herbst 2005:383 (KEY), 384
(DESCR)
M
Wagner et al. 1990:871 (EM)
L
Wagner et al. 1990:871 (L)
H
Wagner et al. 1990:872 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:383, 385 (KEY), 385 (DESCR); Wood & LeGrande 2006:24 (Le)
Wagner et al. 1990:872 (Ni, K, O, Mo, L, M, Ka)
Wagner et al. 1990:873 (O)
L
M
H
Wagner et al. 1990:873 (O? [single coll. in 1981 from abandoned ‘Ewa
canefield], L, M, H); Herbarium Pacificum Staff 1999:4 (O? [additional
seemingly wild O‘ahu colls.]); Staples & Herbst 2005:383, 385 (KEY), 385
(DESCR); Oppenheimer 2011:8 (WM)
Wagner et al. 1990:873 (O)
H
Staples et al. 2002:12 (NEWNAT/H, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 110
Scientific Name
Dicot
Malvaceae
Status
Kl Le Ni K
Ni
M Ka H
nat
Gossypium hirsutum L.
nat
Gossypium tomentosum Nutt. ex Seem.
end
Herissantia crispa (L.) Brizicky
nat
Hibiscadelphus bombycinus C.N.Forbes
end
Hibiscadelphus crucibracteatus Hobdy
end
Hibiscadelphus distans L.E.Bishop & D.R.Herbst
end
Hibiscadelphus giffardianus Rock
end
H
Wagner et al. 1990:879 (H); Lorence & Wagner 1995:186 (KEY); Staples &
Herbst 2005:383 (KEY), 393 (DESCR)
Hibiscadelphus hualalaiensis Rock
end
H
Wagner et al. 1990:879 (H); Lorence & Wagner 1995:186 (KEY); Staples &
Herbst 2005:383 (KEY), 393 (DESCR)
Hibiscadelphus wilderianus Rock
end
Hibiscadelphus woodii Lorence & W.L.Wagner
end
Hibiscus arnottianus A.Gray ssp. arnottianus
end
Hibiscus arnottianus A.Gray ssp. immaculatus
(M.J.Roe) D.M.Bates
end
K
O Mo L
H
Citation
Gossypium barbadense L.
Ni
K
O Mo L
Wagner et al. 1990:875 (Ni, K, O, Mo, L, H); Staples & Herbst 2005:383
(KEY), 386 (DESCR)
O
M
Wagner et al. 1990:876 (O); Starr et al. 2002:21 (EM)
O Mo L
M Ka
Wagner et al. 1990:876 (Ni, K, O, Mo, L, M, Ka); Staples & Herbst 2005:383
(KEY), 386 (DESCR)
M
Wagner et al. 1990:877 (WM)
H
L
Wagner et al. 1990:878 (H); Lorence & Wagner 1995:186 (KEY)
Wagner et al. 1990:878 (L); Lorence & Wagner 1995:186 (KEY)
K
Wagner et al. 1990:878 (K); Lorence & Wagner 1995:186 (KEY); Staples &
Herbst 2005:383 (KEY), 393 (DESCR)
M
K
Wagner et al. 1990:879 (EM); Lorence & Wagner 1995:186 (KEY)
Lorence & Wagner 1995:183 (SPNOV/K, KEY, DESCR)/Herbst & Wagner
1999:23; Wood 2012:95 (EXTINCT?)
O
Wagner et al. 1990:883 (O); Staples & Herbst 2005:383, 387 (KEY), 388
(DESCR)
Mo
Wagner et al. 1990:883 (Mo); Staples & Herbst 2005:383, 387 (KEY), 388
(DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 111
Scientific Name
Dicot
Malvaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Hibiscus arnottianus A.Gray ssp. punaluuensis
(Skottsb.) D.M.Bates
end
Hibiscus brackenridgei A.Gray ssp. brackenridgei
end
Hibiscus brackenridgei A.Gray ssp. mokuleianus
(M.J.Roe) D.M.Bates
end
Hibiscus brackenridgei A.Gray ssp. molokaianus (Rock
ex Caum) F.D.Wilson
end
Hibiscus clayi O.Deg. & I.Deg.
end
K
Hibiscus furcellatus Desr.
ind
K
O
M
H
Wagner et al. 1990:885 (K, O, M, H); Wilson 1993:279 (KEY, DESCR)
Hibiscus kokio Hillebr. ex Wawra ssp. kokio
end
K
O Mo
M
H?
Wagner et al. 1990:887 (K, O, Mo, M, H? [presumed to have been present on
Hawai‘i]); Staples & Herbst 2005:383, 387 (KEY), 388 (DESCR)
Hibiscus kokio Hillebr. ex Wawra ssp. saintjohnianus
(M.J.Roe) D.M.Bates
end
K
Hibiscus macrophyllus Roxb. ex Hornem.
nat
O
?
O?
Frohlich & Lau 2008:7 (NEWNAT/O, DESCR); Wagner et al. 2012:49 (O
record adventive)
nat
O
Wagner et al. 1990:887 (O); Staples & Herbst 2005:383, 387 (KEY), 389
(DESCR)
Hibiscus makinoi Jotani & Ohba
Hibiscus mutabilis L.
O
Citation
Wagner et al. 1990:883 (O); Staples & Herbst 2005:383, 387 (KEY), 388
(DESCR)
Mo L
K
M
H
O
Wagner et al. 1990:884 (Mo, L, M, H); Wilson 1993:277 (KEY, DESCR);
Staples & Herbst 2005:383, 387 (KEY), 388 (DESCR)
Wagner et al. 1990:884 (K, O); Wilson 1993:278 (KEY, DESCR); Staples &
Herbst 2005:383, 387 (KEY), 388 (DESCR)
Mo
Wagner et al. 1990:883 (as Hibiscus brackenridgei var. molokaiana, syn. of
H. brackenridgei subsp. brackenridgei); Wilson 1993:278
(RESURRECT/Mo, COMBNOV & STATNOV:Syn. H. brackenridgei var.
molokaiana, KEY, DESCR)/Herbst & Wagner 1999:23
Wagner et al. 1990:884 (K); Staples & Herbst 2005:383, 387 (KEY), 391
(DESCR)
Wagner et al. 1990:887 (K); Staples & Herbst 2005:383, 387 (KEY), 389
(DESCR)
H
Wagner et al. 1990:887 (O, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 112
Scientific Name
Dicot
Malvaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Hibiscus ovalifolius (Forssk.) Vahl
nat
K
Wagner et al. 1990:884 (K [as Hibiscus calyphyllus]); Staples et al. 2003:14
(NOM:Syn. H. calyphyllus); Staples & Herbst 2005:383, 387 (KEY), 391
(DESCR)
Hibiscus tiliaceus L.
ind?
K
Hibiscus waimeae A.Heller ssp. hannerae (O.Deg. &
I.Deg.) D.M.Bates
end
K
Wagner et al. 1990:889 (K); Staples & Herbst 2005:383, 387 (KEY), 391
(DESCR)
Hibiscus waimeae A.Heller ssp. waimeae
end
K
Wagner et al. 1990:889 (K); Staples & Herbst 2005:383, 387 (KEY), 391
(DESCR)
Kokia cookei O.Deg.
end
Kokia drynarioides (Seem.) Lewton
end
Kokia kauaiensis (Rock) O.Deg. & Duvel
end
Kokia lanceolata Lewton
end
Malachra alceifolia Jacq.
nat
Malva neglecta Wallr.
nat
Malva parviflora L.
nat
Malvastrum americanum (L.) Torr.
nat
O Mo L
M
H
Mo
Wagner et al. 1990:890 (Mo); Staples & Herbst 2005:383 (KEY), 393
(DESCR)
H
K
Ni
K
Wagner et al. 1990:890 (H); Staples & Herbst 2005:383 (KEY), 393
(DESCR)
Wagner et al. 1990:890 (K); Staples & Herbst 2005:383 (KEY), 393
(DESCR)
O
K
Wagner et al. 1990:888 (K, O, Mo, L, M, H); Fryxell 2001:258 (COMBNOV
[Talipariti tiliaceum]; Pfeil et al. 2002/Wagner & Herbst 2003:33 (Talipariti
is nested within Hibiscus and best maintained there, NOM:Syn. T. tiliaceum);
Staples & Herbst 2005:383, 387 (KEY), 390 (DESCR)
Wagner et al. 1990:891 (O)
O
O Mo L
O Mo
M
Wagner et al. 1990:891 (K, O, M)
M
Herbarium Pacificum Staff 1998:10 (NEWNAT/EM, DESCR); Starr & Starr
2011:32 (ELEV)
M Ka
H
Wagner et al. 1990:893 (Ni, K, O, Mo, L, M, Ka, H)
Wagner et al. 1990:894 (O); Starr et al. 2008:47 (Mo)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 113
Scientific Name
Dicot
Malvaceae
Status
Kl Le Ni K
Malvastrum coromandelianum (L.) Garcke ssp.
coromandelianum
nat
Ni
Malvaviscus penduliflorus DC.
nat
Modiola caroliniana (L.) G.Don
nat
Sida acuta Burm.f. ssp. carpinifolia (L.f.) Borss.Waalk.
nat
K
Sida ciliaris L.
nat
Sida cordifolia L.
nat
Sida fallax Walp.
ind
Sida linifolia Cav.
nat
Sida rhombifolia L.
nat?
Sida spinosa L.
nat
Sida urens L.
nat
Sidastrum micranthum (A.St.-Hil.) Fryxell
nat
Sidastrum paniculatum (L.) Fryxell
nat
Kl Le Ni
O Mo L
M Ka H
K
O Mo L
M Ka
H
Wagner et al. 1990:894 (Ni, K, O, Mo, L, M, Ka, H)
K
O Mo
M
H
Wagner et al. 1990:895 (K, M, H); Staples & Herbst 2005:383, 387 (KEY),
391 (DESCR); Oppenheimer 2007:26 (Mo); Oppenheimer 2011:8 (O)
M
H
Wagner et al. 1990:896 (M, H)
O
M
H
Wagner et al. 1990:897 (K, O, M, H)
K
O Mo L
M Ka
K
O Mo L
M
H
Wagner et al. 1990:897 (EM, H); Lorence et al. 1995:41 (K); Herbst et al.
2004:9 (O); Oppenheimer 2007:26 (Mo, L, WM)
K
O Mo L
M Ka
H
Wagner et al. 1990:897 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:383 (KEY), 392 (DESCR); Wood & LeGrande 2006:24 (Le)
Wagner et al. 1997:59 (NEWNAT/O, DESCR); Oppenheimer & Bartlett
2000:6 (WM); Staples et al. 2003:14 (K); Starr et al. 2004:24 (EM); Starr et
al. 2006:36 (Ka); Oppenheimer 2007:26 (L); Oppenheimer 2008:30 (Mo)
Mo
Ni
Citation
Wagner et al. 1990:898 (Mo)
K
O Mo L
M Ka
H
Wagner et al. 1990:898 (Ni, K, O, Mo, L, M, Ka, H)
K
O Mo L
M
H
Wagner et al. 1990:899 (O, H); Lorence & Flynn 1997:10 (K); Oppenheimer
et al. 1999:8 (WM); Staples et al. 2002:12 (L); Oppenheimer 2007:26 (Mo)
M
H
Wagner et al. 1990:899 (H); Starr et al. 2002:21 (EM); Oppenheimer 2007:27
(WM)
M
H
Wagner et al. 1990:901 (O, H); Starr et al. 2003:28 (EM); Lorence & Flynn
2006:3 (K); Oppenheimer 2007:27 (Mo); Oppenheimer 2008:30 (L)
H
Wagner et al. 1990:901 (H); Oppenheimer 2011:8 (L)
K
O Mo L
L
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 114
Scientific Name
Dicot
Malvaceae
Status
Thespesia populnea (L.) Sol. ex Corrêa
ind?
Urena lobata L.
nat
Kl Le Ni K
Kl
Ni
O Mo L
M Ka H
K
O Mo L
M
K
O
M
H
Citation
Wagner et al. 1990:902 (Ni, K, O, Mo, M, H); Staples & Herbst 2005:383
(KEY), 393 (DESCR); Oppenheimer 2007:27 (L)
Wagner et al. 1990:903 (K, O, EM); Staples & Herbst 2005:383 (KEY), 393
(DESCR)
Martyniaceae
Proboscidea louisianica (Mill.) Thell.
?
H?
Kennedy et al. 2010:24 (H? [adventive])
H
Wagner et al. 1990:905 (O, H [as Arthrostema ciliatum] ); Imada et al.
2000:13 (genus spelling corrected); Staples & Herbst 2005:394 (KEY), 397
(DESCR)
Melastomataceae
Arthrostemma ciliatum Pav. ex D.Don
nat
O
Clidemia hirta (L.) D.Don var. hirta
nat
K
O Mo L
M
H
Wagner et al. 1990:906 (K, O, Mo, M, H); Oppenheimer 2007:27 (L)
Dissotis rotundifolia (Sm.) Triana
nat
K
O
M
H
Wagner et al. 1990:907 (O, M, H); Herbarium Pacificum Staff 1996:5 (K);
Staples & Herbst 2005:394 (KEY), 395 (DESCR)
Heterocentron subtriplinervium (Link & Otto) A.Braun
& C.D.Bouché
nat
K
O
M
H
Wagner et al. 1990:909 (K, O, M, H); Staples & Herbst 2005:394 (KEY),
395 (DESCR)
Medinilla cumingii Naudin
nat
O
M
H
Oppenheimer 2004:13 (NEWNAT/EM, DESCR); Staples & Herbst 2005:394
(KEY), 396 (DESCR); Daehler & Baker 2006:15 (O? [adventive], DESCR);
Frohlich & Lau 2010:11 (O); Parker & Parsons 2010:42 (H)
Medinilla heterophylla A.Gray
nat
O
Medinilla magnifica Lindl.
nat
O
Medinilla venosa (Blume) Blume
nat
Frohlich & Lau 2010:12 (NEWNAT/O, DESCR)
H
M
Wagner et al. 1990:909 (widely planted, not known to escape); Staples &
Herbst 2005:394 (KEY), 396 (DESCR); Daehler & Baker 2006:15 (O?
[adventive], DESCR); Frohlich & Lau 2007:10 (NEWNAT/O); Parker &
Parsons 2010:42 (H)
Wagner et al. 1990:909 (EM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 115
Scientific Name
Dicot
Melastomataceae
Status
Kl Le Ni K
O Mo L
Melastoma sanguineum Sims
nat
Melastoma septemnervium Lour.
nat
K
O
Miconia calvescens DC.
nat
K
O
Oxyspora paniculata (D.Don) DC.
nat
Pterolepis glomerata (Rottb.) Miq.
nat
Tetrazygia bicolor (Mill.) Cogn.
nat
Tibouchina granulosa (Desr.) Cogn.
nat
O
Tibouchina herbacea (DC.) Cogn.
nat
O? Mo L
Tibouchina longifolia (Vahl) Baill. ex Cogn.
nat
Tibouchina urvilleana (DC.) Cogn. var. urvilleana
nat
K
Trembleya phlogiformis DC. var. parvifolia Cogn.
nat
K
M Ka H
M
H
Wagner et al. 1990:911 (H); Staples & Herbst 2005:394 (KEY), 397
(DESCR)
H
Wagner et al. 1990:910 (K, H [as Melastoma candidum]); Conant 1996:1 (O);
Conant 1998:1 (O); Staples et al. 2003:15 (NOM:Syn. M. candidum); Staples
& Herbst 2005:394 (KEY), 396 (DESCR)
H
Conant 1996:1 (NEWNAT/K, O); Medeiros et al. 1997 (EM, H, DESCR);
Conant 1998:1 (O); Staples & Herbst 2005:394 (KEY), 397 (DESCR)
O
K
Citation
Wagner et al. 1990:912 (O); Nagata 1995:12 (O); Staples & Herbst 2005:394
(KEY), 397 (DESCR)
O Mo L
H
Wagner et al. 1990:912 (K, O, H); Hughes 1995:7 (Mo); Wood 2006:18 (L)
H
Wagner et al. 1990:913 (H); Staples & Herbst 2005:394 (KEY), 397
(DESCR)
Wagner et al. 1990:915 (cultivated); Staples & Herbst 2005:394 (KEY), 397
(DESCR); Lau & Frohlich 2012:18 (NEWNAT/O, DESCR)
O
M
M
H
Wagner et al. 1990:915 (EM, WM, H); Herbarium Pacificum Staff 1999:5
(L); Wysong et al. 2007:5 (Mo); Frohlich & Lau 2007:10 (O); Wagner et al.
2012:52 (O plants removed, not definitely naturalized)
H
Wagner et al. 1990:916 (H)
H
Wagner et al. 1990:916 (K, O, M, H); Staples & Herbst 2005:394 (KEY),
397 (DESCR)
Wagner et al. 1990:917 (K)
Meliaceae
Azadirachta indica A.Juss.
nat
O
Staples & Herbst 2005:398 (KEY), 400 (DESCR); Lau & Frohlich 2012:19
(NEWNAT/O, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 116
Scientific Name
Dicot
Meliaceae
Status
Cedrela odorata L.
nat
Khaya nyasica Stapf ex Baker f.
nat
Melia azedarach L.
nat
Sandoricum koetjape (Burm.f.) Merr.
nat
Swietenia macrophylla King
nat
Toona ciliata M.Roem.
nat
Kl Le Ni K
O Mo L
M Ka H
M
Staples & Herbst 2005:398 (KEY), 400 (DESCR); Starr et al. 2006:37
(NEWNAT/EM, DESCR)
O
Ni
K
Citation
Daehler & Baker 2006:9 (NEWNAT/O, DESCR)
O Mo L
M
H
Wagner et al. 1990:918 (Ni, K, O, L, M, H); Hughes 1995:7 (Mo); Staples &
Herbst 2005:398 (KEY), 399 (DESCR)
M
Starr et al. 2004:24 (NEWNAT/EM, WM, DESCR); Staples & Herbst
2005:398 (KEY), 401 (DESCR)
O
M
Meidell et al. 1998:6 (NEWNAT/O, WM); Staples & Herbst 2005:398
(KEY), 400 (DESCR)
K
O Mo L
M
H
Wagner et al. 1990:920 (K, O, L, M, H [as Toona ciliata var. australis]);
Herbst & Wagner 1999:24 (NOM:Syn. T. ciliata var. australis); Staples &
Herbst 2005:398 (KEY), 401 (DESCR); Oppenheimer 2007:27 (Mo)
K
O Mo L
M
H
Wagner et al. 1990:921 (Ni, K. O, Mo, L, M, H [as Cocculus trilobus]);
Wagner & Herbst 1995:22 (NOM:Syn. C. trilobus)
Menispermaceae
Cocculus orbiculatus (L.) DC.
Stephania japonica (Thunb.) Miers
ind
Ni
?
O?
nat
O
?
O?
Frohlich & Lau 2007:12 (O? [adventive], DESCR)
Molluginaceae
Mollugo cerviana (L.) Ser.
L
H
Wagner et al. 1990:922 (H); Herbst et al. 2004:9 (O); Imada et al. 2008:13 (L)
Moraceae
Antiaris toxicaria Lesch.
Artocarpus altilis (Parkinson ex Z) Fosberg
Daehler & Baker 2006:16 (O? [adventive])
pol
Non-naturalized Polynesian introduction; Staples & Herbst 2005:403 (KEY,
DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 117
Scientific Name
Dicot
Moraceae
Status
Kl Le Ni K
O Mo L
Artocarpus heterophyllus Lam.
nat
Broussonetia papyrifera (L.) Vent.
pol
Fatoua villosa (Thunb.) Nakai
nat
Ficus macrophylla Desf. ex Pers.
nat
Ficus microcarpa L.f.
nat
Ficus nota (Blanco) Merr.
nat
O
Ficus platypoda (A.Cunn. ex Miq.) A.Cunn. ex Miq.
nat
O
Ficus pumila L.
?
O?
Ficus religiosa L.
?
O?
Ficus rubiginosa Desf. ex Vent.
nat
M Ka H
O
Ni
K
O Mo L
M Ka
H
Staples & Herbst 2005:403 (KEY), 404 (DESCR); Lau & Frohlich 2012:19
(NEWNAT/O, DESCR); Parker & Parsons 2012b:70 (NEWNAT/H)
H
Wagner et al. 1990:924 (Ni, K, O, Mo, L, M. Ka, H); Staples & Herbst
2005:403 (KEY), 404 (DESCR)
O
Wagner & Herbst 1995:22 (NEWNAT/O, DESCR)
Mo
K
Citation
O Mo L
M
H
Wagner et al. 1990:924; Oppenheimer & Bartlett 2000:6 (NEWNAT/WM,
DESCR); Starr et al. 2002:21 (H); Staples & Herbst 2005:403, 407 (KEY),
409 (DESCR); Oppenheimer 2006:12 (Mo)
M Ka
H
Wagner et al. 1990:924 (O, M, H); Lorence et al. 1995:41 (K); Hughes
1995:7 (Mo); Warren & Herbst 1994:2 (Ka); Staples & Herbst 2005:403, 408
(KEY), 409 (DESCR); Oppenheimer 2007:28 (L)
H
Herbarium Pacificum Staff 1998:10 (NEWNAT/O, H, DESCR); Staples &
Herbst 2005:403, 407 (KEY), 411 (DESCR)
H
Oppenheimer & Bartlett 2000:7 (NEWNAT/O, EM, WM, NOM: Ficus
rubiginosa misapplied, in part); Oppenheimer 2003:14 (H); Staples & Herbst
2005:403, 407, 408 (KEY), 410 (DESCR)
M
Staples & Herbst 2005:403, 407 (KEY), 410 (DESCR); Frohlich & Lau
2012:50 (O? [adventive])
H
L
M
Staples & Herbst 2005:403, 408 (KEY), 411 (DESCR); Frohlich & Lau
2008:7 (NEWNAT/O, DESCR); Parker & Parsons 2012b:70 (H); Wagner et
al. 2012:52 (O record adventive)
Wagner et al. 1990:924; Nagata 1995:12 (NEWNAT/O); Oppenheimer &
Bartlett 2000:7 (remove O, spm. reidentified as Ficus cf. platypoda);
Oppenheimer 2003:15 (NEWNAT/WM); Staples & Herbst 2005:403, 407,
408 (KEY), 410 (DESCR); Oppenheimer 2008:30 (L)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 118
Scientific Name
Dicot
Moraceae
Maclura tinctoria (L.) D.Don ex Steud.
Status
Kl Le Ni K
?
O Mo L
M Ka H
O?
Citation
Frohlich & Lau 2012:51 (O? [adventive])
Morus alba L.
nat
K
O Mo
M
H
Wagner et al. 1990:926 (K, O, Mo, M, H); Staples & Herbst 2005:403
(KEY), 412 (DESCR)
Streblus pendulinus (Endl.) F.Muell.
ind
K
O Mo L
M
H
Wagner et al. 1990:927 (K, O, Mo, L, M, H)
Muntingiaceae
Muntingia calabura L.
?
O?
Wagner et al. 1990:588 (O? [based on possibly naturalized coll. in 1946 from
Hälawa Valley], DESCR); Staples & Herbst 2005:336 (KEY), 337 (DESCR)
Myoporaceae
Myoporum sandwicense A.Gray
ind
Myoporum stellatum (G.L.Webster) O.Deg. & I.Deg.
end
Ni
K
O Mo L
M
H
O
Wagner et al. 1990:928 (Ni, K, O, Mo, L, M, H); Staples & Herbst 2005:414
(DESCR); Chinnock 2007:142 (acceptance of Myoporum sandwicense [as
subsp. sandwicense], 144 (proposed RESURRECT/L of syn. M. sandwicense
var. lanaiense [as COMBNOV & STATNOV M. sandwicense subsp.
lanaiense], 145 (proposed RESURRECT/EM of syn. M. degeneri]
Wagner et al. 1990:928 (syn. of Myoporum sandwicense); Chinnock
2007:146/Wagner et al. 2005– (RESURRECT/O)
Myricaceae
Morella cerifera (L.) Small
nat
K?
Morella faya (Aiton) Wilbur
nat
K
O
nat
K
O
L
M
H
Meidell et al. 1997:17 (NEWNAT/WM [as Myrica cerifera]); Wilbur
1994:100/Oppenheimer 2003:15 (NOM:Syn. M. cerifera); Frohlich & Lau
2012:40 (K? [adventive], H)
M
H
Wagner et al. 1990:931 (K, O, L, M, H [as Myrica faya]); Oppenheimer et al.
1999:9 (WM); Wilbur 1994:103/Staples et al. 2002:13 (NOM:Syn. M. faya)
M
H
Wagner et al. 1990:932 (O, H); Oppenheimer 2004:13 (WM); Staples &
Herbst 2005:415 (DESCR); Frohlich & Lau 2012:41 (K)
Myrsinaceae
Ardisia crenata Sims
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 119
Scientific Name
Dicot
Myrsinaceae
Ardisia elliptica Thunb.
Status
Kl Le Ni K
nat
K
O Mo L
M Ka H
O
M
H
Citation
Wagner et al. 1990:932 (K, O, EM, H); Meidell et al. 1998:7 (WM); Staples
& Herbst 2005:415 (DESCR)
Ardisia sieboldii Miq.
?
O?
Daehler & Baker 2006:16 (O? [adventive], DESCR)
Ardisia virens Kurz
?
O?
Daehler & Baker 2006:16 (O? [adventive], DESCR)
Embelia pacifica Hillebr.
end
K
O Mo L
M
H
Wagner et al. 1990:933 (K, O, Mo, L, M, H)
Myrsine alyxifolia Hosaka
end
K
Myrsine degeneri Hosaka
end
Myrsine denticulata (Wawra) Hosaka
end
K
Wagner et al. 1990:938 (K)
Myrsine fernseei (Mez) Hosaka
end
K
Wagner et al. 1990:938 (K)
Myrsine fosbergii Hosaka
end
K
Myrsine helleri (O.Deg. & I.Deg.) H.St.John
end
K
Myrsine juddii Hosaka
end
Myrsine kauaiensis Hillebr.
end
K
Wagner et al. 1990:941 (K)
Myrsine knudsenii (Rock) Hosaka
end
K
Wagner et al. 1990:941 (K)
Myrsine lanaiensis Hillebr.
end
K
O Mo L
M
H
Wagner et al. 1990:941 (K, O, Mo, L, M, H)
Myrsine lessertiana A.DC.
end
K
O Mo L
M
H
Wagner et al. 1990:938 (O, L, EM [as Myrsine emarginata]), 942 (K, O, Mo,
L, M, H [as M. lessertiana]); Wagner & Herbst 2003:35 (NOM:Syn. M.
emarginata)
Myrsine linearifolia Hosaka
end
K
Wagner et al. 1990:937 (K)
O
Wagner et al. 1990:937 (O)
O
Wagner et al. 1990:940 (O); Lorence et al. 1995:41 (K)
Wagner et al. 1990:940 (K)
O
Wagner et al. 1990:940 (O)
Wagner et al. 1990:943 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 120
Scientific Name
Dicot
Myrsinaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Myrsine mezii Hosaka
end
K
Wagner et al. 1990:943 (K); Lorence et al. 1995:42 (REDISCOVER/K)
Myrsine petiolata Hosaka
end
K
Wagner et al. 1990:943 (K)
Myrsine pukooensis (H.Lév.) Hosaka
end
Myrsine punctata (H.Lév.) Wilbur
end
Myrsine sandwicensis A.DC.
end
Myrsine vaccinioides W.L.Wagner, D.R.Herbst &
Sohmer
end
Myrsine wawraea (Mez) Hosaka
end
O Mo L
K
M
Wagner et al. 1990:944 (O, Mo, L, WM)
O
Wagner et al. 1990:944 (K, O)
O Mo L
M
H
M
Wagner et al. 1990:946 (O, Mo, L, M, H)
Wagner et al. 1990:946 (WM)
K
Wagner et al. 1990:946 (K)
Myrtaceae
Angophora costata (Gaertn.) Britten
nat
M
Corymbia calophylla (R.Br. ex Lindl.) K.D.Hill &
L.A.S.Johnson
nat
Corymbia citriodora (Hook.) K.D.Hill & L.A.S.Johnson
nat
K
O Mo
Corymbia ficifolia (F.Muell.) K.D.Hill & L.A.S.Johnson
nat
K
O
Corymbia gummifera (Sol. ex Gaertn.) K.D.Hill &
L.A.S.Johnson
nat
K
O
Eucalyptus albens Benth.
nat
O
Starr et al. 2006:37 (NEWNAT/EM, DESCR)
H
Wagner et al. 1990:951 (O, H [as Eucalyptus calophylla]); Hill & Johnson
1995:240/Herbst & Wagner 1999:24 (COMBNOV:Syn. E. calophylla)
M
H
Wagner et al. 1990:952 (K, O, Mo, M, H [as Eucalyptus citriodora]); Hill &
Johnson 1995:388/Herbst & Wagner 1999:24 (COMBNOV:Syn. E.
citriodora); Staples & Herbst 2005:417, 420 (KEY [as E. citriodora]), 421
(DESCR)
M
H
Wagner et al. 1990:953 (K, O, M, H [as Eucalyptus ficifolia]); Hill &
Johnson 1995:245/Herbst & Wagner 1999:24 (COMBNOV:Syn. E. ficifolia);
Staples & Herbst 2005:417, 420 (KEY [as E. ficifolia]), 421 (DESCR)
Wagner et al. 1990:956 (K, O [as Eucalyptus gummifera]); Hill & Johnson
1995:233/Herbst & Wagner 1999:24 (COMBNOV:Syn. E. gummifera)
O
H
Wagner et al. 1990:950 (O, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 121
Scientific Name
Dicot
Myrtaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Eucalyptus amygdalina Labill.
nat
M
H
Wagner et al. 1990:950 (M, H)
Eucalyptus botryoides Sm.
nat
K
M
H
Wagner et al. 1990:951 (K, H); Oppenheimer 2003:15 (WM); Staples &
Herbst 2005:417, 420 (KEY), 420 (DESCR)
Eucalyptus bridgesiana R.T.Baker
nat
K
O Mo
H
Wagner et al. 1990:951 (K, O, Mo, H)
Eucalyptus camaldulensis Dehnh.
nat
K
O Mo
H
Wagner et al. 1990:951 (K, O, Mo, M, H); Staples & Herbst 2005:417, 420
(KEY), 420 (DESCR)
Eucalyptus cinerea F.Muell. ex Benth.
nat
K
H
Wagner et al. 1990:952 (K, H); Staples & Herbst 2005:417, 420 (KEY), 421
(DESCR)
Eucalyptus cladocalyx F.Muell.
nat
H
Wagner et al. 1990:952 (O?, Mo? [forestry plantings, not yet vouchered as
naturalizing], H); Staples & Herbst 2005:417, 420 (KEY), 423 (DESCR)
Eucalyptus cornuta Labill.
nat
H
Wagner et al. 1990:952 (H); Staples & Herbst 2005:417, 420 (KEY), 423
(DESCR)
Eucalyptus crebra F.Muell.
nat
K
O Mo L
Eucalyptus deanei Maiden
nat
K
O Mo
H
Wagner et al. 1990:953 (K, O, Mo, H)
Eucalyptus deglupta Blume
nat
K
O Mo
H
Wagner et al. 1990:953 (K, O, Mo, H); Staples & Herbst 2005:417, 420
(KEY), 421 (DESCR)
Eucalyptus globulus Labill. ssp. globulus
nat
K
O
M
H
Wagner et al. 1990:954 (K, O, M, H)
Eucalyptus globulus Labill. ssp. maidenii (F.Muell.)
J.B.Kirkp.
nat
M
H
Wagner et al. 1990:954 (M, H)
Eucalyptus gomphocephala DC.
nat
O
Eucalyptus goniocalyx F.Muell. ex Miq.
nat
O
M
O? Mo?
Wagner et al. 1990:953 (K, O, Mo); Staples & Herbst 2005:417, 420 (KEY),
421 (DESCR); Oppenheimer 2008:30 (L)
Wagner et al. 1990:954 (O)
M
H
Wagner et al. 1990:954 (O, H); Starr & Starr 2011:28 (EM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 122
Scientific Name
Dicot
Myrtaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Eucalyptus marginata Donn ex Sm.
nat
K
O Mo
M
H
Wagner et al. 1990:956 (K, O, Mo, M, H)
Eucalyptus microcorys F.Muell.
nat
K
O Mo
M
H
Wagner et al. 1990:956 (K, O, Mo, M, H)
Eucalyptus paniculata Sm.
nat
K
O Mo
M
H
Wagner et al. 1990:956 (K, O, Mo, M, H); Staples & Herbst 2005:417, 420
(KEY), 422 (DESCR)
Eucalyptus pilularis Sm.
nat
K
O Mo L
M
H
Wagner et al. 1990:957 (K, O, Mo, M, H); Oppenheimer 2008:30 (L)
Eucalyptus punctata DC.
nat
M
Oppenheimer 2004:14 (NEWNAT/WM, DESCR)
Eucalyptus raveretiana F.Muell.
nat
M
Wagner et al. 1990:957 (EM)
Eucalyptus resinifera Sm.
nat
Eucalyptus robusta Sm.
nat
Eucalyptus rudis Endl.
O
M
H
Wagner et al. 1990:957 (O, M, H)
K
O Mo L
M
H
Wagner et al. 1990:957 (K, O, Mo, M, H); Staples & Herbst 2005:417, 420
(KEY), 422 (DESCR); Oppenheimer 2007:28 (L)
nat
K
O
M
H
Wagner et al. 1990:958 (K, O, M, H)
Eucalyptus saligna Sm.
nat
K
O Mo
M
H
Wagner et al. 1990:958 (K, O, Mo, M, H); Staples & Herbst 2005:417, 420
(KEY), 422 (DESCR)
Eucalyptus sideroxylon A.Cunn. ex Woolls ssp.
sideroxylon
nat
K
O Mo
M
Eucalyptus tereticornis Sm.
nat
K
O Mo
M
Eucalyptus viminalis Labill. ssp. viminalis
nat
Eugenia brasiliensis Lam.
nat
Eugenia koolauensis O.Deg.
end
O
M
O Mo
Wagner et al. 1990:958 (K, O, Mo, M)
H
Wagner et al. 1990:958 (K, O, Mo, M, H); Staples & Herbst 2005:417, 420
(KEY), 422 (DESCR)
H
Wagner et al. 1990:959 (O, H)
H
Staples & Herbst 2005:417 (KEY), 423 (DESCR); Starr & Starr 2011:29
(NEWNAT/EM, DESCR); Parker & Parsons 2012b:70 (H)
Wagner et al. 1990:960 (O, Mo)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 123
Scientific Name
Dicot
Myrtaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Eugenia reinwardtiana (Blume) DC.
ind
K
O Mo
M
Wagner et al. 1990:960 (K, O, Mo, M)
Eugenia uniflora L.
nat
K
O Mo L
M
Wagner et al. 1990:961 (K, Mo); Oppenheimer 2003:16 (EM, WM); Staples
& Herbst 2005:417 (KEY), 423 (DESCR); Frohlich & Lau 2010:12 (O);
Starr et al. 2010:65 (L)
Leptospermum laevigatum (Sol. ex Gaertn.) F.Muell.
nat
Leptospermum morrisonii Joy Thomps.
nat
Leptospermum petersonii F.M.Bailey
nat
Leptospermum polygalifolium Salisb. ssp.
polygalifolium
nat
Leptospermum scoparium J.R.Forst. & G.Forst.
nat
K
O
L
M
Lophostemon confertus (R.Br.) Peter G.Wilson &
J.T.Waterh.
nat
K?
O Mo L
M
Melaleuca armillaris (Sol. ex Gaertn.) Sm.
nat
Melaleuca quinquenervia (Cav.) S.T.Blake
nat
O
L
Wagner et al. 1990:963 (L); Thompson 1989:372 (KEY, DESCR);
Herbarium Pacificum Staff 1999:5 (O? [vouchered only by cultivated coll. in
1956 from Honolulu], L); Staples et al. 2002:13 (O)
M
Thompson 1989:402 (KEY, DESCR); Herbarium Pacificum Staff 1999:5
(NEWNAT/EM, NOM: Leptospermum laevigatum misapplied, in part)
H
O
Wagner et al. 1990:961 (O [as Leptospermum flavescens]); Thompson
1989:396 (KEY, DESCR)/Herbarium Pacificum Staff 1999:6 (NOM:Syn. L.
flavescens)
Wagner et al. 1990:963 (K, O, L); Thompson 1989:415 (KEY, DESCR);
Oppenheimer 2004:14 (EM); Staples & Herbst 2005:417 (KEY), 424
(DESCR)
H?
M
K
Thompson 1989:393 (KEY, DESCR); Herbarium Pacificum Staff 1999:5
(NEWNAT/H, NOM: Leptospermum flavescens misapplied, in part)
O Mo L
M
Wagner et al. 1990:963 (K?, O, Mo?, L?, M?, H? [widespread forestry
planting, confirmed naturalized only on O‘ahu]); Oppenheimer 2004:14
(EM); Staples & Herbst 2005:417 (KEY), 431 (DESCR); Oppenheimer
2007:28 (L); Oppenheimer 2010:37 (Mo)
Wagner et al. 1990:964 [forestry planting, not documented as naturalizing];
Staples & Herbst 2005:417, 425 (KEY), 425 (DESCR); Starr & Starr 2012:89
(NEWNAT/EM)
H
Wagner et al. 1990:964 (K, O, Mo, M, H); Staples & Herbst 2005:417, 425
(KEY), 425 (DESCR); Oppenheimer 2010:37 (L)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 124
Scientific Name
Dicot
Myrtaceae
Melaleuca styphelioides Sm.
Status
Kl Le Ni K
O Mo L
M Ka H
?
Citation
Wagner et al. 1990:964 [forestry planting, not documented as naturalizing],
DESCR; Staples & Herbst 2005:417, 425 (KEY), 426 (DESCR)
Metrosideros kermadecensis W.R.B.Oliv.
nat
M
Starr et al. 2004:25 (NEWNAT/EM)
Metrosideros macropus Hook. & Arn.
end
Metrosideros polymorpha Gaudich. var. dieteri
J.W.Dawson & Stemmerm.
end
K
Metrosideros polymorpha Gaudich. var. glaberrima
(H.Lév.) H.St.John
end
K
Metrosideros polymorpha Gaudich. var. incana (H.Lév.)
H.St.John
end
Metrosideros polymorpha Gaudich. var. macrophylla
(Rock) H.St.John
end
Metrosideros polymorpha Gaudich. var. newellii (Rock)
H.St.John
end
Metrosideros polymorpha Gaudich. var. polymorpha
end
Metrosideros polymorpha Gaudich. var. pseudorugosa
Skottsb.
end
Metrosideros polymorpha Gaudich. var. pumila
(A.Heller) J.W.Dawson & Stemmerm.
end
Metrosideros rugosa A.Gray
end
O
Wagner et al. 1990:969 (O)
Metrosideros tremuloides (A.Heller) Knuth
end
O
Wagner et al. 1990:970 (O)
O
Wagner et al. 1990:965 (O)
Wagner et al. 1990:967 (K)
O Mo L
M
H
Wagner et al. 1990:967 (K, O, Mo, L, M, H)
O Mo L
M
H
Wagner et al. 1990:969 (O, Mo, L, M, H)
M
H
Wagner et al. 1990:969 (H); Oppenheimer & Bartlett 2002:10 (EM, WM)
H
Wagner et al. 1990:969 (H)
H
Wagner et al. 1990:969 (O, Mo, L, M, H); Staples & Herbst 2005:417
(KEY), 426 (DESCR [as Metrosideros polymorpha])
O Mo L
K
O Mo
M
M
Wagner et al. 1990:969 (WM)
M
Wagner et al. 1990:969 (K, Mo, M); Kennedy et al. 2010:22 (O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 125
Scientific Name
Dicot
Myrtaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Mo L
M
Citation
Metrosideros waialealae (Rock) Rock var. fauriei
(H.Lév.) J.W.Dawson & Stemmerm.
end
Wagner et al. 1990:970 (Mo, L); Oppenheimer et al. 1999:9 (WM)
Metrosideros waialealae (Rock) Rock var. waialealae
end
K
Pimenta dioica (L.) Merr.
nat
K
Pimenta racemosa (Mill.) J.W.Moore
nat
Psidium cattleianum Sabine
nat
K
O Mo L
M
H
Wagner et al. 1990:971 (K, O, Mo, L, M, H); Staples & Herbst 2005:417
(KEY), 428 (DESCR)
Psidium guajava L.
nat
K
O Mo L
M
H
Wagner et al. 1990:972 (K, O, Mo, L, M, H); Staples & Herbst 2005:417
(KEY), 428 (DESCR)
Rhodomyrtus tomentosa (Aiton) Hassk.
nat
K
O
H
Wagner et al. 1990:972 (K, O, H); Staples et al. 2002:13 (L); Staples &
Herbst 2005:417 (KEY), 431 (DESCR)
Syncarpia glomulifera (Sm.) Nied.
nat
K? O? Mo L?
M
H?
Wagner et al. 1990:948 (K?, O?, Mo?, L?, M?, H? [forestry planting,
reportedly escaping at Kamakou, Moloka‘i]); Oppenheimer 2003:16
(NEWNAT/WM, DESCR); Oppenheimer 2010:37 (Mo)
Syzygium cumini (L.) Skeels
nat
K
M
H
Wagner et al. 1990:975 (K, O, Mo, L, M, H); Staples & Herbst 2005:417,
429 (KEY), 430 (DESCR)
Syzygium grande (Wight) Walp.
nat
Syzygium jambos (L.) Alston
nat
K
O Mo L
M
H
Wagner et al. 1990:975 (K, O, Mo, L, M, H); Staples & Herbst 2005:417,
429 (KEY), 429 (DESCR)
Syzygium malaccense (L.) Merr. & L.M.Perry
pol
K
O Mo
M
H
Wagner et al. 1990:975 (K, O, Mo, M, H); Staples & Herbst 2005:417, 429
(KEY), 429 (DESCR)
Wagner et al. 1990:970 (K)
O
M
H
O
Lorence et al. 1995:42 (NEWNAT/K, DESCR); Starr et al. 2003:28 (EM);
Staples & Herbst 2005:417 (KEY), 427 (DESCR); Frohlich & Lau 2012:41
(O); Parker & Parsons 2012b:70 (H)
Staples & Herbst 2005:417 (KEY), 427 (DESCR [as var. racemosa]);
Frohlich & Lau 2012:41 (NEWNAT/O, DESCR)
L
O Mo L
O
Daehler & Baker 2006:9 (NEWNAT/O, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 126
Scientific Name
Dicot
Myrtaceae
Status
Syzygium sandwicense (A.Gray) Müll.Berol.
end
Syzygium syzygioides (Miq.) Merr. & L.M.Perry
nat
Kl Le Ni K
K
O Mo L
M Ka H
O Mo L
M
Citation
Wagner et al. 1990:976 (K, O, Mo, L, M); Wagner et al. 2012:55 (author
change [from " (A.Gray) Nied."] & spelling correction [from "sandwicensis"])
O
Daehler & Baker 2006:9 (NEWNAT/O, DESCR)
Nyctaginaceae
Boerhavia acutifolia (Choisy) J.W.Moore
ind
Ni
Boerhavia coccinea Mill.
nat
Boerhavia herbstii Fosberg
end
Boerhavia repens L.
ind
Mirabilis jalapa L.
nat
Pisonia brunoniana Endl.
ind
Pisonia grandis R.Br.
ind
Pisonia sandwicensis Hillebr.
end
K
O Mo L
M
H
Wagner et al. 1990:986 (K, O, Mo, L, M, H)
Pisonia umbellifera (G.Forst.) Seem.
ind
K
O Mo L
M
H
Wagner et al. 1990:987 (K, O, Mo, L, M, H)
Pisonia wagneriana Fosberg
end
K
Kl Le Ni
K
O Mo L
M Ka
H
Wagner et al. 1990:978 (Ni, O, Mo, L, M, Ka, H [as Boerhavia glabrata]);
Lorence & Flynn 1999:5 (K); Herbst & Wagner 1999:24 (NOM: B. glabrata
misapplied)
K
O Mo L
M Ka
H
Wagner et al. 1990:978 (K, O, M, Ka, H); Oppenheimer & Bartlett 2002:10
(L); Oppenheimer 2003:16 (Mo)
O
L
M Ka
H
Wagner et al. 1990:980 (O, L, M, Ka, H)
K
O Mo L
M Ka
H
Wagner et al. 1990:981 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:24 (Le)
K
O Mo L
M
H
Wagner et al. 1990:982 (K, O, L, M, H); Oppenheimer 2003:16 (Mo); Staples
& Herbst 2005:434 (DESCR)
O Mo L
M
H
Wagner et al. 1990:984 (O, Mo, L, M, H)
M?
Wagner et al. 1990:985 (M? [based on Macrae coll. in 1825, not vouchered
since]); Staples & Herbst 2005:434 (DESCR)
Wagner et al. 1990:988 (K)
Nymphaeaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 127
Scientific Name
Dicot
Nymphaeaceae
Status
Nymphaea Xdaubenyana W.T.Baxter ex Daubeny
nat
Nymphaea caerulea Savigny
nat
Nymphaea capensis Thunb.
nat
Nymphaea lotus L.
nat
Kl Le Ni K
O Mo L
M Ka H
M
Citation
Oppenheimer 2003:17 (NEWNAT/WM); Staples & Herbst 2005:435 (KEY),
436 (DESCR)
H
K
Herbarium Pacificum Staff 1998:11 (NEWNAT/H, DESCR); Staples &
Herbst 2005:435 (KEY, DESCR)
Herbarium Pacificum Staff 1998:11 (NEWNAT/K [as Nymphaea capensis
var. zanzibariensis], DESCR); Staples et al. 2003:15 (NEWNAT/K [as N.
capensis var. capensis], DESCR); Wagner & Herbst 2003:37 (NOM:Syn. N.
capensis var. zanzibariensis); Staples & Herbst 2005:435 (KEY), 436
(DESCR)
O
Staples et al. 2003:16 (NEWNAT/O, DESCR)
Ochnaceae
Ochna serrulata (Hochst.) Walp.
nat
Ochna thomasiana Engl. & Gilg
nat
Sauvagesia erecta L.
nat
H
K
O
L
Mo
Herbarium Pacificum Staff 1998:12 (NEWNAT/H, DESCR); Staples &
Herbst 2005:437 (KEY), 438 (DESCR)
M
Wagner et al. 1990:989 (O? [as Ochna kirkii; noted escaping cultivation]);
Herbarium Pacificum Staff 1998:12 (NOM: O. kirkii misapplied, O?, EM?
[no bona fide naturalized vouchers yet made], DESCR); Imada et al. 2000:13
(NEWNAT/O); Oppenheimer 2003:17 (WM); Oppenheimer 2004:14 (EM);
Staples & Herbst 2005:437 (KEY, DESCR); Starr et al. 2006:37 (L); Frohlich
& Lau 2012:41 (K)
M
Wagner et al. 1990:989 (Mo); Starr et al. 2004:25 (EM)
M
Oppenheimer 2003:17 (NEWNAT/EM, DESCR)
Oleaceae
Fraxinus griffithii C.B.Clarke
nat
Fraxinus uhdei (Wenz.) Lingelsh.
nat
K
O Mo L
M
H
Wagner et al. 1990:991 (K, O, Mo, L, M, H); Staples & Herbst 2005:438
(KEY), 444 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 128
Scientific Name
Dicot
Oleaceae
Status
Jasminum fluminense Vell.
nat
Jasminum multiflorum (Burm.f.) Andrews
Kl Le Ni K
O Mo L
M Ka H
O
M
Citation
H
Wagner et al. 1990:990 (H? [noted spreading in Ho‘okena area]); Herbarium
Pacificum Staff 1996:5 (NEWNAT/O, H); Starr et al. 2002:22 (EM); Staples
& Herbst 2005:438, 439 (KEY), 439 (DESCR)
nat
H
Staples & Herbst 2005:438, 439 (KEY), 439 (DESCR); Parker & Parsons
2012b:71 (NEWNAT/H, DESCR)
Jasminum polyanthum Franch.
nat
H
Imada 2007:38 (NEWNAT/H, DESCR)
Ligustrum lucidum W.T.Aiton
nat
H
Staples et al. 2003:16 (NEWNAT/H, DESCR); Staples & Herbst 2005:438,
441 (KEY), 442 (DESCR)
Ligustrum sinense Lour.
nat
K
H
Lorence & Flynn 1999:5 (NEWNAT/K); Herbarium Pacificum Staff 1999:6
(H); Staples & Herbst 2005:438, 441 (KEY), 442 (DESCR)
Nestegis sandwicensis (A.Gray) O.Deg., I.Deg. &
L.A.S.Johnson
end
K
O Mo L
M
H
Wagner et al. 1990:992 (K, O, Mo, L, M, H)
Noronhia emarginata (Lam.) Poir.
nat
K
O?
M
Olea europaea L. ssp. cuspidata (Wall. ex G.Don) Cif.
nat
K
O
Olea europaea L. ssp. europaea
nat
L
Flynn & Lorence 2002:15 (NEWNAT/K, DESCR); Starr et al. 2003:29 (EM,
WM, DESCR); Staples & Herbst 2005:438 (KEY), 442 (DESCR); Daehler &
Baker 2006:17 (O? [adventive])
M
H
Wagner et al. 1990:992 (H [as Olea europaea subsp. africana]); Lorence et al.
1995:42 (K); Green & Wickens 1989:295/Herbst & Wagner 1999:24
(NOM:Syn. O. europaea subsp. africana); Starr et al. 1999:13 (EM); Staples
& Herbst 2005:438 (KEY), 443 (DESCR); Frohlich & Lau 2010:12 (O);
Starr et al. 2010:65 (L)
M
H
Wagner et al. 1990:992 (H); Starr et al. 1999:13 (EM); Staples & Herbst
2005:438 (KEY), 443 (DESCR)
M
H
Wagner et al. 1990:995 (K, O, M, H); Wagner & Herbst 1995:22 (K)
Onagraceae
Epilobium billardierianum Ser. ssp. cinereum (A.Rich.)
P.H.Raven & Engelhorn
nat
K
O
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 129
Scientific Name
Dicot
Onagraceae
Epilobium ciliatum Raf. ssp. ciliatum
Fuchsia Xhybrida Vilm.
Status
Kl Le Ni K
O Mo L
nat
M
?
Fuchsia boliviana Carrière
nat
K
Fuchsia magellanica Lam.
nat
K
Fuchsia paniculata Lindl.
nat
Ludwigia octovalvis (Jacq.) P.H.Raven
pol?
K
O Mo L
Ludwigia palustris (L.) Elliott
nat
K
O Mo
Oenothera affinis Cambess.
nat
Oenothera biennis L.
nat
Oenothera glazioviana Micheli
M Ka H
Citation
H
Wagner et al. 1990:995 (H); Wood 2007:16 (EM); Oppenheimer 2011:8
(WM); Wagner et al. 2012:56 (add "subsp. ciliatum")
H?
Wagner et al. 1990:997 (H? [as Fuchsia hybrida; cultivated, sometimes
escaping], DESCR); Staples & Herbst 2005:444 (KEY), 445 (DESCR)
Wagner et al. 1990:996 (K); Staples & Herbst 2005:444 (KEY), 445
(DESCR)
M
H
Wagner et al. 1990:996 (K, M, H); Staples & Herbst 2005:444 (KEY), 445
(DESCR)
H
Wagner et al. 1990:997 (H); Staples & Herbst 2005:444 (KEY), 445
(DESCR)
M
H
Wagner et al. 1990:998 (K, O, Mo, L, M, H)
M
H
Wagner et al. 1990:998 (K, O, Mo, M, H)
H
Wagner et al. 1990:1000 (H)
H
Starr et al. 2008:47 (NEWNAT/EM, DESCR); Pratt & Bio 2012:77 (H)
H?
Wagner et al. 1990:1000 (H? [noted as cultivated at Volcano, but not
appearing to be naturalizing], DESCR)
M
?
Oenothera kunthiana (Spach) Munz
nat
O
M
Oenothera laciniata Hill
nat
M
Oenothera speciosa Nutt.
?
M?
Starr et al. 2004:25 (NEWNAT/EM, DESCR); Frohlich & Lau 2010:13 (O)
H
Wagner et al. 1990:1000 (M, H); Starr & Starr 2012:89 (EM, highest elev
record)
Staples & Herbst 2005:446 (DESCR); Starr & Starr 2011:29
(NEWNAT/EM); Wagner et al. 2012:56 (not naturalized; probably persisting
& spreading only by rhizomes)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 130
Scientific Name
Dicot
Onagraceae
Oenothera stricta Ledeb. ex Link ssp. stricta
Status
Kl Le Ni K
O Mo L
M Ka H
nat
Citation
M
H
Wagner et al. 1990:1001 (EM, H)
Oxalidaceae
Oxalis corniculata L.
pol?
Oxalis debilis Kunth var. corymbosa (DC.) Lourteig
nat
Le Ni
K
O Mo L
M Ka
H
Wagner et al. 1990:1002 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:447 (KEY); Wood & LeGrande 2006:24 (Le)
K
O Mo L
M
H
Wagner et al. 1990:1002 (K, O, Mo, L, M, H [as Oxalis corymbosa]); Herbst
& Wagner 1999:25 (NOM:Syn. O. corymbosa = O. debilis); Staples & Herbst
2005:447 (KEY), 448 (DESCR [as O. debilis var. corymbosa])
H
Wagner et al. 1990:1005 (H)
H
Wagner et al. 1990:1005 (Ni, O, Mo, L, M, Ka, H); Lorence et al. 1995:43
(K); Wood & LeGrande 2006:25 (Le)
Papaveraceae
Argemone glauca (Nutt. ex Prain) Pope var. decipiens
Ownbey
end
Argemone glauca (Nutt. ex Prain) Pope var. glauca
end
Argemone mexicana L.
nat
Bocconia frutescens L.
nat
M
H
Wagner et al. 1990:1006 (M, H)
Eschscholzia californica Cham.
nat
M
H
Wagner et al. 1990:1006 (EM, H); Staples & Herbst 2005:449 (DESCR)
Hunnemannia fumariifolia Sweet
nat
M
H
Wagner et al. 1990:1007 (EM); Parker & Parsons 2012a:60 (H)
Le Ni
K
O Mo L
M Ka
K
O Mo
M
Wagner et al. 1990:1005 (K, O, M); Wysong et al. 2007:6 (Mo) [Note:
Argemone ochroleuca Sweet was collected as a roadside weed by the
Degeners at the south end of Kunia Road, O‘ahu, in 1958; in 1985 it was
reidentified as A. mexicana]
Passifloraceae
Passiflora Xviolacea Loisel.
nat
K
O
M
Wagner et al. 1990:1010 (K, O [as Passiflora xcaerulea]); Starr et al. 1999:13
(EM, NOM: P. xcaerulea misapplied); Staples & Herbst 2005:450 (KEY),
451 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 131
Scientific Name
Dicot
Passifloraceae
Status
Passiflora edulis Sims
nat
Passiflora foetida L.
nat
Passiflora laurifolia L.
Kl Le Ni K
O Mo L
M Ka H
Citation
K
O Mo L
M
H
Wagner et al. 1990:1010 (K, O, L, M, H); Hughes 1995:7 (Mo); Staples &
Herbst 2005:450 (KEY, DESCR)
K
O Mo L
M
H
Wagner et al. 1990:1011 (Ni, K, O, M, H); Herbarium Pacificum Staff
1996:6 (Mo); Oppenheimer & Bartlett 2002:10 (L); Staples & Herbst
2005:450 (KEY); Wood & LeGrande 2006:25 (Le)
nat
K
O Mo
M
H
Wagner et al. 1990:1011 (K, O, Mo, H); Meidell et al. 1998:7 (WM); Starr et
al. 1999:13 (EM)
Passiflora ligularis Juss.
nat
K
O
M
H
Wagner et al. 1990:1011 (K, O, L, H); Meidell et al. 1998:7 (WM); Staples &
Herbst 2005:450 (KEY)
Passiflora maliformis L.
nat
K
Lorence et al. 1995:43 (NEWNAT/K, DESCR)
Passiflora manicata (Juss.) Pers.
nat
K
Wagner et al. 1990:1012 (K); Staples & Herbst 2005:450 (KEY), 451
(DESCR)
Passiflora pulchella Kunth
nat
O
H
Wagner et al. 1990:1012 (O, H); Staples & Herbst 2005:450 (KEY)
Passiflora quadrangularis L.
nat
O
H
Wagner et al. 1990:1012 (O, H); Staples & Herbst 2005:450 (KEY)
Passiflora suberosa L.
nat
K
O
H
Wagner et al. 1990:1014 (O, M, H); Lorence et al. 1995:43 (K); Wagner et al.
1997:60 (L); Staples & Herbst 2005:450 (KEY); Starr et al. 2006:38 (EM)
Passiflora subpeltata Ortega
nat
K
O Mo
Passiflora tarminiana Coppens & Barney
nat
K
Passiflora vitifolia Kunth
nat
Le Ni
L
L
M
M
M
O
Wagner et al. 1990:1014 (K, O, M); Oppenheimer 2003:18 (Mo)
H
Wagner et al. 1990:1012 (K, H [as Passiflora mollissima]); Oppenheimer &
Bartlett 2000:7 (EM); Coppens d'Eeckenbrugge et. al. 2001:9/Staples et al.
2003:17 (SPNOV, NOM: P. mollissima misapplied, DESCR); Staples &
Herbst 2005:450 (KEY)
Wagner et al. 1990:1014 (O); Staples & Herbst 2005:450 (KEY), 451
(DESCR)
Phytolaccaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 132
Scientific Name
Dicot
Phytolaccaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Phytolacca dioica L.
nat
Staples & Herbst 2005:452 (DESCR); Frohlich & Lau 2012:43
(NEWNAT/O, DESCR)
Phytolacca octandra L.
nat
K
O Mo L
M
Phytolacca sandwicensis Endl.
end
K
O Mo
M
H
Wagner et al. 1990:1016 (K, O, Mo, M, H)
Rivina humilis L.
nat
K
O Mo
M
H
Wagner et al. 1990:1016 (K, O, H); Meidell et al. 1997:17 (WM);
Oppenheimer 2003:18 (Mo); Staples & Herbst 2005:452 (DESCR);
Oppenheimer 2007:28 (EM)
Wagner et al. 1990:1015 (K, O, Mo, L, M)
Piperaceae
Peperomia alternifolia Yunck.
end
Peperomia blanda (Jacq.) Kunth var. floribunda (Miq.)
H.Huber
ind
Peperomia cookiana C.DC.
end
Peperomia degeneri Yunck.
end
Mo
Peperomia eekana C.DC.
end
Mo
Peperomia ellipticibacca C.DC.
end
Peperomia expallescens C.DC.
end
Peperomia globulanthera C.DC.
end
Peperomia hesperomannii Wawra
end
K
Peperomia hirtipetiola C.DC.
end
K
Peperomia hypoleuca Miq.
end
Ni
Mo L
M
Wagner et al. 1990:1022 (Mo, L, M)
K
O Mo L
M
H
Wagner et al. 1990:1029 (Ni, K, O, Mo, L, M, H [as Peperomia
leptostachya]); Huber 1987:293/Florence 1997:173/Herbst & Wagner
1999:25 (NOM:Syn. P. leptostachya)
K
Mo
M
H
Wagner et al. 1990:1022 (K, Mo, M, H)
Wagner et al. 1990:1022 (Mo)
M
Wagner et al. 1990:1024 (Mo, M)
O
Wagner et al. 1990:1024 (O)
Mo
M
Wagner et al. 1990:1025 (Mo, M)
M
Wagner et al. 1990:1025 (M)
Wagner et al. 1990:1025 (K)
L
M
Wagner et al. 1990:1026 (L, M); Lorence et al. 1995:43 (K)
H
Wagner et al. 1990:1026 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 133
Scientific Name
Dicot
Piperaceae
Status
Kl Le Ni K
Peperomia kipahuluensis H.St.John & Lamoureux
end
Peperomia kokeana Yunck.
end
K
Peperomia latifolia Miq.
end
K
Peperomia ligustrina Hillebr.
O Mo L
M Ka H
M
Citation
Wagner et al. 1990:1028 (EM)
Wagner et al. 1990:1028 (K)
O Mo L
M
H
Wagner et al. 1990:1028 (K, O, Mo, L, M, H)
end
Mo
M
H
Wagner et al. 1990:1029 (Mo, M, H)
Peperomia macraeana C.DC.
end
O Mo L
M
H
Wagner et al. 1990:1031 (O, Mo, L, M, H)
Peperomia mauiensis Wawra
end
Mo L
M
Peperomia membranacea Hook. & Arn.
end
K
O Mo
Peperomia oahuensis C.DC.
end
K
O
Peperomia obovatilimba C.DC.
end
Peperomia pellucida (L.) Kunth
nat
Peperomia remyi C.DC.
end
Peperomia rockii C.DC.
end
Peperomia sandwicensis Miq.
end
Peperomia subpetiolata Yunck.
end
Peperomia tetraphylla (G.Forst.) Hook. & Arn.
ind
Piper aduncum L.
nat
M
Wagner et al. 1990:1031 (Mo, L, WM)
H
Wagner et al. 1990:1032 (K, O)
M
H
O
K
O Mo L
M
H
Wagner et al. 1990:1034 (K, O, Mo, L, M, H)
Wagner et al. 1990:1035 (Mo)
O Mo
Wagner et al. 1990:1035 (K, O, Mo)
M
K
Wagner et al. 1990:1034 (EM, H); Wagner & Herbst 1995:23 (WM);
Oppenheimer & Bartlett 2002:10 (remove WM, incorrectly attributed)
Wagner et al. 1990:1034 (O); Staples & Herbst 2005:453 (KEY), 455
(DESCR)
Mo
K
Wagner et al. 1990:1032 (K, O, Mo, M, H)
O Mo L
M
O?
M
Wagner et al. 1990:1035 (EM); Wood 2012:96 (EXTINCT?)
H
Wagner et al. 1990:1036 (K, O, Mo, L, M, H)
Starr et al. 2004:25 (NEWNAT/EM, DESCR); Frohlich & Lau 2010:14 (O);
Wagner et al. 2012:58 (O plants controlled, not definitely naturalized)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 134
Scientific Name
Dicot
Piperaceae
Status
Piper auritum Kunth
nat
Piper betle L.
nat
Piper methysticum G.Forst.
pol
Kl Le Ni K
K
O Mo L
M Ka H
O
M
H
O
K
Citation
Staples et al. 2006:8 (NEWNAT/ K, O, DESCR); Starr & Starr 2011:29
(EM); Parker & Parsons 2012b:71 (H)
Staples & Herbst 2005:456 (KEY, DESCR); Frohlich & Lau 2008:8
(NEWNAT/O, DESCR)
O Mo
M
Mo
M
H
Wagner et al. 1990:1036 (K, O, Mo, M, H); Staples & Herbst 2005:456
(KEY, DESCR)
Pittosporaceae
Pittosporum argentifolium Sherff
end
Pittosporum confertiflorum A.Gray
end
O
Pittosporum flocculosum (Hillebr.) Sherff
end
O
Pittosporum gayanum Rock
end
K
Pittosporum glabrum Hook. & Arn.
end
K
Pittosporum halophilum Rock
end
Pittosporum hawaiiense Hillebr.
end
H
Wagner et al. 1990:1044 (H)
Pittosporum hosmeri Rock
end
H
Wagner et al. 1990:1044 (H); Staples & Herbst 2005:458 (KEY), 459
(DESCR)
Pittosporum kauaiense Hillebr.
end
K
Wagner et al. 1990:1045 (K)
Pittosporum napaliense Sherff
end
K
Wagner et al. 1990:1045 (K)
Pittosporum pentandrum (Blanco) Merr.
nat
K
L
M
Wagner et al. 1990:1039 (Mo, M)
H
Wagner et al. 1990:1040 (O. L. M, H)
Wagner et al. 1990:1041 (O)
Wagner et al. 1990:1043 (K)
O Mo L
M
Wagner et al. 1990:1043 (K, O, Mo, L, M)
Mo
O
Wagner et al. 1990:1040 (syn. of Pittosporum confertiflorum); Wagner &
Herbst 1999:1887 (RESURRECT/Mo); Wood & Kiehn 2011:469 (KEY,
DESCR)
H
Herbarium Pacificum Staff 1999:6 (NEWNAT/O, H); Staples & Herbst
2005:458 (KEY, DESCR); Frohlich & Lau 2012:43 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 135
Scientific Name
Dicot
Pittosporaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Pittosporum terminalioides Planch. ex A.Gray
end
L
M
H
Wagner et al. 1990:1047 (L, M, H)
Pittosporum undulatum Vent.
nat
L
M
H
Wagner et al. 1990:1048 (L, H); Staples & Herbst 2005:458 (KEY), 459
(DESCR); Starr et al. 2006:38 (EM)
Pittosporum viridiflorum Sims
nat
L
M
H
Wagner et al. 1990:1048 (L, H); Starr et al. 1999:14 (EM); Staples & Herbst
2005:458 (KEY), 459 (DESCR); Frohlich & Lau 2012:43 (O)
H
Wagner et al. 1990:1050 (H)
H
Wagner et al. 1990:1050 (EM, H); Wood 2006:18 (Mo)
H
Wagner et al. 1990:1050 (O); Herbst et al. 2004:9 (H)
H
Wagner et al. 1990:1051 (H)
O
Plantaginaceae
Plantago aristata Michx.
nat
Plantago australis Lam. ssp. hirtella (Kunth) Rahn
nat
Plantago debilis R.Br.
nat
Plantago hawaiensis (A.Gray) Pilg.
end
Plantago lanceolata L.
nat
K
O Mo L
M
H
Wagner et al. 1990:1051 (K, O, Mo, L, M, H)
Plantago major L.
nat
K
O Mo L
M
H
Wagner et al. 1990:1051 (K, O, Mo, L, M, H)
Plantago pachyphylla A.Gray
end
K
O Mo
M
H
Wagner et al. 1990:1052 (K, O, Mo, M, H)
Plantago princeps Cham. & Schltdl. var. anomala Rock
end
K
O
Plantago princeps Cham. & Schltdl. var. laxiflora
A.Gray
end
Plantago princeps Cham. & Schltdl. var. longibracteata
H.Mann
end
Plantago princeps Cham. & Schltdl. var. princeps
end
Mo
M
O
Wagner et al. 1990:1055 (K, O)
Mo
K
M
H
Wagner et al. 1990:1055 (Mo, M, H)
O
Wagner et al. 1990:1055 (K, O)
O
Wagner et al. 1990:1055 (O)
Plumbaginaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 136
Scientific Name
Dicot
Plumbaginaceae
Status
Plumbago auriculata Lam.
nat
Plumbago zeylanica L.
ind
Kl Le Ni K
Kl
Ni
O Mo L
M Ka H
K
O
M
K
O Mo L
M Ka
Citation
Wagner et al. 1990:1056 [cultivated and sometimes persisting]; Herbarium
Pacificum Staff 1999:7 (NEWNAT/EM, DESCR); Staples & Herbst
2005:460 (DESCR); Frohlich & Lau 2012:44 (K, O)
H
Wagner et al. 1990:1056 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:460 (DESCR)
Polemoniaceae
Collomia linearis Nutt.
nat
M
Wagner & Herbst 1995:23 (NEWNAT/EM, DESCR)
Gilia capitata Sims
nat
M
Wagner et al. 1990:1057 (EM)
Polygalaceae
Polygala paniculata L.
nat
Polygala virgata Thunb.
nat
K
O Mo L
M
H
M
Wagner et al. 1990:1058 (O, EM, H); Lorence et al. 1995:48 (K);
Oppenheimer et al. 1999:10 (WM); Oppenheimer 2006:13 (Mo);
Oppenheimer 2007:30 (L)
Starr & Starr 2011:31 (NEWNAT/EM, DESCR)
Polygonaceae
Antigonon leptopus Hook. & Arn.
nat
Coccoloba uvifera (L.) L.
nat
Emex australis Steinh.
nat
Emex spinosa (L.) Campd.
nat
K
O Mo L
M
H
Wagner et al. 1990:1060 (K, O, L, M, H); Staples & Herbst 2005:461 (KEY),
462 (DESCR); Starr et al. 2006:40 (Mo)
O Mo
M
H
Wagner et al. 1990:1059; Herbst 1998:3 (NEWNAT/O); Oppenheimer &
Bartlett 2002:11 (WM); Starr et al. 2002:24 (EM, H); Staples & Herbst
2005:461 (KEY), 462 (DESCR); Wysong et al. 2007:6 (Mo)
H
Parker & Parsons 2012b:71 (NEWNAT/H, DESCR, NOM: Emex spinosa
misapplied, in part)
O Mo
M
Wagner et al. 1990:1060 (O, Mo, M, H); Parker & Parsons 2012b:71 (remove
H, spms. misidentified as Emex australis)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 137
Scientific Name
Dicot
Polygonaceae
Fallopia convolvulus (L.) Á.Löve
Muehlenbeckia complexa (A.Cunn.) Meisn.
Status
Kl Le Ni K
O Mo L
M Ka H
nat
M
?
Wagner et al. 1990:1063 (EM [as Polygonum convolvulus]); Ronse Decraene
& Akeroyd 1988:369/Herbst & Wagner 1999:29 (NOM:Syn. P. convolvulus)
H?
Wagner et al. 1990:1059 (H? [as Muehlenbeckia axillaris; cultivated and
persisting in Hawai‘i Volcanoes National Park], DESCR); Staples & Herbst
2005:461 (KEY), 464 (NOM: M. axillaris misapplied, DESCR)
H
Wagner et al. 1990:1063 (H [as Polygonum capitatum]); Ronse Decraene &
Akeroyd 1988:367/Herbst & Wagner 1999:29 (NOM:Syn. Polygonum
capitatum); Oppenheimer & Bartlett 2002:11 (EM); Staples & Herbst
2005:461 (KEY [as Polygonum]), 464 (DESCR [as P. capitatum]); Frohlich
& Lau 2010:15 (O)
H
Wagner et al. 1990:1063 (H [as Polygonum chinense]); Ronse Decraene &
Akeroyd 1988:367/Herbst & Wagner 1999:29 (NOM:Syn. Polygonum
chinense); Imada et al. 2008:13 (O)
Persicaria capitata (Buch.-Ham. ex D.Don) Masam.
nat
O
Persicaria chinensis (L.) H.Gross
nat
O
Persicaria glabra (Willd.) M.Gómez
nat?
Persicaria punctata (Elliott) Small
nat
Polygonum argyrocoleon Steud. ex Kunze
nat
M
Polygonum aviculare L.
nat
M
H
Wagner et al. 1990:1063 (M, H)
Rumex acetosella L.
nat
K
M
H
Wagner et al. 1990:1066 (K, M, H); Mosyakin & Wagner 1998:42 (KEY)
Rumex albescens Hillebr.
end
K
Rumex brownei Campd.
nat
K
M
Citation
O Mo
M
H
Wagner et al. 1990:1064 (K, O, Mo, M, H [as Polygonum glabrum]); Wilson
1990:632/Herbst & Wagner 1999:30 (NOM:Syn. Polygonum glabrum)
Mo
M
H
Wagner et al. 1990:1064 (H [as Polygonum punctatum]); Herbst & Wagner
1999:30/Hinds & Freeman 2005:586 (NOM:Syn. Polygonum punctatum);
Oppenheimer 2004:16 (EM); Wood 2006:19 (Mo)
Wagner et al. 1990:1062 (EM)
O
Wagner et al. 1990:1066 (K, O); Mosyakin & Wagner 1998:41 (KEY)
M
H
Wagner et al. 1990:1066 (EM, H); Mosyakin & Wagner 1998:42 (KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 138
Scientific Name
Dicot
Polygonaceae
Status
Kl Le Ni K
Rumex conglomeratus Murray
nat
K
Rumex crispus L. ssp. crispus
nat
K
Rumex crispus L. ssp. fauriei (Rech.f.) Mosyakin &
W.L.Wagner
nat
Rumex giganteus W.T.Aiton
end
Rumex obtusifolius L. ssp. obtusifolius
nat
Rumex skottsbergii O.Deg. & I.Deg.
end
Triplaris weigeltiana (Rchb.) Kuntze
nat
O Mo L
M Ka H
Wagner et al. 1997:60 (NEWNAT/K, DESCR); Mosyakin & Wagner
1998:42 (KEY)
O
L
O?
L
Mo
K
Citation
M
H
Wagner et al. 1990:1067 (K, O, L, M, H [as Rumex crispus]); Mosyakin &
Wagner 1998:42 (becomes subsp. crispus with addition of subsp. fauriei,
KEY)
Mosyakin & Wagner 1998:40 (COMBNOV, NEWNAT/O? [single cultivated
coll. in 1993 from Lyon Arboretum], L, NOM:Syn. Rumex fauriei, DESCR)
M
O
H
Wagner et al. 1990:1067 (Mo, M, H); Mosyakin & Wagner 1998:42 (KEY)
H
Wagner et al. 1990:1069 (K, O, H); Mosyakin & Wagner 1998:42 (KEY)
H
Wagner et al. 1990:1069 (H); Mosyakin & Wagner 1998:42 (KEY)
O
Staples & Herbst 2005:461 (KEY), 464 (DESCR); Lau & Frohlich 2012:20
(NEWNAT/O, DESCR)
Portulacaceae
Portulaca lutea Sol. ex G.Forst.
ind
Kl
Ni
K
O Mo L
M
H
Portulaca molokiniensis Hobdy
end
Portulaca oleracea L.
nat
Kl Le Ni
K
O Mo L
M Ka
H
Wagner et al. 1990:1072 (Ni, K, O, Mo, L, M, H); Staples & Herbst
2005:465 (KEY), 466 (DESCR); Wood & LeGrande 2006:27 (Le);
Oppenheimer 2006:13 (Ka)
Portulaca pilosa L.
nat
Kl Le? Ni
K
O Mo
M
H
Wagner et al. 1990:1072 (Ni, K, O, Mo, M, H); Staples & Herbst 2005:465
(KEY), 466 (DESCR); Wood & LeGrande 2006:27 (Le? [reported but
unvouchered])
Portulaca sclerocarpa A.Gray
end
H
Wagner et al. 1990:1074 (L, H)
M Ka
L
Wagner et al. 1990:1071 (O, Mo, L, M, H); Wood 2006:19 (K); Imada
2007:39 (Ni)
Wagner et al. 1990:1072 (EM, Ka); Staples & Herbst 2005:465 (KEY,
DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 139
Scientific Name
Dicot
Portulacaceae
Status
Portulaca sp.A
nat
Portulaca villosa Cham.
end
Talinum fruticosum (L.) Juss.
nat
Talinum paniculatum (Jacq.) Gaertn.
nat
Kl Le Ni K
O Mo L
M Ka H
M
Kl Le?
O Mo L
M Ka
O
Citation
Wagner et al. 1990:1075 (EM)
H
Wagner et al. 1990:1074 (O, Mo, L, M, Ka, H); Wood & LeGrande 2006:27
(Le? [reported by Caum in 1936 but unvouchered])
H
Wagner et al. 1990:1076 (H [as Talinum triangulare]); Wagner & Herbst
1995:25 (O, NOM:Syn. T. triangulare); Staples & Herbst 2005:467 (DESCR)
K
O
M
H
Wagner et al. 1990:1075 (K, O, H); Oppenheimer & Bartlett 2000:8 (WM);
Staples & Herbst 2005:467 (DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:1077 (K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:28 (Le)
Primulaceae
Anagallis arvensis L.
nat
Le
Lysimachia daphnoides (A.Gray) Hillebr.
end
K
Lysimachia filifolia C.N.Forbes & Lydgate
end
K
Lysimachia forbesii Rock
end
Lysimachia glutinosa Rock
end
K
Lysimachia hillebrandii Hook.f. ex A.Gray
end
K
Wagner et al. 1990:1080 (K); St. John 1987g:44 (SPNOV/K [as Lysimachia
kahiliensis], DESCR); Marr & Bohm 1997:265 ([as L. daphnoides] KEY,
DESCR, 272 [as L. kahiliensis] KEY, DESCR); Herbst & Wagner 1999:30
(NOM:Syn. L. kahiliensis)
O
Wagner et al. 1990:1080 (K, O); Marr & Bohm 1997:266 (KEY, DESCR);
Wood 2012:97 (REDISCOVER/K)
O
Wagner et al. 1990:1080 (O); Marr & Bohm 1997:267 (KEY, DESCR)
Wagner et al. 1990:1081 (K); Marr & Bohm 1997:268 (KEY, DESCR)
O Mo L
M
Wagner et al. 1990:1081 (K, O, Mo, L, M); St. John 1987g:45 (SPNOV/K
[as Lysimachia haupuensis], DESCR), 48 (SPNOV/K [as L. ovoidea],
DESCR), 50 (SPNOV/O [as L. waianaeensis], DESCR); Marr & Bohm
1997:268 ([as L. haupuensis] KEY, DESCR), 269 ([as L. hillebrandii] KEY,
DESCR), 274 ([as L. ovoidea] KEY, DESCR), 284 ([as L. waianaeensis]
KEY, DESCR); Herbst & Wagner 1999:30 (NOM:Syns: L. haupuensis, L.
ovoidea, L. waianaeensis)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 140
Scientific Name
Dicot
Primulaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Lysimachia iniki K.L.Marr
end
K
Marr & Bohm 1997:270 (SPNOV/K, KEY, DESCR)/Herbst & Wagner
1999:30
Lysimachia kalalauensis Skottsb.
end
K
Wagner et al. 1990:1082 (K); Marr & Bohm 1997:272 (KEY, DESCR)
Lysimachia lydgatei Hillebr.
end
Lysimachia mauritiana Lam.
ind
Lysimachia maxima (R.Knuth) H.St.John
end
Lysimachia pendens K.L.Marr
end
Lysimachia remyi Hillebr.
end
Lysimachia scopulensis K.L.Marr
end
K
Marr & Bohm 1997:265 (SPNOV/K, KEY, DESCR)/Herbst & Wagner
1999:30
Lysimachia venosa (Wawra) H.St.John
end
K
Wagner et al. 1990:1083 (K); Marr & Bohm 1997:284 (KEY, DESCR);
Wood 2012:98 (EXTINCT?)
M
Ni
K
Mo
M
Wagner et al. 1990:1082 (WM); Marr & Bohm 1997:280 (NOM:syn. of
Lysimachia remyi subsp. remyi [not followed by US website 2012])
H
Mo
Wagner et al. 1990:1082 (Ni, K, Mo, EM, WM, H)
Wagner et al. 1990:1083 (Mo); Marr & Bohm 1997:273 (KEY, DESCR)
K
Marr & Bohm 1997:275 (SPNOV/K, KEY, DESCR)/Herbst & Wagner
1999:30
Mo
M
Wagner et al. 1990:1083 (Mo, M); St. John 1987g:43 (SPNOV/M [as
Lysimachia caliginis], DESCR); Marr & Bohm 1997:275 (KEY, DESCR),
SSPNOV: 277 [as Lysimachia remyi subsp. caliginis/EM, WM, NOM:Syn.
L. caliginis, KEY, DESCR], 278 [as L. remyi subsp. kipahuluensis/EM,
KEY, DESCR], 279 [as L. remyi subsp. remyi/L, WM, KEY, DESCR], 281
[as L. remyi subsp. subherbacea/O, Mo, NOM:Syn. L. subherbacea, KEY,
DESCR]); Herbst & Wagner 1999:31 (NOM:Syns. L. remyi subspp.
caliginis, kipahuluensis, subherbacea)
Proteaceae
Banksia integrifolia L.f.
nat
Grevillea banksii R.Br.
nat
M
Ni
K
O Mo
M
Starr et al. 2006:40 (NEWNAT/EM, DESCR)
H
Wagner et al. 1990:1086 (Ni, K, O, Mo, M, H); Staples & Herbst 2005:468
(KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 141
Scientific Name
Dicot
Proteaceae
Status
Grevillea robusta A.Cunn. ex R.Br.
nat
Macadamia integrifolia Maiden & Betche
nat
Macadamia tetraphylla L.A.S.Johnson
nat
Kl Le Ni K
K
O Mo L
M Ka H
O Mo L
M Ka
H
Mo
Citation
Wagner et al. 1990:1086 (K, O, M, H); Herbarium Pacificum Staff 1999:8
(L); Oppenheimer 2003:22 (Mo); Staples & Herbst 2005:468 (KEY), 469
(DESCR); Starr et al. 2006:40 (Ka)
Wagner et al. 1990:1085; Staples & Herbst 2005:468 (KEY), 470 (DESCR);
Oppenheimer 2010:38 (NEWNAT/Mo, DESCR)
M
Wagner et al. 1990:1085; Oppenheimer 2004:16 (NEWNAT/EM, DESCR);
Staples & Herbst 2005:468 (KEY), 470 (DESCR)
Ranunculaceae
Anemone hupehensis Lemoine var. japonica (Thunb.)
Bowles & W.L.Stern
nat
H
Wagner et al. 1990:1087 (H)
Ranunculus hawaiensis A.Gray
end
M
H
Wagner et al. 1990:1088 (EM, H)
Ranunculus mauiensis A.Gray
end
M
H
Wagner et al. 1990:1089 (K, O? [reported by Hillebrand in 1888 from Mount
Ka‘ala], Mo, M, H)
Ranunculus muricatus L.
nat
H
Wagner et al. 1990:1089 (H)
Ranunculus parviflorus L.
nat
H
Wagner et al. 1990:1089 (H)
Ranunculus plebeius R.Br. ex DC.
nat
H
Wagner et al. 1990:1089 (H)
Ranunculus repens L.
nat
H
Wagner et al. 1990:1090 (H)
K
O? Mo
Rhamnaceae
Alphitonia excelsa (Fenzl) Reissek ex Benth.
nat
Alphitonia ponderosa Hillebr.
end
Colubrina asiatica (L.) Brongn.
ind
O
Ni
Daehler & Baker 2006:10 (NEWNAT/O, DESCR)
K
O Mo L
M
K
O Mo L
M
H
Wagner et al. 1990:1092 (K, O, Mo, L, M, H)
Wagner et al. 1990:1093 (Ni, K, O, Mo); Starr et al. 2003:31 (WM);
Oppenheimer 2007:30 (L)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 142
Scientific Name
Dicot
Rhamnaceae
Status
Colubrina oppositifolia Brongn. ex H.Mann
end
Gouania hillebrandii Oliv.
end
Gouania meyenii Steud.
end
Gouania vitifolia A.Gray
end
Rhamnus californica Eschsch. var. californica
nat
Ziziphus mauritiana Lam.
Kl Le Ni K
O Mo L
M Ka H
O
M
Mo L
K
H
M Ka
Citation
Wagner et al. 1990:1094 (O, H); Lau 1995:10 (WM)
Wagner et al. 1990:1095 (Mo, L, M, Ka)
O
Wagner et al. 1990:1095 (O); Lorence et al. 1995:48 (K)
O
M
H
Wagner et al. 1990:1097 (O, WM, H); Herbarium Pacificum Staff 1996:6
(REDISCOVER/O); Herbarium Pacificum Staff 1998:12 (REDISCOVER/H)
H
Wagner et al. 1990:1097 (H)
?
O?
Staples & Herbst 2005:473 (DESCR); Frohlich & Lau 2010:15
(NEWNAT/O, DESCR); Wagner et al. 2012:61 (O record adventive)
?
O?
Wagner et al. 1990:1099 (O? [collected twice in 1948 at He‘eia Bridge,
present status unknown], DESCR)
Bruguiera sexangula (Lour.) Poir.
nat
O
Wagner et al. 1990:1099 (O [as Bruguiera gymnorrhiza]); Herbst & Wagner
1999:31 (NOM: B. gymnorrhiza misapplied]
Rhizophora mangle L.
nat
K
Acaena exigua A.Gray
end
K
M
Cotoneaster pannosus Franch.
nat
K
M
Rhizophoraceae
Bruguiera parviflora (Roxb.) Wight & Arn. ex Griffith
O Mo L
M
H
Wagner et al. 1990:1099 (K, O, Mo, L, H); Herbarium Pacificum Staff
1999:8 (M); Oppenheimer 2003:22 (EM)
Rosaceae
Wagner et al. 1990:1102 (K, WM); Meidell et al. 1998:6
(REDISCOVER/WM); Wood 2012:98 (EXTINCT?)
H
Wagner et al. 1990:1100 (EM?, H? [persisting near cultivated plants in
Volcano, Hawai‘i and Kula, Maui]); Lorence et al. 1995:49 (NEWNAT/K,
DESCR); Herbarium Pacificum Staff 1999:8 (EM); Staples & Herbst
2005:474 (KEY), 482 (DESCR); Parker & Parsons 2012b:72 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 143
Scientific Name
Dicot
Rosaceae
Status
Kl Le Ni K
K
O Mo L
Eriobotrya japonica (Thunb.) Lindl.
nat
O
Fragaria chiloensis (L.) Duchesne ssp. sandwicensis
(Decne.) Staudt
end
Fragaria vesca L.
nat
Heteromeles arbutifolia (Lindl.) M.Roem.
nat
Osteomeles anthyllidifolia (Sm.) Lindl.
ind
Photinia davidiana (Decne.) Cardot
nat
Prunus campanulata Maxim.
nat
Prunus grisea (C.Müll.) Kalkman
nat
O
Prunus persica (L.) Batsch
nat
O?
Pyracantha angustifolia (Franch.) C.K.Schneid.
nat
M Ka H
L
M
H
Wagner et al. 1990:1100 (H? [occasionally escaping cultivation in Hawai‘i
Volcanoes National Park]); Lorence et al. 1995:49 (NEWNAT/K, DESCR);
Herbarium Pacificum Staff 1999:8 (EM); Staples & Herbst 2005:474 (KEY,
DESCR); Oppenheimer 2007:31 (L); Frohlich & Lau 2008:8 (O); Parker &
Parsons 2012b:72 (H)
M
H
Wagner et al. 1990:1103 (EM, H)
H
Wagner et al. 1990:1104 (K, H)
H
Herbst & Wagner 1996:12 (NEWNAT/H, DESCR); Staples & Herbst
2005:474 (KEY), 482 (DESCR)
M
H
Wagner et al. 1990:1104 (K, O, Mo, L, M, H); Staples & Herbst 2005:474
(KEY), 482 (DESCR)
M?
H
Herbarium Pacificum Staff 1996:6 (NEWNAT/EM? [possibly naturalizing in
Kula Forest Reserve], H); Staples & Herbst 2005:474 (KEY), 483 (DESCR)
K
K
Citation
O Mo L
K
Lorence & Flynn 2006:4 (NEWNAT/K, DESCR)
Daehler & Baker 2006:11 (NEWNAT/O)
K
M
H?
Wagner et al. 1990:1100 (O?, H? [reported in the wild at Peacock Flats,
O‘ahu and Kïpuka Puaulu, Hawai‘i, perhaps persisting plantings or escapes]);
Oppenheimer 2003:22 (NEWNAT/EM, DESCR); Staples & Herbst 2005:474
(KEY), 476 (DESCR)
M
H
Wagner et al. 1990:1105 (K, H); Herbarium Pacificum Staff 1999:9 (K,
remove H [reidentified as Pyracantha koidzumii]); Staples & Herbst
2005:474 (KEY), 477 (DESCR); Starr et al. 2008:48 (EM); Pratt & Bio
2012:78 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 144
Scientific Name
Dicot
Rosaceae
Status
Kl Le Ni K
Pyracantha crenatoserrata (Hance) Rehder
nat
K
Pyracantha koidzumii Rehder
nat
K
Rosa Xdamascena Mill.
nat
Rosa laevigata Michx.
nat
Rubus argutus Link
nat
Rubus discolor Weihe & Nees
O Mo L
M Ka H
Mo
H
Herbarium Pacificum Staff 1999:9 (NEWNAT/K, H, DESCR, NOM:
Pyracantha angustifolia misapplied, in part); Staples & Herbst 2005:474
(KEY), 477 (DESCR)
M
H
Herbarium Pacificum Staff 1999:9 (NEWNAT/K, H, DESCR, NOM:
Pyracantha angustifolia misapplied, in part); Starr et al. 2003:31 (EM);
Staples & Herbst 2005:474 (KEY), 477 (DESCR)
M
H
Staples et al. 2002:14 (NEWNAT/Mo, EM, H); Oppenheimer & Bartlett
2002:11 (WM); Staples & Herbst 2005:474, 480 (KEY), 480 (DESCR)
H
Nagata 1995:12 (NEWNAT/L); Staples & Herbst 2005:474, 480 (KEY), 481
(DESCR); Parker & Parsons 2012a:61 (H)
H
Wagner et al. 1990:1107 (K, O, M, H); Hughes 1995:9 (Mo)
L
K
O Mo
M
nat
O
M
Rubus ellipticus Sm. var. obcordatus Focke
nat
O?
Rubus glaucus Benth.
nat
Rubus hawaiensis A.Gray
end
Rubus macraei A.Gray
end
Rubus niveus Thunb.
nat
K
Rubus rosifolius Sm.
nat
K
K
Mo
O Mo L
Citation
Wagner et al. 1990:1106 (O?, WM? [patches noted at Lanipö Trail, O‘ahu
and Kaulalewelewe, West Maui]); Conant 1996:1 (NEWNAT/O); Starr &
Starr 2011:32 (EM)
H
Wagner et al. 1990:1107 (H); Frohlich & Lau 2007:10 (O); Wagner et al.
2012:62 (O plants controlled, not definitely naturalized)
M
H?
Wagner et al. 1990:1106 (H? [reported as naturalizing in Puna District along
Wright Road, but fertile voucher needed to confirm ID]); Starr et al. 2003:31
(NEWNAT/EM)
M
H
Wagner et al. 1990:1109 (K, Mo, M, H)
M
H
Wagner et al. 1990:1109 (EM, H)
M
H
Wagner et al. 1990:1110 (EM, H); Nagata 1995:12 (H); Flynn & Lorence
1998:6 (K)
M
H
Wagner et al. 1990:1110 (K, O, Mo, L, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 145
Scientific Name
Dicot
Rosaceae
Rubus sieboldii Blume
Rubus ulmifolius Schott var. inermis (Willd.) Focke
Status
Kl Le Ni K
nat
O Mo L
M Ka H
K
?
Citation
H
Wagner et al. 1990:1110 (K); Parker & Parsons 2012b:72 (H)
H?
Wagner et al. 1990:1106 (H? [based on report by Fosberg in 1969 of escape
in Hawai‘i Volcanoes National Park residential area; no vouchers seen])
Rubiaceae
Atractocarpus fitzalanii (F.Muell.) Puttock
nat
Bobea brevipes A.Gray
end
Bobea elatior Gaudich.
end
Bobea sandwicensis (A.Gray) Hillebr.
end
Bobea timonioides (Hook.f.) Hillebr.
end
Cinchona pubescens Vahl
nat
Coffea arabica L.
nat
Coffea liberica Hiern
nat
Coprosma cymosa Hillebr.
end
Coprosma elliptica W.R.B.Oliv.
end
Coprosma ernodeoides A.Gray
end
Coprosma foliosa A.Gray
end
O
Daehler & Baker 2006:11 (NEWNAT/O [as Randia fitzalanii], DESCR);
Puttock 1999:285 (NOM:Syn. R. fitzalanii)
K
O
Wagner et al. 1990:1115 (K, O)
K
O Mo L
M
O Mo L
M
O
M
H
Wagner et al. 1990:1118 (K, O, EM, H); Lorence et al. 1995:49 (K)
M
H
Wagner et al. 1990:1120 (EM, H); Staples & Herbst 2005:485 (KEY), 486
(DESCR)
M Ka
H
Wagner et al. 1990:1120 (K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:484 (KEY), 486 (DESCR)
K
K
O Mo L
H
Wagner et al. 1990:1117 (O, Mo, L, M)
O
Staples et al. 2002:14 (NEWNAT/O, DESCR); Staples & Herbst 2005:484
(KEY), 487 (DESCR)
H
K
Wagner et al. 1990:1124 (H)
Wagner et al. 1990:1124 (K)
M
K
Wagner et al. 1990:1115 (K, O, Mo, L, M, H)
O Mo L
M
H
Wagner et al. 1990:1125 (EM, WM? [single disjunct coll. in 1917 from
Mount ‘Eke bog], H)
Wagner et al. 1990:1125 (K, O, Mo, L, M)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 146
Scientific Name
Dicot
Rubiaceae
Status
Kl Le Ni K
O Mo L
Coprosma kauensis (A.Gray) A.Heller
end
Coprosma longifolia A.Gray
end
Coprosma menziesii A.Gray
end
Coprosma montana Hillebr.
end
Mo
Coprosma ochracea W.R.B.Oliv.
end
Coprosma pubens A.Gray
end
Coprosma rhynchocarpa A.Gray
end
Coprosma ternata W.R.B.Oliv.
end
Coprosma waimeae Wawra
end
Diodia apiculata (Roem. & Schult.) K.Schum.
nat
Galium aparine L.
nat
Galium divaricatum Pourr. ex Lam.
nat
Gardenia brighamii H.Mann
end
O Mo L
Gardenia mannii H.St.John & Kuykendall
end
O
Gardenia remyi H.Mann
end
K
Gynochthodes trimera (Hillebr.) Razafim. & B.Bremer
end
K
M Ka H
K
Citation
Wagner et al. 1990:1126 (K)
O
Wagner et al. 1990:1126 (O)
H
Wagner et al. 1990:1128 (H)
M
H
Wagner et al. 1990:1128 (EM, H); Wysong et al. 2007:7 (Mo)
O Mo L
M
H
Wagner et al. 1990:1129 (O, Mo, L, M, H)
Mo L
M
H
Wagner et al. 1990:1129 (Mo, L, M, H)
M
H
Wagner et al. 1990:1130 (H); Herbarium Pacificum Staff 1996:6 (EM)
Mo
M
Wagner et al. 1990:1130 (Mo); Wagner & Herbst 1995:25 (WM)
K
Wagner et al. 1990:1130 (K)
O
Imada et al. 2000:15 (NEWNAT/O)
Mo
M
M
H
Wagner et al. 1990:1131 (EM); Wagner & Herbst 1995:25 (H)
M
H
Wagner et al. 1990:1132 (O, Mo, L, WM, H)
Wagner et al. 1990:1133 (O)
Mo
O
Starr & Starr 2011:31 (NEWNAT/EM); Frohlich & Lau 2012:45 (Mo, NOM:
Sherardia arvensis misapplied, in part)
M
L
M
H
Wagner et al. 1990:1133 (K, Mo, M, H)
Wagner et al. 1990:1157 (O, L, M [as Morinda trimera]); Lorence et al.
1995:50 (K); Razafimandimbison & Bremer 2011:295/Wagner et al. 2012:65
(COMBNOV:Syn. M. trimera)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 147
Scientific Name
Dicot
Rubiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Kadua acuminata Cham. & Schltdl.
end
K
O Mo L
M
H
Wagner et al. 1990:1138 (K, O, Mo, L, M, H [as Hedyotis acuminata]);
Terrell et al. 2005:832/Kennedy et al. 2010:23 (NOM:Syn. H. acuminata)
Kadua affinis DC.
end
K
O Mo L
M
H
Wagner et al. 1990:1152 (K, O, Mo, L, M, H [as Hedyotis terminalis]);
Terrell et al. 2005:832/Kennedy et al. 2010:23 (NOM:Syn. H. terminalis)
Kadua axillaris (Wawra) W.L.Wagner & Lorence
end
Mo
M
H
Wagner et al. 1990:1146 (Mo, M, H [as Hedyotis hillebrandii]); Terrell et al.
2005:832/Kennedy et al. 2010:23 (COMBNOV:Syn. H. hillebrandii)
Kadua centranthoides Hook. & Arn.
end
K
O Mo L
M
H
Wagner et al. 1990:1139 (K, O, Mo, L, M, H [as Hedyotis centranthoides]);
Terrell et al. 2005:832/Kennedy et al. 2010:23 (NOM:Syn. H. centranthoides)
Kadua cookiana Cham. & Schltdl.
end
K
H
Wagner et al. 1990:1140 (K, H [as Hedyotis cookiana]); Terrell et al.
2005:833/Kennedy et al. 2010:23 (NOM:Syn. H. cookiana)
Kadua cordata Cham. & Schltdl. ssp. cordata
end
K
Kadua cordata Cham. & Schltdl. ssp. remyi (Hillebr.)
W.L.Wagner & Lorence
end
Kadua cordata Cham. & Schltdl. ssp. waimeae (Wawra)
W.L.Wagner & Lorence
end
Kadua coriacea (Sm.) W.L.Wagner & Lorence
end
O Mo
M
Wagner et al. 1990:1151 (K, O, Mo [as Hedyotis schlechtendahliana var.
schlechtendahliana]); Wagner & Lorence 1998:316 (changed from varietal to
subspecific status, KEY); Terrell et al. 2005:833/Kennedy et al. 2010:23
(NOM:Syn. H. schlechtendahliana subsp. schlechtendahliana)
L
Wagner et al. 1990:1151 (L [as Hedyotis schlechtendahliana var. remyi]);
Wagner & Lorence 1998:315 (restored from varietal to subspecific status,
KEY)/Herbst & Wagner 1999:32; Terrell et al. 2005:833/Kennedy et al.
2010:23 (COMBNOV & STATNOV:Syn. H. schlechtendahliana subsp.
remyi)
K
Wagner et al. 1990:1150 (as Hedyotis glaucifolia var. waimeae, syn. of H.
schlechtendahliana var. schlechtendahliana); Wagner & Lorence 1998:316
(RESURRECT/K, COMBNOV & STATNOV [as H. schlechtendahliana
subsp. waimeae, NOM:Syn. H. glaucifolia var. waimeae], KEY/Herbst &
Wagner 1999:31; Terrell et al. 2005:833/Kennedy et al. 2010:23
(COMBNOV:Syn. H. schlechtendahliana subsp. waimeae)
O
M
H
Wagner et al. 1990:1141 (O, M, H [as Hedyotis coriacea]); Terrell et al.
2005:832/Kennedy et al. 2010:23 (COMBNOV:Syn. H. coriacea)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 148
Scientific Name
Dicot
Rubiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Kadua degeneri (Fosberg) W.L.Wagner & Lorence ssp.
coprosmifolia (Fosberg) W.L.Wagner & Lorence
end
O
Wagner et al. 1990:1142 (O [as Hedyotis degeneri var. coprosmifolia]);
Terrell et al. 2005:833/Kennedy et al. 2010:23 (COMBNOV &
STATNOV:Syn. H. degeneri var. coprosmifolia)
Kadua degeneri (Fosberg) W.L.Wagner & Lorence ssp.
degeneri
end
O
Wagner et al. 1990:1142 (O [as Hedyotis degeneri var. degeneri]); Terrell et
al. 2005:833/Kennedy et al. 2010:24 (COMBNOV:Syn. H. degeneri var.
degeneri)
Kadua elatior (H.Mann) A.Heller
end
K
O Mo
Kadua fluviatilis C.N.Forbes
end
K
O
Kadua flynnii (W.L.Wagner & Lorence) W.L.Wagner &
Lorence
end
K
Wagner & Lorence 1998:311 (SPNOV/K, DESCR)/Herbst & Wagner
1999:31 [as Hedyotis flynnii]; Terrell et al. 2005:833/Kennedy et al. 2010:24
(COMBNOV:Syn. H. flynnii)
Kadua foggiana (Fosberg) W.L.Wagner & Lorence
end
K
Wagner et al. 1990:1144 (K [as Hedyotis foggiana]); Terrell et al.
2005:832/Kennedy et al. 2010:24 (COMBNOV:Syn. H. foggiana)
Kadua foliosa Hillebr.
end
M
Wagner et al. 1990:1145 (EM [as Hedyotis foliosa]); Terrell et al.
2005:833/Kennedy et al. 2010:24 (NOM:Syn. H. foliosa)
Kadua formosa Hillebr.
end
M
Wagner et al. 1990:1145 (WM [as Hedyotis formosa]); Terrell et al.
2005:832/Kennedy et al. 2010:24 (NOM:Syn. H. formosa)
Kadua fosbergii (W.L.Wagner & D.R.Herbst)
W.L.Wagner & Lorence
end
Kadua haupuensis Lorence & W.L.Wagner
end
K
Lorence et al. 2010:138 (SPNOV/K, DESCR); Wood 2012:98 (EXTINCT?)
Kadua knudsenii Hillebr.
end
K
Wagner et al. 1990:1147 (K [as Hedyotis knudsenii]); Terrell et al.
2005:832/Kennedy et al. 2010:24 (NOM:Syn. H. knudsenii)
O
M
Wagner et al. 1990:1142 (K, O, Mo, EM [as Hedyotis elatior]); Terrell et al.
2005:833/Kennedy et al. 2010:24 (COMBNOV:Syn. H. elatior)
Wagner et al. 1990:1142 (K, O [as Hedyotis fluviatilis]); Terrell et al.
2005:832/Kennedy et al. 2010:24 (NOM:Syn. H. fluviatilis)
L
Wagner et al. 1990:1145 (O, L [as Hedyotis fosbergii]); Terrell et al.
2005:832/Kennedy et al. 2010:24 (COMBNOV:Syn. H. fosbergii)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 149
Scientific Name
Dicot
Rubiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Mo L
Kadua laxiflora H.Mann
end
Kadua littoralis Hillebr.
end
Kadua parvula A.Gray
end
Kadua st.-johnii (B.C.Stone & Lane) W.L.Wagner &
Lorence
end
K
Wagner et al. 1990:1150 (K [as Hedyotis st.-johnii]); Terrell et al.
2005:833/Kennedy et al. 2010:24 (COMBNOV:Syn. H. st.-johnii)
Kadua tryblium (D.R.Herbst & W.L.Wagner)
W.L.Wagner & Lorence
end
K
Wagner et al. 1990:1155 (K [as Hedyotis tryblium]); Terrell et al.
2005:832/Kennedy et al. 2010:24 (COMBNOV:Syn. H. tryblium)
Mitracarpus hirtus (L.) DC.
nat
Morinda citrifolia L.
pol
Nertera granadensis (L.f.) Druce
K
O Mo
M
Citation
M
Wagner et al. 1990:1148 (Mo, L, WM [as Hedyotis mannii]); Terrell et al.
2005:833/Kennedy et al. 2010:24 (NOM:Syn. H. mannii)
H
O
Ni
Wagner et al. 1990:1147 (K, O, Mo, EM, H [as Hedyotis littoralis]); Terrell
et al. 2005:833/Kennedy et al. 2010:24 (NOM:Syn. H. littoralis)
Wagner et al. 1990:1148 (O [as Hedyotis parvula]); Terrell et al.
2005:833/Kennedy et al. 2010:24 (NOM:Syn. H. parvula)
M
H
Wagner et al. 1990:1156 (H); Starr et al. 2003:31 (EM)
K
O Mo L
M
H
Wagner et al. 1990:1157 (Ni, K, O, Mo, L, M, H); Staples & Herbst
2005:483 (KEY), 493 (DESCR)
ind
K
O Mo
M
H
Wagner et al. 1990:1158 (K, O, Mo, WM, H [as Nertera granadensis]); Heads
1996:388 (COMBNOV {as Coprosma granadensis}, NOM:Syn. N.
granadensis)/Herbst & Wagner 1999:31; Oppenheimer & Bartlett 2000:8
(EM); Wagner et al. 2012:65 (NOM:Syn. C. granadensis)
Oldenlandia corymbosa L.
nat
K
O
M Ka
H
Wagner et al. 1990:1141 (O, EM, H [as Hedyotis corymbosa]); Lorence et al.
1995:50 (K); Oppenheimer & Bartlett 2002:12 (WM); Starr et al. 2002:24
(H); Starr et al. 2006:40 (Ka); Groeninchkx et al. 2009:125/Wagner et al.
2012:64 (NOM:Syn. H. corymbosa)
Oldenlandiopsis callitrichoides (Griseb.) Terrell &
W.H.Lewis
nat
O
M
Wagner et al. 1990:1139 ( O, M [as Hedyotis callitrichoides]); Oppenheimer
& Bartlett 2002:12 (WM); Terrell & Lewis 1990:185/Wagner et al. 2012:63
(COMBNOV:Syn. H. callitrichoides)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 150
Scientific Name
Dicot
Rubiaceae
Status
Kl Le Ni K
K
O Mo L
M Ka H
O
Citation
Paederia foetida L.
nat
M
H
Wagner et al. 1990:1160 (K, O, H [as Paederia scandens]); Wagner & Herbst
1995:25 (NOM:Syn. P. scandens); Starr et al. 1999:14 (EM); Oppenheimer
2003:23 (WM)
Pentas lanceolata (Forssk.) Deflers
nat
M
H
Nagata 1995:12 (NEWNAT/H); Starr et al. 2002:24 (EM); Staples & Herbst
2005:485 (KEY), 495 (DESCR)
Psychotria fauriei (H.Lév.) Fosberg
end
Psychotria grandiflora H.Mann
end
K
Wagner et al. 1990:1163 (K)
Psychotria greenwelliae Fosberg
end
K
Wagner et al. 1990:1164 (K); Herbarium Pacificum Staff 1996:6 (O); Kiehn
& Imada 2010:27 (remove O, spms. reidentified as Psychotria mariniana & P.
hathewayi)
Psychotria hathewayi Fosberg var. brevipetiolata
Fosberg
end
O
Wagner et al. 1990:1165 (O)
Psychotria hathewayi Fosberg var. hathewayi
end
O
Wagner et al. 1990:1165 (O)
Psychotria hawaiiensis (A.Gray) Fosberg var.
hawaiiensis
end
Mo
M
H
Wagner et al. 1990:1165 (Mo, M, H)
Psychotria hawaiiensis (A.Gray) Fosberg var.
hillebrandii (Rock) Fosberg
end
Mo
M
H
Wagner et al. 1990:1165 (Mo, M, H)
Psychotria hawaiiensis (A.Gray) Fosberg var. scoriacea
(Rock) Fosberg
end
M
H
Wagner et al. 1990:1165 (M, H)
Psychotria hexandra H.Mann ssp. hexandra var.
hexandra
end
K
Wagner et al. 1990:1166 (K)
Psychotria hexandra H.Mann ssp. hexandra var. hirta
Wawra
end
K
Wagner et al. 1990:1166 (K)
O
Wagner et al. 1990:1163 (O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 151
Scientific Name
Dicot
Rubiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
K
Citation
Psychotria hexandra H.Mann ssp. hexandra var. kealiae
Fosberg
end
Wagner et al. 1990:1166 (K)
Psychotria hexandra H.Mann ssp. oahuensis O.Deg. &
Fosberg var. hosakana Fosberg
end
O
Wagner et al. 1990:1166 (O)
Psychotria hexandra H.Mann ssp. oahuensis O.Deg. &
Fosberg var. oahuensis
end
O
Wagner et al. 1990:1166 (O)
Psychotria hexandra H.Mann ssp. oahuensis O.Deg. &
Fosberg var. rockii Fosberg
end
O
Wagner et al. 1990:1166 (O)
Psychotria hobdyi Sohmer
end
K
Psychotria kaduana (Cham. & Schltdl.) Fosberg
end
K
O Mo L
M
Wagner et al. 1990:1168 (K, O, Mo, L, M)
Psychotria mariniana (Cham. & Schltdl.) Fosberg
end
K
O Mo L
M
Wagner et al. 1990:1169 (K, O, Mo, L, M)
Psychotria mauiensis Fosberg
end
K
Mo L
M
Psychotria wawrae Sohmer
end
K
Psydrax odorata (G.Forst.) A.C.Sm. & S.P.Darwin
ind
K
O Mo L
M
H
Wagner et al. 1990:1119 (K, O, Mo, L, M, H [as Canthium odoratum]);
Smith 1988:230 (COMBNOV:Syn. C. odoratum)/Herbst & Wagner 1999:31;
Staples & Herbst 2005:484 (KEY), 499 (DESCR)
Richardia brasiliensis Gomes
nat
K
O
M
H
Wagner et al. 1990:1171 (K, O, L, EM, H); Oppenheimer 2006:13 (Mo);
Imada et al. 2008:14 (add WM, remove Mo, EM [spms. reidentified as
Richardia scabra])
Richardia scabra L.
nat
K
O Mo
M
H
Lorence et al. 1995:50 (NEWNAT/K, DESCR); Oppenheimer 2003:23
(WM); Imada et al. 2008:14 (O, Mo, EM, NOM: Richardia brasiliensis
misapplied, in part); Pratt & Bio 2012:79 (H)
Wagner et al. 1990:1166 (K)
H
Wagner et al. 1990:1169 (K, Mo, L, M, H)
Wagner et al. 1990:1170 (K)
L
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 152
Scientific Name
Dicot
Rubiaceae
Status
Serissa japonica (Thunb.) Thunb.
nat
Sherardia arvensis L.
nat
Spermacoce assurgens Ruiz & Pav.
nat
Spermacoce exilis (L.O.Williams) C.D.Adams
nat
Spermacoce latifolia Aubl.
nat
Spermacoce prostrata Aubl.
nat
Kl Le Ni K
K
O Mo L
M Ka H
O Mo L
K
Mo
O
Citation
H
Wagner et al. 1997:61 (NEWNAT/H); Staples & Herbst 2005:484, 485
(KEY), 498 (DESCR)
M
H
Wagner et al. 1990:1171 (EM, H); Oppenheimer 2008:34 (Mo); Frohlich &
Lau 2012:45 (remove Mo, spm. reidentified as Galium aparine)
M
H
Wagner et al. 1990:1173 (K, O, M, H); Lorence et al. 1995:51 (KEY);
Oppenheimer 2003:23 (Mo, L); Lorence & Flynn 2006:4 (KEY)
M
H
Wagner et al. 1990:1173 (M, H [as Spermacoce mauritiana]); Lorence et al.
1995:51 (H, NOM:Syn. S. mauritiana, KEY); Wagner et al. 1997:61
(NEWNAT/H [as S. capitata], DESCR; Lorence & Flynn 2006:4 (NOM: S.
capitata misapplied, KEY)
M
Lorence et al. 1995:51 (NEWNAT/K, KEY); Oppenheimer 2004:17 (EM);
Lorence & Flynn 2006:4 (KEY); Oppenheimer 2010:38 (Mo)
M
H
Wagner et al. 1990:1173 (H? [reportedly collected in Hilo, but voucher not
seen]); Lorence et al. 1995:52 (NEWNAT/H [as Spermacoce ovalifolia],
KEY, DESCR); Wagner et al. 1997:61 (O); Oppenheimer 2003:23 (EM,
WM, NOM: S. ovalifolia misapplied); Lorence & Flynn 2006:4 (KEY)
H
Wagner et al. 1990:1175 (H); Oppenheimer 2003:24 (WM); Starr et al.
2004:27 (EM); Staples & Herbst 2005:500 (KEY), 509 (DESCR)
Rutaceae
Flindersia brayleyana F.Muell.
nat
M
Melicope adscendens (H.St.John & E.P.Hume)
T.G.Hartley & B.C.Stone
end
M
Melicope anisata (H.Mann) T.G.Hartley & B.C.Stone
end
Melicope balloui (Rock) T.G.Hartley & B.C.Stone
end
K
Wagner et al. 1990:1183 (EM [as Pelea adscendens]); Hartley & Stone
1989:120 (COMBNOV:Syn. P. adscendens)
Wagner et al. 1990:1183 (K [as Pelea anisata]); Hartley & Stone 1989:122
(COMBNOV:Syn. P. anisata)
M
Wagner et al. 1990:1183 (EM [as Pelea balloui]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. balloui)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 153
Scientific Name
Dicot
Rutaceae
Status
Kl Le Ni K
O Mo L
M Ka H
K
Citation
Melicope barbigera A.Gray
end
Wagner et al. 1990:1184 (K [as Pelea barbigera]); Hartley & Stone 1989:122
(COMBNOV:Syn. P. barbigera)
Melicope christophersenii (H.St.John) T.G.Hartley &
B.C.Stone
end
O
Wagner et al. 1990:1184 (O [as Pelea christophersenii]); Hartley & Stone
1989:121 (COMBNOV:Syn. P. christophersenii)
Melicope cinerea A.Gray
end
O
Wagner et al. 1990:1185 (O [as Pelea cinerea]); Hartley & Stone 1989:122
(COMBNOV:Syn. P. cinerea)
Melicope clusiifolia (A.Gray) T.G.Hartley & B.C.Stone
end
K
Melicope cruciata (A.Heller) T.G.Hartley & B.C.Stone
end
K
Wagner et al. 1990:1186 (K [as Pelea cruciata]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. cruciata)
Melicope degeneri (B.C.Stone) T.G.Hartley & B.C.Stone
end
K
Wagner et al. 1990:1186 (K [as Pelea degeneri]); Hartley & Stone 1989:122
(COMBNOV:Syn. P. degeneri)
Melicope elleryana (F.Muell.) T.G.Hartley
nat
O
Melicope elliptica A.Gray
end
O Mo
Melicope feddei (H.Lév.) T.G.Hartley & B.C.Stone
end
Melicope haleakalae (B.C.Stone) T.G.Hartley &
B.C.Stone
end
Melicope haupuensis (H.St.John) T.G.Hartley &
B.C.Stone
end
Melicope hawaiensis (Wawra) T.G.Hartley & B.C.Stone
end
O Mo L
M
H
Wagner et al. 1990:1185 (K, O, Mo, L, M, H [as Pelea clusiifolia]); Hartley
& Stone 1989:122 (COMBNOV:Syn. P. clusiifolia)
Frohlich & Lau 2012:46 (NEWNAT/O, DESCR)
M
Wagner et al. 1990:1188 (O, WM [as Pelea elliptica]); Hartley & Stone
1989:122 (COMBNOV:Syn. P. elliptica); Herbarium Pacificum Staff 1996:7
(Mo)
K
Wagner et al. 1990:1188 (K [as Pelea feddei]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. feddei)
M
Wagner et al. 1990:1189 (EM [as Pelea haleakalae]); Hartley & Stone
1989:122 (COMBNOV:Syn. P. haleakalae)
K
Wagner et al. 1990:1189 (K [as Pelea haupuensis]); Hartley & Stone
1989:120 (COMBNOV:Syn. P. haupuensis)
Mo L
M
H
Wagner et al. 1990:1189 (Mo, L, M, H [as Pelea hawaiensis]); Hartley &
Stone 1989:121 (COMBNOV:Syn. P. hawaiensis)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 154
Scientific Name
Dicot
Rutaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Melicope hiiakae (B.C.Stone) T.G.Hartley & B.C.Stone
end
O
Wagner et al. 1990:1206 (as Pelea hiiakae, syn. of P. wawraeana); Hartley &
Stone 1989:122 (COMBNOV:Syn. P. hiiakae); Wagner & Herbst 1999:1894
(RESURRECT/O)
Melicope hosakae (H.St.John) W.L.Wagner &
R.K.Shannon
end
O
Wagner et al. 1990:1190 (O [as Pelea honoluluensis]); Hartley & Stone
1989:121 (COMBNOV [as Melicope honoluluensis]:Syn. P. honoluluensis);
Wagner & Shannon 1998:16 (COMBNOV for P. hosakae, next available
replacement name for invalidly published syns. P. honoluluensis & Melicope
honoluluensis)
Melicope kaalaensis (H.St.John) T.G.Hartley &
B.C.Stone
end
O
Wagner et al. 1990:1190 (O [as Pelea kaalaensis]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. kaalaensis)
Melicope kavaiensis (H.Mann) T.G.Hartley & B.C.Stone
end
K
Melicope knudsenii (Hillebr.) T.G.Hartley & B.C.Stone
end
K
Melicope lydgatei (Hillebr.) T.G.Hartley & B.C.Stone
end
Melicope macropus (Hillebr.) T.G.Hartley & B.C.Stone
end
Melicope makahae (B.C.Stone) T.G.Hartley &
B.C.Stone
end
Melicope molokaiensis (Hillebr.) T.G.Hartley &
B.C.Stone
end
Mo
M
Wagner et al. 1990:1194 (Mo, EM, WM [as Pelea molokaiensis]); Hartley &
Stone 1989:121 (COMBNOV:Syn. P. molokaiensis)
Melicope mucronulata (H.St.John) T.G.Hartley &
B.C.Stone
end
Mo
M
Wagner et al. 1990:1196 (Mo, EM [as Pelea mucronulata]); Hartley & Stone
1989:121 (COMBNOV:Syn. P. mucronulata)
Wagner et al. 1990:1192 (K [as Pelea kavaiensis]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. kavaiensis)
M
O
Wagner et al. 1990:1192 (K, EM [as Pelea knudsenii]); Hartley & Stone
1989:121 (COMBNOV:Syn. P. knudsenii)
Wagner et al. 1990:1193 (O [as Pelea lydgatei]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. lydgatei)
K
Wagner et al. 1990:1193 (K [as Pelea macropus]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. macropus); Wood 2012:98 (EXTINCT?)
O
Wagner et al. 1990:1194 (O [as Pelea makahae]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. makahae)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 155
Scientific Name
Dicot
Rutaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Mo L
Citation
Melicope munroi (H.St.John) T.G.Hartley & B.C.Stone
end
Wagner et al. 1990:1196 (L [as Pelea munroi]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. munroi); Herbst & Wagner 1999:32 (Mo)
Melicope nealae (B.C.Stone) T.G.Hartley & B.C.Stone
end
Melicope oahuensis (H.Lév.) T.G.Hartley & B.C.Stone
end
Melicope obovata (H.St.John) T.G.Hartley & B.C.Stone
end
L? M?
Melicope orbicularis (Hillebr.) T.G.Hartley & B.C.Stone
end
M
Wagner et al. 1990:1197 (EM, WM [as Pelea orbicularis]); Hartley & Stone
1989:121 (COMBNOV:Syn. P. orbicularis)
Melicope ovalis (H.St.John) T.G.Hartley & B.C.Stone
end
M
Wagner et al. 1990:1198 (EM [as Pelea ovalis]); Hartley & Stone 1989:122
(COMBNOV:Syn. P. ovalis)
Melicope ovata (H.St.John & E.P.Hume) T.G.Hartley &
B.C.Stone
end
K
O
Wagner et al. 1990:1198 (K [as Pelea ovata]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. ovata); Wagner & Shannon 1998:17 (O)
Melicope pallida (Hillebr.) T.G.Hartley & B.C.Stone
end
K
O
Wagner et al. 1990:1198 (K, O [as Pelea pallida]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. pallida); Lorence et al. 1995:52 (REDISCOVER/K)
Melicope paniculata (H.St.John) T.G.Hartley &
B.C.Stone
end
K
Melicope peduncularis (H.Lév.) T.G.Hartley &
B.C.Stone
end
K
Melicope pseudoanisata (Rock) T.G.Hartley &
B.C.Stone
end
K
Wagner et al. 1990:1196 (K [as Pelea nealae]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. nealae); Wood 2012:99 (EXTINCT?)
O
Wagner et al. 1990:1196 (O [as Pelea oahuensis]); Hartley & Stone 1989:122
(COMBNOV:Syn. P. oahuensis)
Wagner et al. 1990:1197 (L?, M? [as Pelea obovata; known only from the
type, from Maui or possibly Läna‘i]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. obovata)
Wagner et al. 1990:1199 (K [as Pelea paniculata]); Hartley & Stone 1989:122
(COMBNOV:Syn. P. paniculata); Lorence et al. 1995:53 (REDISCOVER/K)
O Mo
M
Wagner et al. 1990:1199 (K, O, Mo, M [as Pelea peduncularis]); Hartley &
Stone 1989:122 (COMBNOV:Syn. P. peduncularis)
H
Wagner et al. 1990:1200 (H [as Pelea pseudoanisata]); Hartley & Stone
1989:121 (COMBNOV:Syn. P. pseudoanisata)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 156
Scientific Name
Dicot
Rutaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Melicope puberula (H.St.John) T.G.Hartley & B.C.Stone
end
K
Wagner et al. 1990:1202 (K [as Pelea puberula]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. puberula)
Melicope quadrangularis (H.St.John & E.P.Hume)
T.G.Hartley & B.C.Stone
end
K
Wagner et al. 1990:1202 (K [as Pelea quadrangularis]); Hartley & Stone
1989:122 (COMBNOV:Syn. P. quadrangularis); Lorence et al. 1995:53
(REDISCOVER/K); Wood 2012:99 (EXTINCT?)
Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone
end
Melicope reflexa (H.St.John) T.G.Hartley & B.C.Stone
end
Melicope rotundifolia (A.Gray) T.G.Hartley &
B.C.Stone
end
O
Wagner et al. 1990:1203 (O [as Pelea rotundifolia]); Hartley & Stone
1989:122 (COMBNOV:Syn. P. rotundifolia)
Melicope saint-johnii (E.P.Hume) T.G.Hartley &
B.C.Stone
end
O
Wagner et al. 1990:1203 (O [as Pelea saint-johnii]); Hartley & Stone
1989:121 (COMBNOV:Syn. P. saint-johnii)
Melicope sandwicensis (Hook. & Arn.) T.G.Hartley &
B.C.Stone
end
O
Wagner et al. 1990:1204 (O [as Pelea sandwicensis]); Hartley & Stone
1989:121 (COMBNOV:Syn. P. sandwicensis)
Melicope sessilis (H.Lév.) T.G.Hartley & B.C.Stone
end
Mo
M
Melicope volcanica (A.Gray) T.G.Hartley & B.C.Stone
end
Mo L
M
Melicope waialealae (Wawra) T.G.Hartley & B.C.Stone
end
Melicope wailauensis (H.St.John) T.G.Hartley &
B.C.Stone
end
H
Mo
K
Wagner et al. 1990:1202 (H [as Pelea radiata]); Hartley & Stone 1989:121
(COMBNOV:Syn. P. radiata)
Wagner et al. 1990:1203 (Mo [as Pelea reflexa]); Hartley & Stone 1989:122
(COMBNOV:Syn. P. reflexa)
Wagner et al. 1990:1199 (as Pelea sessilis, syn. of P. parvifolia); Hartley &
Stone 1989:121 (NOMNOV: Melicope mauii, Syn. P. parvifolia); Herbst &
Wagner 1999:32/Hartley 2001:288 (COMBNOV:Syns. P. parvifolia, P.
sessilis, Melicope mauii (nom. illeg.)
H
Wagner et al. 1990:1205 (L, EM, H [as Pelea volcanica]); Hartley & Stone
1989:122 (COMBNOV:Syn. P. volcanica); Oppenheimer 2008:34 (Mo)
Wagner et al. 1990:1206 (K [as Pelea waialealae]); Hartley & Stone 1989:122
(COMBNOV:Syn. P. waialealae)
Mo
Wagner et al. 1990:1206 (Mo [as Pelea wailauensis]); Hartley & Stone
1989:122 (COMBNOV:Syn. P. wailauensis)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 157
Scientific Name
Dicot
Rutaceae
Status
Kl Le Ni K
K
O Mo L
M Ka H
O
Citation
Melicope wawraeana (Rock) T.G.Hartley & B.C.Stone
end
Wagner et al. 1990:1206 (K, O [as Pelea wawraeana]); Hartley & Stone
1989:122 (COMBNOV:Syn. P. wawraeana)
Melicope zahlbruckneri (Rock) T.G.Hartley &
B.C.Stone
end
Murraya paniculata (L.) Jack
nat
Platydesma cornuta Hillebr. var. cornuta
end
O
Wagner et al. 1990:1210 (O)
Platydesma cornuta Hillebr. var. decurrens B.C.Stone
end
O
Wagner et al. 1990:1210 (O)
Platydesma remyi (Sherff) O.Deg., I.Deg, Sherff &
B.C.Stone
end
Platydesma rostrata Hillebr.
end
K
Platydesma spathulata (A.Gray) B.C.Stone
end
K
Ruta graveolens L.
nat
Triphasia trifolia (Burm.f.) P.Wilson
nat
Zanthoxylum dipetalum H.Mann var. dipetalum
end
Zanthoxylum dipetalum H.Mann var. tomentosum Rock
end
Zanthoxylum hawaiiense Hillebr.
end
K
Mo L
Zanthoxylum kauaense A.Gray
end
K
O Mo L
H
K
O
M
Kraus 2003:77 (NEWNAT/O, DESCR); Starr et al. 2003:32 (EM, DESCR);
Staples & Herbst 2005:500 (KEY), 508 (DESCR); Frohlich & Lau 2012:46
(K)
H
Wagner et al. 1990:1210 (H)
Wagner et al. 1990:1210 (K)
O
M
H
M
Wagner et al. 1990:1211 (K, O, M, H)
Starr et al. 2004:27 (NEWNAT/EM, DESCR); Staples & Herbst 2005:500
(KEY), 510 (DESCR)
O
K
Wagner et al. 1990:1208 (H [as Pelea zahlbruckneri]); Hartley & Stone
1989:122 (COMBNOV:Syn. P. zahlbruckneri)
Staples & Herbst 2005:500 (KEY), 510 (DESCR); Lau & Frohlich 2012:22
(NEWNAT/O, DESCR)
O Mo
H
Wagner et al. 1990:1214 (K, O, Mo, H)
H
Wagner et al. 1990:1214 (H)
M
H
Wagner et al. 1990:1214 (Mo, L, M, H); Lorence et al. 1995:54 (K); Wood
2007:16 (WM)
M
H
Wagner et al. 1990:1215 (K, O, Mo, L, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 158
Scientific Name
Dicot
Rutaceae
Zanthoxylum oahuense Hillebr.
Status
Kl Le Ni K
end
O Mo L
M Ka H
O
Citation
Wagner et al. 1990:1216 (O)
Santalaceae
Exocarpos gaudichaudii A.DC.
end
Ni
O Mo L
M Ka
H
Exocarpos luteolus C.N.Forbes
end
Exocarpos menziesii Stauffer
end
Santalum album L.
nat
Santalum ellipticum Gaudich.
end
Santalum freycinetianum Gaudich.
end
Santalum haleakalae Hillebr. var. haleakalae
end
Santalum haleakalae Hillebr. var. lanaiense (Rock)
Harbaugh
end
Santalum involutum H.St.John
end
Santalum paniculatum Hook. & Arn. var. paniculatum
end
H
Wagner et al. 1990:1222 (H)
Santalum paniculatum Hook. & Arn. var. pilgeri (Rock)
Stemmerm.
end
H
Wagner et al. 1990:1222 (H)
K
Wagner et al. 1990:1218 (K)
L
Ni
Wagner et al. 1990:1218 (Ni, O, Mo, L, M, Ka, H)
H
Wagner et al. 1990:1218 (L, H)
K?
O
Wagner et al. 1990:1220 (K?, O? [forestry planting, but no naturalized
colls.]); Staples & Herbst 2005:511 (DESCR); Frohlich & Lau 2008:9
(NEWNAT/O, DESCR)
K
O Mo L
M Ka
O
M
Wagner et al. 1990:1221 (O, Mo [as Santalum freycinetianum var.
freycinetianum]); Oppenheimer 2008:34 (WM); Harbaugh et al. 2010:833
(vars. lanaiense & pyrularium transferred out of S. freycinetianum, remove
Mo [now called S. haleakalae var. lanaiense], KEY, DESCR)
M
Wagner et al. 1990:1222 (EM [as Santalum haleakalae]); Harbaugh et al.
2010:833 (NOM: now var. haleakalae with addition of S. haleakalae var.
lanaiense, KEY, DESCR)
M
Wagner et al. 1990:1221 (L, M [as Santalum freycinetianum var. lanaiense]);
Harbaugh et al. 2010:834 (COMBNOV:Syn. S. freycinetianum var.
lanaiense, add Mo, KEY, DESCR)
Mo L
H
K
Wagner et al. 1990:1220 (Ni, K, O, Mo, L, M, Ka, H)
Wagner et al. 1990:1221 (syn. of Santalum freycinetianum var. pyrularium);
Harbaugh et al. 2010:835 (RESURRECT/K, KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 159
Scientific Name
Dicot
Santalaceae
Santalum pyrularium A.Gray
Status
Kl Le Ni K
end
O Mo L
M Ka H
K
Citation
Wagner et al. 1990:1221 (K [as Santalum freycinetianum var. pyrularium]);
Harbaugh et al. 2010:836 (NOM:Syn. S. freycinetianum var. pyrularium,
KEY, DESCR)
Sapindaceae
Alectryon macrococcus Radlk. var. auwahiensis
G.Linney
end
Alectryon macrococcus Radlk. var. macrococcus
end
Allophyllus cobbe (L.) Raeusch.
K
O Mo
M
Wagner et al. 1990:1225 (EM)
M
Wagner et al. 1990:1225 (K, O, Mo, WM); Oppenheimer 2003:24 (EM)
?
O?
Frohlich & Lau 2008:9 (NEWNAT/O); Wagner et al. 2012:69 (O record
adventive)
Cardiospermum grandiflorum Sw.
nat
O
Wagner et al. 1990:1226 (O); Staples & Herbst 2005:512 (KEY), 517
(DESCR)
Cardiospermum halicacabum L.
nat?
Cupaniopsis anacardioides (A.Rich.) Radlk.
nat
Dodonaea viscosa Jacq.
ind
Filicium decipiens (Wight & Arn.) Thwaites
nat
Koelreuteria elegans (Seem.) A.C.Sm. ssp. formosana
(Hayata) F.G.Mey.
nat
Majidea zanquebarica J.Kirk ex Oliv.
nat
Ni
Ni
K
O Mo
M
H
Wagner et al. 1990:1226 (Ni, K, O, Mo, M, H); Staples & Herbst 2005:512
(KEY), 517 (DESCR)
O
M
K
O Mo L
M Ka
H
Wagner et al. 1990:1227 (Ni, K, O, Mo, L, M, H); Staples & Herbst
2005:512 (KEY), 513 (DESCR); Starr et al. 2006:41 (Ka)
K
O
M
H
Wagner et al. 1990:1224; Staples et al. 2002:15 (NEWNAT/O, DESCR);
Oppenheimer 2003:24 (WM, H); Starr et al. 2004:28 (EM); Staples & Herbst
2005:512 (KEY), 514 (DESCR); Frohlich & Lau 2012:46 (K)
O
M
Staples & Herbst 2005:512 (KEY), 517 (DESCR); Frohlich & Lau 2010:15
(NEWNAT/O, DESCR); Starr & Starr 2011:32 (EM, DESCR)
Oppenheimer 2003:25 (NEWNAT/WM, DESCR); Staples & Herbst
2005:512 (KEY), 515 (DESCR); Starr et al. 2008:48 (EM); Frohlich & Lau
2010:16 (O)
H
Staples & Herbst 2005:512 (KEY), 517 (DESCR); Parker & Parsons
2012a:61 (NEWNAT/H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 160
Scientific Name
Dicot
Sapindaceae
Status
Sapindus oahuensis Hillebr. ex Radlk.
end
Sapindus saponaria L.
ind
Kl Le Ni K
K
O Mo L
M Ka H
O
Citation
Wagner et al. 1990:1229 (K, O); Staples & Herbst 2005:512 (KEY), 517
(DESCR)
H
Wagner et al. 1990:1229 (H); Staples & Herbst 2005:512 (KEY), 517
(DESCR)
Sapotaceae
Chrysophyllum oliviforme L.
nat
Ni? K
Planchonella sandwicensis (A.Gray) Pierre
end
K
Sideroxylon persimile (Hemsl.) T.D.Penn.
nat
Sideroxylon polynesicum (Hillebr.) Smedmark &
Anderb.
ind
K
O
M
H
Wagner et al. 1990:1231 (Ni?, K?, O?, M? [vouchered as escape from
cultivation, but not truly naturalized]); Lorence et al. 1995:54 (NEWNAT/K,
DESCR); Wagner et al. 1997:62 (O, H); Oppenheimer 2003:25 (EM, WM);
Staples & Herbst 2005:518 (KEY), 519 (DESCR)
O Mo L
M
H
Wagner et al. 1990:1233 (K, O, Mo, L, M, H [as Pouteria sandwicensis]);
Swenson et al. 2007/Wagner et al. 2012:69 (NOM:Syn. P. sandwicensis)
O
L?
M
O Mo L
M
O
M Ka
O Mo L
M
Starr et al. 2002:24 (NEWNAT/O, L? [single coll. in 1952 from forestry
planting north of Läna‘i City], EM, DESCR); Staples et al. 2003:18 (O,
DESCR); Staples & Herbst 2005:518 (KEY), 522 (DESCR)
H
Wagner et al. 1990:1232 (K, O, Mo, L, M, H [as Nesoluma polynesicum]);
Smedmark & Anderberg 2007:1502/Wagner et al. 2012:69
(COMBNOV:Syn. N. polynesicum)
Scrophulariaceae
Antirrhinum orontium L.
nat
Bacopa monnieri (L.) Wettst.
ind
Buchnera pusilla Kunth
nat
Castilleja arvensis Cham. & Schltdl.
nat
Ni
K
K
O
L
M
Wagner et al. 1990:1236 (O, WM); Staples et al. 2002:15 (Ka); Starr et al.
2002:25 (EM); Starr et al. 2006:41 (Ka); Oppenheimer 2007:31 (WM)
H
Wagner et al. 1990:1237 (Ni, K, O, Mo, L, M, H); Staples & Herbst
2005:524 (KEY), 525 (DESCR)
H
Wagner et al. 1990:1239 (H)
H
Wagner et al. 1990:1239 (K, M, H); Staples et al. 2003:19 (O); Oppenheimer
2007:31 (L)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 161
Scientific Name
Dicot
Scrophulariaceae
Status
Kl Le Ni K
O Mo L
M Ka H
M
Citation
Cymbalaria muralis P.Gaertn., B.Mey. & Scherb.
nat
Wagner et al. 1990:1240 (EM)
Dopatrium junceum (Roxb.) Buch.-Ham. ex Benth.
nat
Hebe speciosa (A.Cunn.) Cockayne & Allan
nat
Linaria canadensis (L.) Dum.Cours. var. texana
(Scheele) Pennell
nat
Lindernia antipoda (L.) Alston
nat
Lindernia crustacea (L.) F.Muell.
nat
Lindernia procumbens (Krock.) Philcox
nat
Lophospermum erubescens D.Don
nat
O
Maurandya antirrhiniflora Humb. & Bonpl. ex Willd.
ssp. antirrhiniflora
nat
O
Mazus pumilus (Burm.f.) Steenis
nat
O?
Orthocarpus purpurascens Benth.
nat
H
Wagner et al. 1990:1245 (H)
Otacanthus azureus (Linden) Ronse
nat
H
Staples & Herbst 2005:524 (KEY), 525 (DESCR); Parker & Parsons
2012a:61 (NEWNAT/H)
Parentucellia viscosa (L.) Caruel
nat
H
Wagner et al. 1990:1246 (H)
Paulownia tomentosa (Thunb.) Steud.
nat
H
Parker & Parsons 2012a:61 (NEWNAT/H)
K
Wagner et al. 1990:1240 (K)
O
O
H
Wagner et al. 1990:1235 (H? [cultivated and perhaps escaping, as at Hawai‘i
Volcanoes National Park]); Shannon & Wagner 1996:14 (NEWNAT/H)
M
H
Wagner et al. 1990:1241 (O, H); Starr et al. 2002:25 (EM)
M
H
Wagner et al. 1990:1242 (H); Oppenheimer 2004:17 (WM)
M
H
Wagner et al. 1990:1242 (O, H); Oppenheimer 2003:25 (WM)
K
Lorence et al. 1995:54 (NEWNAT/K, DESCR)
M
H
Wagner et al. 1990:1244 (O, H); Starr et al. 1999:14 (EM)
Wagner et al. 1990:1244 (O)
M
Wagner et al. 1990:1235 (O? [as Mazus rugosus; reported by Hillebrand in
1888 as naturalized along Honolulu watercourses, not vouchered since];
Oppenheimer 2004:17 (NEWNAT/EM, DESCR, NOM:Syn. M. rugosus)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 162
Scientific Name
Dicot
Scrophulariaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Torenia asiatica L.
nat
Torenia glabra Osbeck
nat
Verbascum blattaria L.
nat
Verbascum thapsus L.
nat
Verbascum virgatum Stokes
nat
Veronica arvensis L.
nat
H
Wagner et al. 1990:1246 (H); Staples et al. 2002:15, 16 (H, KEY); Staples &
Herbst 2005:524 (KEY), 527 (DESCR)
H
Staples et al. 2002:15, 16 (NEWNAT/O, H, NOM: Torenia asiatica
misapplied, KEY); Staples & Herbst 2005:524 (KEY), 527 (DESCR)
H
Wagner et al. 1990:1247 (H)
H
Wagner et al. 1990:1247 (EM, H)
H
Wagner et al. 1990:1247 (H)
M
H
Wagner et al. 1990:1248 (Mo, M, H)
Veronica peregrina L. ssp. xalapensis (Kunth) Pennell
nat
M
H
Wagner et al. 1990:1250 (H); Lorence et al. 1995:54 (K); Starr et al. 2002:25
(EM)
Veronica plebeia R.Br.
nat
M
H
Wagner et al. 1990:1250 (M, H)
Veronica serpyllifolia L.
nat
K
M
H
Wagner et al. 1990:1250 (K, H); Nagata 1995:13 (L); Starr et al. 2003:32
(EM); Oppenheimer 2006:14 (Mo); Lau & Frohlich 2012:22 (O)
nat
K
Brugmansia Xcandida Pers.
nat
K
O Mo
M
Capsicum annuum L.
nat
K
O Mo
M
O
M
Mo
K
O Mo L
Citation
Simaroubaceae
Ailanthus altissima (Mill.) Swingle
Frohlich & Lau 2012:47 (NEWNAT/K, DESCR)
Solanaceae
H
Wagner et al. 1990:1252 (K, O, M, H [as Brugmansia candida]); Hughes
1995:9 (Mo); Staples & Herbst 2005:528 (KEY), 530 (DESCR [as B.
xcandida])
Wagner et al. 1990:1253 (K, O, Mo, M [as Capsicum frutescens]); Staples &
Herbst 2005:528 (KEY), 532 (NOM:Syn. C. frutescens, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 163
Scientific Name
Dicot
Solanaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Cestrum aurantiacum Lindl.
nat
O
Cestrum diurnum L.
nat
K
O Mo
Cestrum nocturnum L.
nat
K
O
Datura metel L.
nat
Datura stramonium L.
nat
Lycium sandwicense A.Gray
ind
Nicandra physalodes (L.) Gaertn.
nat
Nicotiana glauca Graham
nat
Nicotiana tabacum L.
nat
Nothocestrum breviflorum A.Gray
end
Nothocestrum latifolium A.Gray
end
K
O Mo L
M
Nothocestrum longifolium A.Gray
end
K
O Mo L
M
Nothocestrum peltatum Skottsb.
end
K
Physalis angulata L.
nat
K
Kl
Ni
Ni
M
H
Citation
Wagner et al. 1990:1254 (O, M, H); Staples & Herbst 2005:528, 533 (KEY),
533 (DESCR)
Wagner et al. 1990:1254 (K, O, Mo); Staples & Herbst 2005:528, 533
(KEY), 533 (DESCR)
M
H
Wagner et al. 1990:1254 (K, O); Oppenheimer & Bartlett 2000:8 (EM); Starr
et al. 2003:32 (H); Staples & Herbst 2005:528, 533 (KEY), 533 (DESCR);
Oppenheimer 2007:31 (WM); Oppenheimer 2010:38 (L)
O Mo
M
H
Wagner et al. 1990:1255 (O, Mo, M, H)
K
O Mo
M
H
Wagner et al. 1990:1257 (K, O, Mo, M, H)
K
O Mo L
M Ka
H
Wagner et al. 1990:1257 (Ni, K, O, Mo, L, M, Ka, H)
K
O Mo L
M
H
Wagner et al. 1990:1260 (K, O, Mo, M, H); Oppenheimer 2010:38 (L)
O Mo L
M Ka
H
Wagner et al. 1990:1260 (O, L, M, Ka); Wagner & Herbst 1995:26 (H);
Oppenheimer 2003:25 (Mo); Staples & Herbst 2005:528 (KEY), 540
(DESCR)
O Mo L
M Ka
H
Wagner et al. 1990:1262 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:528 (KEY), 540 (DESCR)
H
Wagner et al. 1990:1263 (H)
K
L
Wagner et al. 1990:1263 (K, O, Mo, L, M)
H
Wagner et al. 1990:1264 (K, O, Mo, L, M, H)
Wagner et al. 1990:1264 (K)
O Mo
H
Wagner et al. 1990:1265 (K); Imada et al. 2000:15 (O); Oppenheimer
2003:26 (H); Staples et al. 2006:8 (Mo)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 164
Scientific Name
Dicot
Solanaceae
Physalis ixocarpa Brot. ex Hornem.
Status
nat
Physalis peruviana L.
nat
M Ka H
Mo?
M
K
ind?
Solanum aviculare G.Forst.
nat
Solanum capsicoides All.
nat
Solanum elaeagnifolium Cav.
nat
Solanum incompletum Dunal
end
Solanum linnaeanum Hepper & P.-M.L.Jaeger
nat
Solanum lycopersicum L. var. cerasiforme (Dunal)
D.M.Spooner, G.J.Anderson & R.K.Jansen
nat
Kl Le Ni
?
K
O Mo L
M
O Mo L
M Ka
Wagner & Herbst 1995:26 (NEWNAT/EM, DESCR)
H
Wagner et al. 1990:1265 (K, O, Mo, L, M, H); Staples & Herbst 2005:528
(KEY), 534 (DESCR)
H?
Wagner et al. 1990:1251 (H? [cultivated and persisting near Hilo, but not
naturalized], DESCR); Staples & Herbst 2005:528 (KEY), 535 (DESCR)
H
Wagner et al. 1990:1268 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:528, 536 (KEY), 539 (DESCR); Wood & LeGrande 2006:28 (Le)
L
K
Ni
K
Wagner et al. 1990:1269 (L)
O Mo L
M
O Mo
M
K
Citation
Nagata 1995: 13 (NEWNAT/Mo); Wagner & Herbst 2003:47 [single
individual, unclear if naturalized]
?
Solanum americanum Mill.
Solanum mammosum L.
O Mo L
?
Physalis lanceifolia Nees
Solandra maxima (Sessé & Moç.) P.S.Green
Kl Le Ni K
Mo L
H
Wagner et al. 1990:1269 (K, O, Mo, L, H); Oppenheimer & Bartlett 2000:8
(WM); Staples & Herbst 2005:528, 536 (KEY), 536 (DESCR)
Wagner et al. 1990:1270 (O, Mo, M); Oppenheimer et al. 1999:10 (WM)
M
H
Wagner et al. 1990:1270 (K, Mo, L, M, H); Shaw et al. 1995:372
(REDISCOVER/H)
O Mo L
M Ka
H
Wagner et al. 1990:1271 (O, Mo, L, M, H); Herbarium Pacificum Staff
1998:13 (Ka)
O Mo L
M Ka
H
Wagner et al. 1990:1258 (O, Ka?, H? [as Lycopersicon esculentum; reported
but not vouchered from Ka, H], 1259 (Ni, K, O, Mo, L, M, Ka, H [as L.
pimpinellifolium]); Wagner et al. 1997:62 (NOM: L. esculentum, L.
pimpinellifolium misapplied); Staples & Herbst 2005:528, 535 (KEY), 536
(DESCR)
O?
Staples & Herbst 2005:528, 536 (KEY), 538 (DESCR); Frohlich & Lau
2012:51 (O? [adventive])
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 165
Scientific Name
Dicot
Solanaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Solanum mauritianum Scop.
nat
O
Wagner et al. 1990:1271 (O)
Solanum melongena L.
nat
O
Staples & Herbst 2005:528, 536 (KEY), 538 (DESCR); Frohlich & Lau
2012:47 (NEWNAT/O, DESCR)
Solanum nelsonii Dunal
end
Solanum nigrescens Mart. & Galeotti
nat
Solanum pseudocapsicum L.
nat
Solanum robustum H.Wendl.
nat
M
?
M?
Solanum rostratum Dunal
Ni
K
O Mo
M
O Mo
M
H
Wagner et al. 1990:1273 (Ni, K, O, Mo, M, H)
H
Wagner et al. 1990:1274 (H)
H
Wagner et al. 1990:1274 (O, Mo, M, H); Staples & Herbst 2005:528, 536
(KEY), 538 (DESCR)
Wagner et al. 1990:1274 (EM)
H?
Wagner et al. 1990:1267 (H?, all plts removed from Pöhakuloa area in 1977),
DESCR; Starr et al. 2004:28 (EM?, all plts removed in ‘Ulupalakua in 2001)
Solanum sandwicense Hook. & Arn.
end
K
O
Wagner et al. 1990:1275 (K, O)
Solanum seaforthianum Andrews
nat
K
O Mo
M
H
Wagner et al. 1990:1275 (K, O, Mo, M); Staples et al. 2003:19 (H); Staples
& Herbst 2005:528, 535 (KEY), 538 (DESCR)
Solanum torvum Sw.
nat
K
O
M
H
Wagner et al. 1990:1276 (O, WM); Oppenheimer et al. 1999:10 (EM); Starr
et al. 2003:32 (H); Staples & Herbst 2005:528, 536 (KEY), 539 (DESCR);
Frohlich & Lau 2012:48 (K)
Solanum tuberosum L.
nat
Solanum villosum Mill.
nat
Solanum viride G.Forst. ex Spreng.
pol?
K
Streptosolen jamesonii (Benth.) Miers
nat
K
M
Wagner et al. 1990:1276 (EM); Staples & Herbst 2005:528, 536 (KEY), 539
(DESCR)
H
Imada et al. 2008:14 (NEWNAT/H, DESCR)
Wagner et al. 1990:1278 (K)
H
Lorence & Flynn 1997:12 (NEWNAT/K, DESCR); Staples & Herbst
2005:528 (KEY), 540 (DESCR); Parker & Parsons 2012a:61 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 166
Scientific Name
Dicot
Sterculiaceae
Status
Kl Le Ni K
Heritiera littoralis Dryand.
nat
K
Melochia umbellata (Houtt.) Stapf
nat
K
Sterculia apetala (Jacq.) H.Karst.
O Mo L
M Ka H
Staples & Herbst 2005:541 (KEY), 542 (DESCR); Lorence & Flynn 2006:3
(NEWNAT/K, DESCR)
O
M
nat
O
M
Sterculia foetida L.
nat
O
Waltheria indica L.
ind?
Le Ni
K
Citation
H
Wagner et al. 1990:1279 (H); Oppenheimer 2004:17 (EM, WM); Frohlich &
Lau 2007:11 (O); Frohlich & Lau 2012:48 (K)
Oppenheimer 2003:26 (NEWNAT/WM, DESCR); Staples & Herbst
2005:541 (KEY), 543 (DESCR); Lau & Frohlich 2012:22 (O)
Staples & Herbst 2005:541 (KEY), 543 (DESCR); Frohlich & Lau 2010:16
(NEWNAT/O)
O Mo L
M Ka
H
Wagner et al. 1990:1280 (Ni, K, O, Mo, L, M, Ka, H [incl. syn. Waltheria
pyrolaefolia]); Wood & LeGrande 2006:28 (Le); Saunders 2011:79
(RESURRECT/WM, W. pyrolifolia); Wagner et al. 2012:71 (NOM:Syn. W.
pyrifolia)
Tamaricaceae
Tamarix aphylla (L.) H.Karst.
?
O? Mo? L? M? Ka? H?
Wagner et al. 1990:1280 (O?, Mo?, L?, M?, Ka?, H? [planted as ornamentals,
for erosion control, and as windbreaks, but not known to produce seed],
DESCR); Staples & Herbst 2005:544 (DESCR)
Theaceae
Camellia sinensis (L.) Kuntze
nat
K
Eurya sandwicensis A.Gray
end
K
O Mo
M
H
Wagner et al. 1990:1281; Staples & Herbst 2005:546 (DESCR); Lorence &
Flynn 2006:5 (NEWNAT/K, DESCR); Staples et al. 2006:8 (H)
H
Wagner et al. 1990:1281 (K, O, Mo, M, H)
Thymelaeaceae
Wikstroemia bicornuta Hillebr.
end
Wikstroemia forbesii Skottsb.
end
Wikstroemia furcata (Hillebr.) Rock
end
L
Mo
K
M
Wagner et al. 1990:1285 (L, WM)
Wagner et al. 1990:1285 (Mo)
Wagner et al. 1990:1285 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 167
Scientific Name
Dicot
Thymelaeaceae
Status
Kl Le Ni K
O Mo L
M Ka H
K
Citation
Wikstroemia hanalei Wawra
end
Wagner et al. 1990:1286 (K)
Wikstroemia monticola Skottsb.
end
Wikstroemia oahuensis (A.Gray) Rock var. oahuensis
end
K
Wikstroemia oahuensis (A.Gray) Rock var. palustris
(Hochr.) B.Peterson
end
K
Wikstroemia phillyreifolia A.Gray
end
H
Wagner et al. 1990:1288 (H)
Wikstroemia pulcherrima Skottsb.
end
H
Wagner et al. 1990:1289 (H)
Wikstroemia sandwicensis Meisn.
end
H
Wagner et al. 1990:1289 (H)
Wikstroemia skottsbergiana Sparre
end
K
Wagner et al. 1990:1290 (K)
Wikstroemia uva-ursi A.Gray var. kauaiensis Skottsb.
end
K
Wagner et al. 1990:1290 (K)
Wikstroemia uva-ursi A.Gray var. uva-ursi
end
Wikstroemia villosa Hillebr.
end
O Mo L
M
Wagner et al. 1990:1286 (EM)
M
Wagner et al. 1990:1288 (K, O, Mo, L, M)
Wagner et al. 1990:1288 (K)
O Mo
M
Wagner et al. 1990:1290 (O, Mo, M); Staples & Herbst 2005:547 (DESCR)
M
Wagner et al. 1990:1290 (EM); Oppenheimer 2011:9 (REDISCOVER/EM)
Tiliaceae
Corchorus olitorius L.
?
Grewia micrantha Bojer
nat
Heliocarpus popayanensis Kunth
nat
Trichospermum richii (A.Gray) Seem.
?
K? O?
Wagner et al. 1990:1291 (K?, O? [once naturalized but eradicated in Läwa‘i
Valley, Kaua‘i; escaping along Hale‘iwa, O‘ahu lotus ponds], DESCR);
Staples & Herbst 2005:548 (DESCR)
O
K
O
Lau & Frohlich 2012:22 (NEWNAT/O, DESCR)
L
O?
H
Wagner et al. 1990:1292 (K, O, H); Staples & Herbst 2005:549 (DESCR);
Oppenheimer 2007:31 (L)
Daehler & Baker 2006:17 (O? [adventive], DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 168
Scientific Name
Dicot
Tiliaceae
Status
Kl Le Ni K
O Mo L
Triumfetta rhomboidea Jacq.
nat
Triumfetta semitriloba Jacq.
nat
K
O Mo
nat
K
nat
M Ka H
O Mo
Citation
H
Wagner et al. 1990:1292 (O, Mo, H)
M
H
Wagner et al. 1990:1294 (K, O, M, H); Wysong et al. 2007:7 (Mo)
Mo
M
H
Wagner et al. 1990:1295 (K, Mo, M, H); Staples & Herbst 2005:550
(DESCR)
K
O Mo
M
H
Wagner et al. 1990:1295 (K, Mo); Oppenheimer 2004:18 (EM, WM, H);
Staples & Herbst 2005:550 (DESCR); Frohlich & Lau 2007:11 (O)
nat
K
O Mo L
M
H
Wagner et al. 1990:1296 (K, O, Mo, L, M, H); Staples & Herbst 2005:551
(DESCR)
Boehmeria grandis (Hook. & Arn.) A.Heller
end
K
O Mo L
M
H
Wagner et al. 1990:1300 (K, O, Mo, L, M, H)
Boehmeria nivea (L.) Gaudich.
nat
H
Wagner et al. 1990:1300; Parker & Parsons 2012a:62 (NEWNAT/H)
Hesperocnide sandwicensis (Wedd.) Wedd.
end
H
Wagner et al. 1990:1301 (H)
Neraudia angulata R.S.Cowan var. angulata
end
O
Wagner et al. 1990:1303 (O)
Neraudia angulata R.S.Cowan var. dentata O.Deg. &
R.S.Cowan
end
O
Wagner et al. 1990:1303 (O)
Neraudia kauaiensis (Hillebr.) R.S.Cowan
end
K
Neraudia melastomifolia Gaudich.
end
K
Neraudia ovata Gaudich.
end
Tropaeolaceae
Tropaeolum majus L.
Turneraceae
Turnera ulmifolia L.
Ulmaceae
Trema orientalis (L.) Blume
Urticaceae
Wagner et al. 1990:1303 (K)
O Mo
M
Wagner et al. 1990:1303 (K, O, Mo, WM); Oppenheimer 2008:35
(REDISCOVER/Mo)
H
Wagner et al. 1990:1304 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 169
Scientific Name
Dicot
Urticaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Neraudia sericea Gaudich.
end
Pilea microphylla (L.) Liebm.
nat
Pilea peploides (Gaudich.) Hook. & Arn.
ind
Pipturus albidus (Hook. & Arn.) A.Gray
end
Pipturus forbesii Krajina
end
Pipturus kauaiensis A.Heller
end
K
Wagner et al. 1990:1309 (K)
Pipturus ruber A.Heller
end
K
Wagner et al. 1990:1309 (K)
Procris pedunculata (J.R.Forst. & G.Forst.) Wedd.
Mo L
Le
M Ka
Citation
Wagner et al. 1990:1304 (Mo, L, M, Ka)
K
O Mo L
M
H
Wagner et al. 1990:1306 (K, O, M, H); Oppenheimer & Bartlett 2002:12 (L);
Staples & Herbst 2005:553 (KEY), 554 (DESCR); Oppenheimer 2006:14
(Mo)
K
O Mo L
M
H
Wagner et al. 1990:1306 (K, O, Mo, L, M, H); Wood & LeGrande 2006:28
(Le)
K
O Mo L
M
H
Wagner et al. 1990:1308 (K, O, Mo, L, M, H)
M
?
Wagner et al. 1990:1309 (EM)
O?
Frohlich & Lau 2008:9 (NEWNAT/O, DESCR); Wagner et al. 2012:72 (O
record adventive)
Touchardia latifolia Gaudich.
end
K
O Mo L
M
H
Wagner et al. 1990:1310 (K, O, Mo, L, M, H)
Urera glabra (Hook. & Arn.) Wedd.
end
K
O Mo L
M
H
Wagner et al. 1990:1312 (K, O, Mo, L, M, H)
Urera kaalae Wawra
end
O
Urtica urens L.
nat
O
Wagner et al. 1990:1313 (O)
H
Wagner et al. 1990:1314 (H); Herbarium Pacificum Staff 1996:7 (O)
Valerianaceae
Centranthus ruber (L.) DC. ssp. ruber
nat
M
Wagner et al. 1990:1315 (EM)
Verbenaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 170
Scientific Name
Dicot
Verbenaceae
Status
Citharexylum caudatum L.
nat
Citharexylum spinosum L.
Kl Le Ni K
M Ka H
M
nat
O Mo
M
Clerodendrum buchananii (Roxb.) Walp. var. fallax
(Lindl.) Bakh.
nat
O?
M
H
Wagner et al. 1990:1319 (O? [as Clerodendrum speciosissimum; garden
escape]); Staples & Herbst 2005:555, 556, 559 (KEY), 559 (NOM:Syn. C.
speciosissimum, DESCR); Starr et al. 2006:41 (NEWNAT/EM, DESCR);
Frohlich & Lau 2007:11 (O); Parker & Parsons 2010:43 (H); Wagner et al.
2012:72 (O record adventive)
Clerodendrum chinense (Osbeck) Mabb.
nat
K
O Mo L
M
H
Wagner et al. 1990:1319 (K, O, Mo, L, M, H [as Clerodendrum
philippinum]); Wagner & Herbst 1995:26 (NOM:Syn. C. philippinum);
Staples & Herbst 2005:555, 556, 559 (KEY), 560 (DESCR)
Clerodendrum glabrum E.Mey.
nat
K
Clerodendrum macrostegium Schauer
Clerodendrum myricoides (Hochst.) Vatke
Duranta erecta L.
?
nat
K
?
K?
nat
K
H
Citation
O
Clerodendrum indicum (L.) Kuntze
K
O Mo L
Wagner et al. 1990:1317 (O); Nagata 1995:13 (O); Starr et al. 1999:14 (EM);
Oppenheimer 2003:26 (H); Starr et al. 2004:28 (K); Staples & Herbst
2005:556 (KEY), 558 (DESCR)
Wagner et al. 1990:1317; Herbarium Pacificum Staff 1998:13 (NEWNAT/O,
DESCR); Starr et al. 2002:26 (EM); Staples & Herbst 2005:556 (KEY), 558
(DESCR); Starr et al. 2006:41 (Mo)
Staples & Herbst 2005:555, 556, 559 (KEY), 560 (DESCR); Frohlich & Lau
2012:48 (NEWNAT/K, DESCR)
O?
Wagner et al. 1990:1319 (O? [adventive], DESCR); Staples & Herbst
2005:555, 556, 559 (KEY), 560 (DESCR)
O
Lorence & Flynn 1997:12 (NEWNAT/K, DESCR); Staples & Herbst
2005:555, 556, 559 (KEY), 561 (DESCR); Daehler & Baker 2006:11 (O)
Staples & Herbst 2005:555, 556, 559 (KEY), 560 (DESCR); Frohlich & Lau
2012:51 (K? [adventive])
O
Lorence et al. 1995:55 (NEWNAT/K, DESCR); Herbarium Pacificum Staff
1998:13 (O); Staples & Herbst 2005:556 (KEY), 562 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 171
Scientific Name
Dicot
Verbenaceae
Status
Lantana camara L.
nat
Lantana montevidensis (Spreng.) Briq.
nat
Phyla nodiflora (L.) Greene
nat
Priva aspera Kunth
Kl Le Ni K
Le? Ni
O Mo L
K
O Mo L
K
L
M Ka H
O Mo
?
M Ka
M
Wagner et al. 1990:1316; Starr et al. 2002:26 (NEWNAT/WM); Staples &
Herbst 2005:555 (KEY), 566 (DESCR); Kennedy et al. 2010:24 (O, Mo)
O?
Wagner et al. 1990:1316 (O? [reported by Hillebrand in 1888 as a widesperad
weed on western O‘ahu, but no vouchers seen], DESCR)
nat
K
O
Stachytarpheta cayennensis (Rich.) Vahl
nat
K
O Mo
?
K?
Stachytarpheta jamaicensis (L.) Vahl
nat
K
Stachytarpheta mutabilis (Jacq.) Vahl
nat
K
Verbena bonariensis L.
nat
K
O? Mo? L
M
Verbena litoralis Kunth
nat
K
O Mo L
M Ka
Ni
Wagner et al. 1990:1320 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:556 (KEY), 564 (DESCR); Wood & LeGrande 2006:29 (Le? [reported
by Caum in 1936 but unvouchered])
Wagner et al. 1990:1320 (L); Lorence et al. 1995:55 (K); Staples & Herbst
2005:556 (KEY), 564 (DESCR)
Stachytarpheta australis Moldenke
Stachytarpheta incana Moldenke
H
Citation
L
M
H
Wagner et al. 1990:1321 (K, O, L, M, H [as Stachytarpheta dichotoma]);
Munir 1992:156/Herbst & Wagner 1999:32 (NOM: S. dichotoma misapplied)
M
H
Wagner et al. 1990:1322 (K, O, Mo, M, H [as Stachytarpheta urticifolia]);
Munir 1992:145/Herbst & Wagner 1999:32 (NOM:Syns. S. dichotoma, S.
urticifolia); Staples & Herbst 2005:555 (KEY), 569 (DESCR)
Wagner et al. 1990:1321 (K? [reported by Moldenke in 1874 as present, but
no vouchers seen])
O Mo L
M Ka
H
Wagner et al. 1990:1322 (K, O, L, M, Ka, H); Oppenheimer 2003:26 (Mo)
Wagner et al. 1990:1322 (K); Staples & Herbst 2005:555 (KEY), 569
(DESCR)
Wagner et al. 1990:1325 (Mo? [reported by Moldenke in 1962, but no
vouchers seen], L, EM); Lorence et al. 1995:55 (K); Frohlich & Lau 2008:9
(O); Wagner et al. 2012:73 (O plants controlled, not definitely naturalized)
H
Wagner et al. 1990:1325 (Ni, K, O, Mo, L, M, Ka, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 172
Scientific Name
Dicot
Verbenaceae
Status
Kl Le Ni K
O Mo L
Verbena rigida Spreng.
nat
O?
Vitex parviflora Juss.
nat
O
Vitex rotundifolia L.f.
ind
Vitex trifolia L.
nat
Ni
K
M Ka H
H
Citation
Wagner et al. 1990:1325 (H); Staples & Herbst 2005:555 (KEY), 567
(DESCR); Frohlich & Lau 2008:10 (O); Wagner et al. 2012:73 (O record
adventive)
Wagner et al. 1990:1326; Staples & Herbst 2005:555, 567 (KEY), 567
(DESCR); Frohlich & Lau 2008:10 (NEWNAT/O, DESCR)
O Mo L
M
H
O Mo
Wagner et al. 1990:1326 (Ni, K, O, Mo, L, M, H); Staples & Herbst
2005:556, 567 (KEY), 568 (DESCR)
Wagner et al. 1990:1326; Staples & Herbst 2005:555, 567 (KEY), 568
(DESCR); Wysong et al. 2007:7 (NEWNAT/Mo); Lau & Frohlich 2012:23
(O)
Violaceae
Isodendrion hosakae H.St.John
end
H
Isodendrion laurifolium A.Gray
end
K
O
Wagner et al. 1990:1329 (K, O)
Isodendrion longifolium A.Gray
end
K
O
Wagner et al. 1990:1329 (K, O)
Isodendrion pyrifolium A.Gray
end
Viola chamissoniana Ging. ssp. chamissoniana
end
Viola chamissoniana Ging. ssp. robusta (Hillebr.)
W.L.Wagner, D.R.Herbst & Sohmer
end
Viola chamissoniana Ging. ssp. tracheliifolia (Ging.)
W.L.Wagner, D.R.Herbst & Sohmer
end
K
Viola helenae C.N.Forbes & Lydgate
end
K
Ni
O Mo L
M?
O
H
Wagner et al. 1990:1328 (H)
Wagner et al. 1990:1331 (Ni, O, Mo, L, M? [reported by Hillebrand in 1888,
but no vouchers seen], H); Herbarium Pacificum Staff 1996:7
(REDISCOVER/H)
Wagner et al. 1990:1333 (O)
Mo
O Mo
Wagner et al. 1990:1333 (Mo)
M
Wagner et al. 1990:1333 (K, O, Mo, M)
Wagner et al. 1990:1333 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 173
Scientific Name
Dicot
Violaceae
Status
Kl Le Ni K
Viola kauaensis A.Gray var. kauaensis
end
K
Viola kauaensis A.Gray var. wahiawaensis C.N.Forbes
end
K
Viola lanaiensis W.Becker
end
Viola maviensis H.Mann
end
Viola oahuensis C.N.Forbes
end
Viola odorata L.
nat
K
Viola wailenalenae (Rock) Skottsb.
end
K
Korthalsella complanata (Tiegh.) Engl.
ind
K
Korthalsella cylindrica (Tiegh.) Engl.
Korthalsella degeneri Danser
O Mo L
M Ka H
O
Citation
Wagner et al. 1990:1334 (K, O)
Wagner et al. 1990:1334 (K)
L
Mo
M
M
Wagner et al. 1990:1334 (L); Havran et al. 2012 (WM)
H
O
Wagner et al. 1990:1336 (Mo, M, H)
Wagner et al. 1990:1336 (O)
L?
H
Wagner et al. 1990:1332 (K? [reported but unvouchered in 1987 along stream
and in wet forest on southwestern Kaua‘i], L? [single coll. in 1910 along
watercourse in Mahana Valley]); Shannon & Wagner 1996:15
(NEWNAT/H); Staples & Herbst 2005:570 (KEY, DESCR); Imada 2007:39
(K)
Wagner et al. 1990:1336 (K)
Viscaceae
O Mo L
M
H
Wagner et al. 1990:1338 (K, O, Mo, L, M, H); Molvray 1997:270
(NOM:Syn. Korthalsella complanata = K. taenioides fma. taenioides, status
change from "end" to "ind"); Herbst & Wagner 1999:32 (recommend against
adopting Molvray 1997 pending further evaluation)
end
O Mo L
M
H
Wagner et al. 1990:1339 (O, Mo, L, M, H); Molvray 1997:271 (status change
from "end" to "ind"); Herbst & Wagner 1999:32 (recommend against
adopting Molvray 1997 pending further evaluation)
end
O
Wagner et al. 1990:1339 (O); Molvray 1997:270, 271(COMBNOV &
STATNOV [as Korthalsella taenioides fma. horneana], NOM:Syn. K.
degeneri, status change from "end" to "ind"); Herbst & Wagner 1999:32
(recommend against adopting Molvray 1997 pending further evaluation)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 174
Scientific Name
Dicot
Viscaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Korthalsella latissima (Tiegh.) Danser
end
K
O
H
Korthalsella platycaula (Tiegh.) Engl.
ind
K
O
L
M
Korthalsella remyana Tiegh.
end
K
O Mo L
M
Citation
Wagner et al. 1990:1339 (K, O, H); Molvray 1997:271 (COMBNOV [as
Korthalsella taenioides fma. pendula], NOM:Syn. K. latissima, status change
from "end" to "ind"); Herbst & Wagner 1999:32 (recommend against
adopting Molvray 1997 pending further evaluation)
Wagner et al. 1990:1340 (K, O, L, EM); Molvray 1997:270 (NOM:Syn.
Korthalsella platycaula = K. taenioides fma. taenioides, status change from
"end" to "ind"); Herbst & Wagner 1999:32 (recommend against adopting
Molvray 1997 pending further evaluation)
H
Wagner et al. 1990:1340 (K, O, Mo, L, M, H); Molvray 1997:271
(COMBNOV & STATNOV [as Korthalsella taenioides fma. remyana],
NOM:Syn. K. remyana, status change from "end" to "ind"); Herbst & Wagner
1999:32 (recommend against adopting Molvray 1997 pending further
evaluation)
H?
Wagner et al. 1990:1342 (H? [noted escaping in 1985 in vicinity of the
DLNR Arboretum in Hilo], DESCR)
H
Wagner et al. 1990:1342 (K, O, H); Staples & Herbst 2005:571 (KEY), 572
(DESCR)
Vitaceae
Cayratia trifolia (L.) Domin
?
Cissus nodosa Blume
nat
K
O
Cissus quadrangularis L.
nat
O
Wagner et al. 1990:1342 (O? [as Cissus quadrangula; cultivated and escaping
in Koko Crater]); Staples & Herbst 2005:571 (KEY), 573 (DESCR); Frohlich
& Lau 2008:10 (NEWNAT/O)
Cissus repens Lam.
nat
O
Lau & Frohlich 2012:23 (NEWNAT/O, DESCR)
Cissus rhombifolia Vahl
nat
Cissus rotundifolia (Forssk.) Vahl
nat
M
K
O
M
Starr et al. 2004:28 (NEWNAT/WM, DESCR); Staples & Herbst 2005:571
(KEY), 572 (DESCR)
H
Lorence & Flynn 1997:12 (NEWNAT/K, DESCR); Herbst 1998:4 (O); Starr
et al. 2002:26 (EM); Staples & Herbst 2005:571 (KEY), 572 (DESCR);
Parker & Parsons 2012a:62 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 175
Scientific Name
Dicot
Vitaceae
Status
Kl Le Ni K
Cissus verticillata (L.) Nicolson & C.E.Jarvis
nat
K
Tetrastigma pubinerve Merr. & Chun
nat
K
Tetrastigma voinieranum (Baltet) Pierre ex Gagnep.
?
Vitis vinifera L.
?
O Mo L
M Ka H
O
Citation
Staples & Herbst 2005:571 (KEY), 573 (DESCR); Frohlich & Lau 2012:48
(NEWNAT/K, O, DESCR)
M
Oppenheimer & Bartlett 2000:9 (NEWNAT/WM, DESCR); Staples & Herbst
2005:573 (DESCR); Frohlich & Lau 2012:49 (K)
O?
Staples & Herbst 2005:573 (DESCR); Frohlich & Lau 2007:12 (O?
[adventive], DESCR)
K?
H?
Wagner et al. 1990:1342 [sometimes persisting after cultivation]; Staples &
Herbst 2005:574 (DESCR); Frohlich & Lau 2012:51 (K?, H? [adventive])
Zygophyllaceae
Guaiacum officinale L.
nat
Tribulus cistoides L.
ind
Tribulus terrestris L.
nat
O
Kl Le Ni
Staples & Herbst 2005:575 (DESCR); Lau & Frohlich 2012:24
(NEWNAT/O, DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:1343 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:576 (DESCR); Wood & LeGrande 2006:29 (Le)
K
O Mo
M
H
Wagner et al. 1990:1344 (K, O, M, H); Oppenheimer 2003:27 (Mo)
Gymnosperm
Araucariaceae
Araucaria columnaris (G.Forst.) Hook.
nat
Mo L
M
Oppenheimer 2002:20 (NEWNAT/WM, KEY); Staples & Herbst 2005:58
(DESCR, KEY); Oppenheimer 2007:19 (Mo); Oppenheimer 2008:23 (L)
M
Oppenheimer 2002:20 (NEWNAT/WM, KEY); Staples & Herbst 2005:60
(KEY), 62 (DESCR); Frohlich & Lau 2012:34 (O)
Cupressaceae
Callitris columellaris F.Muell.
nat
O
Callitris endlicheri (Parl.) F.M.Bailey
nat
O
Lau & Frohlich 2012:14 (NEWNAT/O, DESCR)
Cupressus lusitanica Mill.
nat
O
Staples & Herbst 2005:60, 64 (KEY), 64 (DESCR); Frohlich & Lau 2012:34
(NEWNAT/O, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 176
Scientific Name
Gymnosperm
Cupressaceae
Status
Cupressus macrocarpa Hartw.
nat
Juniperus bermudiana L.
nat
Kl Le Ni K
O Mo L
M Ka H
L
Citation
M
Oppenheimer 2002:19 (EM? [sparingly reproducing in Polipoli]); Staples &
Herbst 2005:60, 64 (KEY), 64 (DESCR); Starr et al. 2008:44
(NEWNAT/EM, DESCR)
M
Oppenheimer 2002:20, 21 (NEWNAT/WM, KEY); Staples & Herbst
2005:60, 65 (KEY), 66 (DESCR); Oppenheimer 2008:26 (L)
M
Oppenheimer 2003:19 (NEWNAT/EM, WM, KEY); Staples & Herbst
2005:70 (KEY), 71 (DESCR) [as to the species]
Pinaceae
Pinus caribaea Morelet var. hondurensis (Sénécl.)
W.H.Barrett & Golfari
nat
Pinus elliottii Engelm.
nat
Pinus jeffreyi A.Murray bis
K
O Mo
Wysong et al. 2007:6 (NEWNAT/Mo); Frohlich & Lau 2012:43 (K, O)
?
M?
Oppenheimer 2003:19 (M? [naturalizing but unvouchered], KEY)
nat
M
Oppenheimer 2002:20, 21 (NEWNAT/EM, KEY); Oppenheimer 2003:19
(KEY); Staples & Herbst 2005:70 (KEY), 71 (DESCR)
Pinus pinaster Aiton
?
M?
Oppenheimer 2002:19 (EM? [sparingly reproducing in Polipoli/Kula Forest
Reserve area]); Oppenheimer 2003:19 (KEY)
Pinus radiata D.Don
nat
M
Oppenheimer 2002:20, 21 (NEWNAT/EM, KEY); Oppenheimer 2003:19
(KEY)
Pinus taeda L.
nat
Pinus patula Schiede ex Schltdl. & Cham.
K?
L
H
Staples & Herbst 2005:70 (KEY), 71 (DESCR); Oppenheimer 2008:31
(NEWNAT/L, DESCR); Frohlich & Lau 2012:51 (K? [adventive]); Parker &
Parsons 2012b:71 (H)
Podocarpaceae
Afrocarpus mannii (Hook.f.) C.N.Page
Dacrycarpus imbricatus (Blume) de Laub.
?
O?
Daehler & Baker 2006:17 (O? [adventive] , DESCR)
nat
O
Staples & Herbst 2005:71 (KEY), 72 (DESCR); Daehler & Baker 2006:10
(NEWNAT/O, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 177
Scientific Name
Gymnosperm
Podocarpaceae
Status
Kl Le Ni K
O Mo L
M Ka H
M
Citation
Nageia falcata (Thunb.) Carrière
nat
Oppenheimer 2002:20, 22 (NEWNAT/EM, KEY); Staples & Herbst 2005:71
(KEY), 73 (DESCR)
Podocarpus elatus R.Br. ex Endl.
nat
O
Staples & Herbst 2005:71 (KEY), 74 (DESCR); Lau & Frohlich 2012:20
(NEWNAT/O, DESCR)
Podocarpus macrophyllus (Thunb.) Sweet
nat
O
Staples & Herbst 2005:71 (KEY), 74 (DESCR); Lau & Frohlich 2012:20
(NEWNAT/O, DESCR)
Taxodiaceae
Cryptomeria japonica (Thunb. ex L.f.) D.Don
nat
M
Oppenheimer 2002:20, 22 (NEWNAT/WM, KEY); Staples & Herbst
2005:75 (DESCR, KEY)
Monocot
Agavaceae
Agave sisalana Perrine
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:1348 (K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:578, 580 (KEY), 581 (DESCR)
Cordyline fruticosa (L.) A.Chev.
pol
K
O Mo L
M
H
Wagner et al. 1990:1348 (Ni, K, O, Mo, L, M, H); Staples & Herbst
2005:578 (KEY), 581 (DESCR)
Furcraea foetida (L.) Haw.
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:1350 (K, O, Mo, L, M, H); Staples & Herbst 2005:578
(KEY), 586 (DESCR); Starr & Starr 2011:23 (Ka)
Phormium tenax J.R.Forst. & G.Forst.
nat
K
Mo
Pleomele aurea (H.Mann) N.E.Br.
end
K
Ni
Wagner et al. 1990:1350 (K, Mo); Staples & Herbst 2005:578 (KEY), 589
(DESCR)
Wagner et al. 1990:1352 (K); Jankalski 2008:20 (proposed transfer to
Dracaena aurea), 21 (proposed NOMNOV Dracaena halemanuensis (K) to
replace Pleomele stenophylla [St. John 1987i:55, SPNOV/K, DESCR], as the
binomial Dracaena stenophylla is already published); Wagner et al. 2005–
(NOM:Syn. P. stenophylla)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 178
Scientific Name
Monocot
Agavaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Mo
M
Citation
Pleomele auwahiensis H.St.John
end
Wagner et al. 1990:1352 (Mo, M); Jankalski 2008:21 (proposed syn. under
COMBNOV Dracaena rockii [Pleomele rockii listed as syn. of P. auwahiensis
in Wagner et al. 1990:1352])
Pleomele fernaldii H.St.John
end
Pleomele forbesii O.Deg.
end
O
Wagner et al. 1990:1352 (O); Jankalski 2008:21 (proposed transfer to
COMBNOV Dracaena forbesii)
Pleomele halapepe H.St.John
end
O
Wagner et al. 1990:1352 (O); Jankalski 2008:21 (proposed transfer to
COMBNOV Dracaena halaapepe [note spelling change])
Pleomele hawaiiensis O.Deg. & I.Deg.
end
Sansevieria trifasciata Prain
nat
L
Wagner et al. 1990:1352 (L); Jankalski 2008:20 (proposed transfer to
COMBNOV Dracaena fernaldii)
H
K
O
M
Wagner et al. 1990:1354 (H); Jankalski 2008:21 (proposed syn. under
COMBNOV Dracaena konaensis [Pleomele konaensis listed as syn. of P.
hawaiiensis in Wagner et al. 1990:1354])
Wagner et al. 1990:1347; Imada et al. 2000:9 (NEWNAT/O); Flynn &
Lorence 2002:14 (K); Starr et al. 2003:23 (EM); Staples & Herbst 2005:578
(KEY), 588 (DESCR)
Alismataceae
Echinodorus berteroi (Speng.) Fassett
Sagittaria latifolia Willd.
?
O?
Wagner et al. 1990:1354 (O? [as Echinodorus rostratus, now a syn. of E.
berteroi; single coll. in 1939 as a pond escape in Kahuku, not vouchered
since], DESCR)
nat
K
O
M
nat
K
O Mo L
M
H
Wagner et al. 1990:1355 (K, O, M); Imada et al. 2000:9 (H); Staples &
Herbst 2005:591 (DESCR)
Aloeaceae
Aloe vera (L.) Burm.f.
Lorence et al. 1995:21 (NEWNAT/K, DESCR); Herbarium Pacificum Staff
1999:3 (O); Oppenheimer 2003:4 (WM); Staples & Herbst 2005:592
(DESCR); Wysong et al. 2007:1 (Mo); Oppenheimer 2011:5 (L)
Araceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 179
Scientific Name
Monocot
Araceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Aglaonema commutatum Schott
nat
O
Alocasia cucullata (Lour.) G.Don
nat
O Mo
M
H
Staples et al. 2003:8 (NEWNAT/EM, H, DESCR); Staples & Herbst
2005:595, 598 (KEY), 598 (DESCR); Oppenheimer 2007:18 (Mo); Imada
2007:35 (O)
Alocasia macrorrhizos (L.) G.Don
pol
O Mo
M
H
Wagner et al. 1990:1356 (K, O, Mo, M); Staples & Woolliams 1997:15
(KEY); Imada et al. 2000:10 (H); Staples & Herbst 2005:595, 598 (KEY),
599 (DESCR)
Anthurium pentaphyllum (Aubl.) G.Don var.
bombacifolium (Schott) Madison
K
?
Staples & Herbst 2005:594, 595 (KEY), 597 (DESCR); Imada 2007:34
(NEWNAT/O, DESCR)
O?
Caladium bicolor (Aiton) Vent.
nat
Colocasia esculenta (L.) Schott
pol
Dieffenbachia maculata (Loddiges) G.Don
nat
K
Epipremnum pinnatum (L.) Engl.
nat
K
Philodendron erubescens K.Koch & Augustin
nat
K
Philodendron lacerum (Jacq.) Schott
nat
Philodendron pinnatifidum (Jacq.) Schott
nat
Ni
K
Daehler & Baker 2006:13 (O? [adventive], DESCR)
O Mo L
M
H
Staples & Herbst 2005:594 (KEY), 601 (DESCR); Resslar 2010:44
(NEWNAT/H, DESCR)
H
Wagner et al. 1990:1357 (Ni, K, O, Mo, L, M, H); Staples & Herbst
2005:594 (KEY), 602 (DESCR)
M
O Mo
O?
K
M
Staples & Herbst 2005:594, 603 (KEY), 604 (DESCR); Lorence & Flynn
2006:2 (NEWNAT/K); Oppenheimer 2007:18 (WM)
H
Wagner et al. 1990:1359 (K, O, M, H); Staples & Herbst 2005:594, 595
(KEY), 604 (DESCR); Oppenheimer 2007:18 (Mo)
M
Flynn & Lorence 2002:14 (NEWNAT/K, DESCR); Staples & Herbst
2005:594, 608 (KEY), 608 (DESCR); Oppenheimer 2007:19 (EM)
M
Oppenheimer 2004:9 (NEWNAT/EM); Staples & Herbst 2005:594, 607
(KEY), 610 (DESCR); Daehler & Baker 2006:13 (O? [adventive], DESCR)
Flynn & Lorence 2002:15 (NEWNAT/K); Staples & Herbst 2005:594, 607
(KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 180
Scientific Name
Monocot
Araceae
Philodendron scandens K.Koch & Sello
Philodendron scandens K.Koch & Sello ssp. scandens f.
micans (K.Koch) G.S.Bunting
Status
Kl Le Ni K
nat
O Mo L
M Ka H
K
?
M
Staples & Herbst 2005:594, 608 (KEY), 610 (DESCR); Oppenheimer
2008:23 (NEWNAT/K, EM, DESCR)
O?
Pistia stratiotes L.
nat
K
Syngonium podophyllum Schott
nat
Xanthosoma robustum Schott
nat
K
Zantedeschia aethiopica (L.) Spreng.
nat
K
Citation
Staples & Herbst 2005:594, 608 (KEY); Daehler & Baker 2006:13 (O?
[adventive], DESCR)
O Mo
M
Wagner et al. 1990:1359 (K, O, Mo, M); Staples & Herbst 2005:594 (KEY),
610 (DESCR)
O
M
Staples & Herbst 2005:594 (KEY), 612 (DESCR); Oppenheimer 2006:10
(NEWNAT/WM); Oppenheimer 2011:6 (O)
O
M
H
M
Staples & Woolliams 1997:13 (NEWNAT/K [as Xanthosoma roseum],
KEY); Imada et al. 2000:10 (H); Oppenheimer & Bartlett 2000:2 (EM, WM);
Herbst et al. 2004:3 (NOM:Syn. X. roseum); Staples & Herbst 2005:595
(KEY), 613 (DESCR)
Wagner et al. 1990:1360 (K, M)
Arecaceae
Archontophoenix alexandrae (F.Muell.) H.Wendl. &
Drude
nat
O
Arenga pinnata (Wurmb) Merr.
nat
O
Caryota mitis Lour.
nat
O
Staples & Herbst 2005:616 (KEY), 619 (DESCR); Daehler & Baker 2006:5
(NEWNAT/O, DESCR)
Coccothrinax barbadensis (Lodd. ex Mart.) Becc.
nat
K
O
Staples & Herbst 2005:617 (KEY), 620 (DESCR); Frohlich & Lau 2012:29
(NEWNAT/K, O, DESCR)
Cocos nucifera L.
pol
K
O Mo L
Ni
M
M Ka
H
Wagner et al. 1990:1362 (H); Staples & Herbst 2005:616, 618 (KEY), 618
(DESCR); Daehler & Baker 2006:4 (O, DESCR); Oppenheimer 2011:6 (EM)
H
Daehler & Baker 2006:5 (NEWNAT/O, DESCR); Parker & Parsons
2012b:66 (H)
H
Wagner et al. 1990:1362 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst
2005:617 (KEY), 620 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 181
Scientific Name
Monocot
Arecaceae
Status
Licuala spinosa Wurmb
nat
Livistona chinensis (Jacq.) R.Br. ex Mart.
nat
Phoenix hybrid
Kl Le Ni K
O Mo L
M Ka H
O
K
Citation
Lau & Frohlich 2012:8 (NEWNAT/O, DESCR)
O
M
Wagner et al. 1990:1364 (O); Oppenheimer 2003:5 (WM); Staples & Herbst
2005:617, 623 (KEY), 623 (DESCR); Daehler & Baker 2006:12 (O);
Oppenheimer 2011:6 (K)
nat
O? Mo?
M
Wagner et al. 1990:1361 (O? [possibly reproducing along Kamehameha Hwy
between Hale‘iwa and Waimea, and near Käne‘ohe]); Oppenheimer 2003:6
(NEWNAT/EM, Mo? [observed naturalized but unvouchered], DESCR);
Staples & Herbst 2005:616, 624 (KEY), 625 (DESCR)
Pinanga coronata (Blume ex Mart.) Blume
nat
O
Pritchardia arecina Becc.
end
Pritchardia bakeri Hodel
end
Pritchardia beccariana Rock
end
Pritchardia flynnii Lorence & Gemmill
end
Pritchardia forbesiana Rock
end
Pritchardia glabrata Becc. & Rock
end
Pritchardia gordonii Hodel
end
Pritchardia hardyi Rock
end
H
M
Staples & Herbst 2005:617 (KEY); Daehler & Baker 2006:5 (NEWNAT/O,
DESCR); Parker & Parsons 2012b:66 (H)
Wagner et al. 1990:1367 (EM); Hodel 2007:S-10 (KEY, DESCR)
O
Hodel 2009:173 (SPNOV/O, DESCR)
H
K
Wagner et al. 1990:1369 (H); Staples & Herbst 2005:617, 625 (KEY); Hodel
2007:S-11 (KEY, DESCR)
Lorence & Gemmill 2004:185 (SPNOV/K, DESCR); Hodel 2007:S-12
(KEY, DESCR)
Mo
L
M
Wagner et al. 1990:1369 (WM); Hodel 2007:S-13 (Mo); Hodel 2007:S-13
(KEY, DESCR)
M
Wagner et al. 1990:1365 (Pritchardia lanaiensis, P. elliptica unplaced, ID
doubtful), 1369 (WM [as P. glabrata]); Herbst & Wagner 1999:13 (P.
lanaiensis, RESURRECT/L); Hodel 2007:S-14 (L, NOM:Syn. P. elliptica, P.
lanaiensis, KEY, DESCR)
H
K
Hodel 2007:S-15 (SPNOV/H, KEY, DESCR)
Wagner et al. 1990:1370 (K); Hodel 2007:S-18 (KEY, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 182
Scientific Name
Monocot
Arecaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Mo
Citation
Pritchardia hillebrandii Becc.
end
Wagner et al. 1990:1370 (Mo); Staples & Herbst 2005:617, 625 (KEY), 626
(DESCR); Hodel 2007:S-18 (KEY, DESCR)/Wagner et al. 2012:77 (author
correction [from "(Kuntze) Becc."])
Pritchardia kaalae Rock
end
Pritchardia lanigera Becc.
end
Pritchardia lowreyana Rock
end
Pritchardia maideniana Becc.
end
Pritchardia martii (Gaudich.) H.Wendl.
end
Pritchardia minor Becc.
end
Pritchardia munroi Rock
end
Pritchardia napaliensis H.St.John
end
K
St. John 1988a:177 (SPNOV/K [as Pritchardia limahuliensis], DESCR);
Wagner et al. 1990:1373 (K [as P. napaliensis]); Herbst & Wagner 1999:13
(accept P. limahuliensis as a new species); Hodel 2007:S-37 (NOM:Syn. P.
limahuliensis, KEY, DESCR)
Pritchardia perlmanii Gemmill
end
K
Gemmill 1998:18 (SPNOV/K, DESCR); Herbst & Wagner 1999:13
(DESCR); Hodel 2007:S-38 (KEY, DESCR)
Pritchardia remota Becc.
end
O
Wagner et al. 1990:1370 (O); Hodel 2007:S-20 (KEY, DESCR)
H
O Mo
Wagner et al. 1990:1371 (H); Hodel 2007:S-22 (KEY, DESCR)
Wagner et al. 1990:1371 (Mo); Hodel 2007:S-23 (KEY, DESCR); Hodel
2009:177 (O)
H
O
Wagner et al. 1990:1367 (H [as Pritchardia affinis]); Staples & Herbst
2005:617, 625 (KEY); Hodel 2007:S-26 (NOM:Syn. P. affinis, KEY,
DESCR)
Wagner et al. 1990:1372 (O [incl. syn. Pritchardia kahukuensis]); Hodel
2007:S-20 (RESURRECT/O [as P. kahukuensis], KEY, DESCR), S-27 (as P.
martii, KEY, DESCR); Wagner et al. 2012:77 (P. kahukuensis not recognized)
K
Wagner et al. 1990:1372 (K); Hodel 2007:S-33 (KEY, DESCR)
Mo
Ni
M
Wagner et al. 1990:1373 (Mo); Hodel 2007:S-36 (WM, KEY, DESCR)
Wagner et al. 1990:1367 [Ni [as Pritchardia aylmer-robinsonii]), 1373 (N [as
P. remota]); Hodel 2007:S-39 (Ni, NOM:Syn. P. aylmer-robinsonii, KEY,
DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 183
Scientific Name
Monocot
Arecaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Pritchardia schattaueri Hodel
end
Pritchardia viscosa Rock
end
K
Wagner et al. 1990:1374 (K); Hodel 2007:S-43 (KEY, DESCR)
Pritchardia waialealeana Read
end
K
Wagner et al. 1990:1374 (K); Hodel 2007:S-43 (KEY, DESCR)
Pritchardia woodii Hodel
end
Ptychosperma macarthurii (Veitch) H.Wendl. ex Hook.f.
nat
O
Staples & Herbst 2005:617 (KEY), 626 (DESCR); Daehler & Baker 2006:6
(NEWNAT/O, DESCR)
?
O?
Wagner et al. 1990:1361 (O? [possibly sparingly naturalized in Kalihi and
Nu‘uanu Valleys, and in Käne‘ohe])
Roystonea regia (Kunth) O.F.Cook
nat
O
Wagner et al. 1990:1361 (O? [possibly sparingly naturalized in Kalihi and
Nu‘uanu Valleys, and in Käne‘ohe]); Staples & Herbst 2005:616 (KEY), 628
(DESCR); Frohlich & Lau 2007:9 (NEWNAT/O)
Syagrus romanzoffiana (Cham.) Glassman
nat
O
Staples & Herbst 2005:617 (KEY), 628 (DESCR); Daehler & Baker 2006:6
(NEWNAT/O, DESCR)
Washingtonia filifera (Linden ex André) H.Wendl. ex
de Bary
nat
O
M
Oppenheimer & Bartlett 2002:3 (NEWNAT/WM, KEY); Staples & Herbst
2005:617 (KEY), 629 (DESCR); Frohlich & Lau 2010:5 (O)
Washingtonia robusta H.Wendl.
nat
O
M
Oppenheimer & Bartlett 2002:3 (NEWNAT/WM, KEY); Staples & Herbst
2005:617 (KEY), 630 (DESCR); Frohlich & Lau 2010:5 (O)
Aechmea bracteata (Sw.) Griseb.
?
O?
Staples & Herbst 2005:631, 634 (KEY); Frohlich & Lau 2008:4
(NEWNAT/O, DESCR); Wagner et al. 2012:77 (O record adventive)
Guzmania lindenii (André) Mez
nat
O
Frohlich & Lau 2010:6 (NEWNAT/O, DESCR)
Roystonea elata (W.Bartram) F.Harper
H
Citation
M
Wagner et al. 1990:1374 (H); Hodel 2007:S-41 (KEY, DESCR)
Hodel 2007:S-44 (SPNOV/EM, KEY, DESCR); Hodel 2007:S-44 (KEY,
DESCR)
Bromeliaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 184
Scientific Name
Monocot
Bromeliaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Guzmania monostachia (L.) Rusby ex Mez
nat
O
Staples & Herbst 2005:631, 638 (KEY); Frohlich & Lau 2010:7
(NEWNAT/O, DESCR)
Tillandsia balbisiana Schult.f.
nat
O
Frohlich & Lau 2010:7 (NEWNAT/O, DESCR)
Tillandsia bulbosa Hook.
nat
O
Staples & Herbst 2005:631, 641 (KEY); Frohlich & Lau 2010:7
(NEWNAT/O, DESCR)
Tillandsia caput-medusae E.Morren
nat
Tillandsia fasciculata Sw.
nat
O
Staples & Herbst 2005:631, 642 (KEY); Frohlich & Lau 2010:8
(NEWNAT/O, DESCR)
Tillandsia gardneri Lindl.
nat
O
Staples & Herbst 2005:631, 642 (KEY); Frohlich & Lau 2010:8
(NEWNAT/O, DESCR)
?
O?
Staples & Herbst 2005:631, 642 (KEY); Frohlich & Lau 2010:8
(NEWNAT/O, DESCR); Wagner et al. 2012:78 (O record adventive)
Tillandsia polystachia (L.) L.
nat
O
Staples & Herbst 2005:631, 642 (KEY); Frohlich & Lau 2010:9
(NEWNAT/O, DESCR)
Tillandsia recurvata (L.) L.
nat
O
Staples & Herbst 2005:631, 641 (KEY); Frohlich & Lau 2010:9
(NEWNAT/O, DESCR)
Tillandsia stricta Sol.
nat
O
Staples & Herbst 2005:631, 642 (KEY); Frohlich & Lau 2010:10
(NEWNAT/O, DESCR)
Tillandsia usneoides (L.) L.
?
O?
Staples & Herbst 2005:631, 641 (KEY), 643 (DESCR); Frohlich & Lau
2007:9 (NEWNAT/O); Wagner et al. 2012:78 (O record adventive)
Werauhia gladioliflora (H.Wendl.) J.R.Grant
?
O?
Frohlich & Lau 2012:50 (O? [adventive])
nat
O
Lau & Frohlich 2012:11 (NEWNAT/O, DESCR)
Tillandsia juncea (Ruiz & Pav.) Poir.
Werauhia sanguinolenta (Cogn. & Marchal) J.R.Grant
M
Staples & Herbst 2005:631, 641 (KEY); Oppenheimer 2006:11
(NEWNAT/EM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 185
Scientific Name
Monocot
Cannaceae
Canna indica L.
Status
Kl Le Ni K
O Mo L
M Ka H
H
Citation
nat
K
O Mo L
M
Wagner et al. 1990:1375 (K, O, Mo, L, M, H); Staples & Herbst 2005:645
(DESCR)
Callisia fragrans (Lindl.) Woodson
nat
K
O
M
Callisia repens L.
nat
Commelina benghalensis L.
nat
K
O Mo
M
H
Wagner et al. 1990:1379 (K, O, M, H); Oppenheimer 2003:9 (Mo)
Commelina diffusa Burm.f.
nat
K
O Mo L
M
H
Wagner et al. 1990:1379 (K, O, Mo, L, M, H)
Commelinaceae
Wagner et al. 1990:1378 (K, O); Starr et al. 2003:26 (EM); Staples & Herbst
2005:646 (KEY), 647 (DESCR)
O
Staples & Herbst 2005:646 (KEY), 647 (DESCR); Frohlich & Lau 2010:10
(NEWNAT/O, DESCR)
Commelina erecta L.
?
O?
Wagner et al. 1990:1379 (O? [reportedly sparingly naturalized in 1925 in
upper Mänoa Valley; no vouchers seen], DESCR)
Gibasis geniculata (Jacq.) Rohweder
?
O?
Wagner et al. 1990:1376 (O? [as Tradescantia geniculata; last reported from
Pauoa Valley in1919, not vouchered since]) [Note: The Plant List website
(2012) now lists T. geniculata as a synonym]
Murdannia nudiflora (L.) Brenan
nat
Palisota bracteosa C.B.Clarke
nat
Tradescantia fluminensis Vell.
nat
Tradescantia spathacea Sw.
nat
H
O
Herbst et al. 2004:5 (NEWNAT/O, DESCR)
M
K
Wagner et al. 1990:1380 (H)
O
H
Staples & Herbst 2005:646, 648 (KEY), 650 (DESCR); Staples et al. 2006:6
(NEWNAT/EM, H, DESCR)
Wagner et al. 1990:1376 [occasional escape]; Staples & Herbst 2005:646,
648 (KEY), 649 (DESCR); Frohlich & Lau 2010:10 (NEWNAT/O, DESCR);
Frohlich & Lau 2012:33 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 186
Scientific Name
Monocot
Commelinaceae
Tradescantia zebrina Hort. ex Bosse
Status
Kl Le Ni K
O Mo L
M Ka H
Mo
Citation
nat
K
M
Wagner et al. 1990:1376 [occasional escape]; Lorence & Flynn 1997:10
(NEWNAT/K, DESCR); Oppenheimer & Bartlett 2000:4 (WM); Starr et al.
2004:21 (EM); Staples & Herbst 2005:646, 648 (KEY), 649 (DESCR);
Oppenheimer 2007:22 (Mo)
Cheilocostus speciosus (J.König) C.Specht
nat
K
M
Wagner et al. 1990:1381 (L [as Costus speciosus]); Wagner & Herbst
1995:18 (K); Oppenheimer & Bartlett 2000:4 (EM, WM); Staples & Herbst
2005:651 (KEY), 652 (DESCR); Frohlich & Lau 2008:5 (O); Specht &
Stevenson 2006:159/Wagner et al. 2012:79 (COMBNOV:Syn. C. speciosus)
Costus dubius (Afzel.) K.Schum.
nat
O
Daehler & Baker 2006:6 (NEWNAT/O, DESCR)
Costus pulverulentus C.Presl
nat
O
Wagner et al. 1990:1381 (O? [an escape in Lyon Arboretum, Mänoa Valley],
DESCR); Staples & Herbst 2005:651 (KEY), 652 (DESCR); Daehler &
Baker 2006:6 (NEWNAT/O, DESCR)
Costus scaber Ruiz & Pav.
nat
O?
Staples & Herbst 2005:651 (KEY), 653 (DESCR); Daehler & Baker 2006:14
(O? [adventive], DESCR); Frohlich & Lau 2012:33 (NEWNAT/K, DESCR)
Costus woodsonii Maas
nat
O
?
O?
Costaceae
Dimerocostus strobilaceus Kuntze
K
O
L
M
Wagner et al. 1990:1381 (O? [an escape in Lyon Arboretum, Mänoa Valley],
DESCR); Oppenheimer 2004:11 (NEWNAT/EM); Staples & Herbst
2005:651 (KEY), 652 (DESCR); Daehler & Baker 2006:7 (O, DESCR)
Staples & Herbst 2005:651 (KEY), 653 (DESCR); Daehler & Baker 2006:14
(O? [adventive])
Cyperaceae
Bolboschoenus maritimus (L.) Palla ssp. paludosus
(A.Nelson) T.Koyama
ind
Bulbostylis capillaris (L.) C.B.Clarke
nat
Carex alligata Boott
end
Ni
K
K
O Mo
O Mo
M
M
H
Wagner et al. 1990:1384 (Ni, K, O, Mo, M); Staples et al. 2002:9 (H)
H
Wagner et al. 1990:1386 (H)
H
Wagner et al. 1990:1388 (K, O, Mo, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 187
Scientific Name
Monocot
Cyperaceae
Status
Kl Le Ni K
O Mo L
M Ka H
K
M
Citation
Carex echinata Murray
ind
H
Wagner et al. 1990:1388 (K, M, H)
Carex inversa R.Br.
nat
H
Shaw & Douglas 1995:249/Herbst & Wagner 1999:19 (NEWNAT/H)
Carex kauaiensis R.W.Krauss
end
Carex longii Mack.
nat
Carex macloviana d'Urv. ssp. subfusca (W.Boott)
T.Koyama
ind
Carex meyenii Nees
ind
Carex montis-eeka Hillebr.
end
Carex nealae R.W.Krauss
end
Carex wahuensis C.A.Mey. ssp. herbstii T.Koyama
end
Carex wahuensis C.A.Mey. ssp. rubiginosa
(R.W.Krauss) T.Koyama
end
K
O Mo L
M
H
Wagner et al. 1990:1392 (K, O, L, M, H); Kennedy et al. 2010:21 (Mo)
Carex wahuensis C.A.Mey. ssp. wahuensis
end
K
O Mo L
M
H
Wagner et al. 1990:1392 (K, O, Mo, L, M, H)
Cladium jamaicense Crantz
ind
K
O Mo
M
H
Wagner et al. 1990:1393 (K, O, Mo, M, H)
Cyperus compressus L.
nat
K
Mo
M
H
Wagner et al. 1990:1395 (H); Wagner & Herbst 1995:18 (EM); Oppenheimer
2006:12 (Mo); Wood 2006:17 (K, Mo)
K
Wagner et al. 1990:1388 (K)
Mo
Ni
K
O Mo L
K
Mo
M
H
Wagner et al. 1990:1387 (H? [as Carex ovalis; single coll. in 1956 from
Kahua Ranch, Kohala]); Strong & Wagner 1997:37 (NEWNAT/EM, H [as C.
longii], DESCR, KEY); Wagner & Herbst 2003:51 (NOM: C. ovalis
misapplied); Oppenheimer 2010:35 (Mo)
M
H
Wagner et al. 1990:1389 (M, H)
M Ka
H
Wagner et al. 1990:1389 (Ni, K, O, Mo, L, M, Ka, H)
M
M
Wagner et al. 1990:1390 (K, Mo, M)
H
O
Wagner et al. 1990:1390 (M, H [as Carex thunbergii, syn. C. nealae]);
Dragon & Barrington 2009:1898, 1906/Wagner et al. 2012:79 (NOM:C.
thunbergii misapplied; status change from "ind?" to "end")
Wagner et al. 1990:1392 (O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 188
Scientific Name
Monocot
Cyperaceae
Status
Kl Le Ni K
O Mo L
Cyperus confertus Sw.
nat
Cyperus congestus Vahl
nat
Cyperus cyperinus (Retz.) Suringar
ind
Cyperus cyperoides (L.) Kuntze
nat
Cyperus difformis L.
nat
Cyperus eragrostis Lam.
nat
Cyperus esculentus L.
nat
Cyperus fauriei Kük.
end
Cyperus gracilis R.Br.
nat
K
O Mo L
Cyperus haspan L.
nat
K
Cyperus hillebrandii Boeck. var. decipiens (Hillebr.)
Kük.
end
K
M Ka H
H
O
K
Herbarium Pacificum Staff 1996:3 (NEWNAT/H); Strong & Wagner 1997:39
(DESCR)
Wagner et al. 1990:1416 (O [as Mariscus congestus]); Tucker
1994:87/Wagner & Herbst 1999:1899 (NOM:Syn. M. congestus)
O Mo
M
Wagner et al. 1990:1416 (K, O, Mo [as Mariscus cyperinus]); Tucker
1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M. cyperinus); Starr et al.
2006:34 (EM)
H
K
Citation
O
M
Herbarium Pacificum Staff 1996:4 (NEWNAT/H); Strong & Wagner 1997:40
(DESCR, KEY)
Wagner et al. 1990:1395 (K, O); Starr et al. 2002:19 (WM); Starr et al.
2006:35 (EM)
H
Staples et al. 2003:10 (NEWNAT/H, DESCR)
H
Wagner et al. 1990:1396 (H); Strong & Wagner 1997:41 (H); Wood 2006:17
(K)
H
Wagner et al. 1990:1417 (Mo, L, H [as Mariscus fauriei]); Tucker
1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M. fauriei)
M Ka
H
Wagner et al. 1990:1396 (K, O, M); Hughes 1995:3 (Mo); Oppenheimer &
Bartlett 2002:7 (L); Starr et al. 2003:27 (H); Staples & Herbst 2005:656
(KEY, DESCR); Starr & Starr 2011:25 (Ka)
O Mo L
M
H
Wagner et al. 1990:1396 (K, M, H [as Cyperus halpan]); Tucker
1994:50/Hughes 1995:4 (Mo, comment on "halpan" spelling); Wood 2006:17
(O); Oppenheimer 2008:27 (L)
O
M
K
Mo L
L
Wagner et al. 1990:1418 (K, O, L, WM [as Mariscus hillebrandii subsp.
decipiens]); Tucker 1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M.
hillebrandii subsp. decipiens)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 189
Scientific Name
Monocot
Cyperaceae
Status
Kl Le Ni K
O Mo L
O
M Ka H
Cyperus hillebrandii Boeck. var. hillebrandii
end
Cyperus hypochlorus Hillebr. var. brevior Kük.
end
K
O Mo
M
Cyperus hypochlorus Hillebr. var. hypochlorus
end
K
O Mo
M
H
Wagner et al. 1990:1418 (K, O, Mo, M, H [as Mariscus hypochlorus subsp.
hypochlorus]); Tucker 1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M.
hypochlorus subsp. hypochlorus)
Cyperus involucratus Rottb.
nat
K
O Mo L
M
H
Wagner et al. 1990:1395 (K, O, M [as Cyperus alternifolius subsp.
flabelliformis]); Tucker 1994:20/Strong & Wagner 1997:41 (NOM:Syn. C.
alternifolius subsp. flabelliformis); Imada et al. 2000:11 (H); Staples &
Herbst 2005:656 (KEY, DESCR); Oppenheimer 2007:23 (Mo); Imada et al.
2008:12 (L)
Cyperus javanicus Houtt.
ind
Le Ni
K
O Mo L
M
H
Wagner et al. 1990:1420 (Ni, K. O, Mo, L, M, H [as Mariscus javanicus]);
Tucker 1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M. javanicus);
Wood & LeGrande 2006:23 (Le)
Cyperus laevigatus L.
ind
Ni
O Mo
M
H
Wagner et al. 1990:1398 (Ni, O, Mo, M, H); Wagner et al. 1990:1398
Cyperus meyenianus Kunth
nat
O Mo
M
H
Wagner et al. 1990:1420 (K, O [as Mariscus meyenianus]); Hughes 1995:4
(Mo); Tucker 1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M.
meyenianus); Oppenheimer 2003:10 (H); Oppenheimer 2006:12 (WM)
Cyperus neokunthianus Kük.
end
Cyperus odoratus L.
ind
K
L
M
H
Citation
Wagner et al. 1990:1418 (K, O, Mo, M [as Mariscus hypochlorus subsp.
brevior]); Tucker 1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M.
hypochlorus subsp. brevior)
M
K
O Mo
M
Wagner et al. 1990:1418 (O, L, EM, H [as Mariscus hillebrandii subsp.
hillebrandii]); Tucker 1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M.
hillebrandii subsp. hillebrandii)
Wagner et al. 1990:1420 (WM [as Mariscus kunthianus]); Tucker
1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M. kunthianus)
H
Wagner et al. 1990:1434 (K, O, Mo, M, H [as Torulinium odoratum subsp.
auriculatum]); Tucker 1994:180/Herbst & Wagner 1999:20/Wagner & Herbst
1999:1901 (NOM:Syn. T. odoratum subsp. auriculatum)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 190
Scientific Name
Monocot
Cyperaceae
Status
Cyperus papyrus L.
nat
Cyperus pennatiformis Kük. var. bryanii Kük.
end
Cyperus pennatiformis Kük. var. pennatiformis
end
Cyperus phleoides (Nees ex Kunth) Hillebr. var.
hawaiiensis (H.Mann) Kük.
end
Cyperus phleoides (Nees ex Kunth) Hillebr. var.
phleoides
Kl Le Ni K
O Mo L
M Ka H
K
H
Citation
Wagner et al. 1990:1398 (K); Staples et al. 2003:10 (H); Staples & Herbst
2005:656 (KEY, DESCR)
Only in Northwestern Hawaiian Islands; see NWHI checklist
K
O
M
K
O Mo L
M
end
K
Mo L
Cyperus pilosus Vahl
nat
K
Cyperus polystachyos Rottb.
ind
Cyperus prolifer Lam.
nat
K
Strong & Wagner 1997:41 (NEWNAT/K, KEY, DESCR); Staples & Herbst
2005:656 (KEY), 657 (DESCR)
Cyperus rockii Kük.
end
K
Wagner et al. 1990:1423 (K [as Mariscus rockii]); Tucker 1994:87/Wagner &
Herbst 1999:1901 (NOM:Syn. M. rockii)
Cyperus rotundus L.
nat
Cyperus sandwicensis Kük.
end
Ni
Le Ni
Ni
K
M Ka
H
Wagner et al. 1990:1421 (K, O, M, H [as Mariscus pennatiformis subsp.
pennatiformis]); Tucker 1994:87/Wagner & Herbst 1999:1900 (NOM:Syn.
M. pennatiformis subsp. pennatiformis)
Wagner et al. 1990:1423 (Ni, K, O, Mo, L, M [as Mariscus phleoides subsp.
hawaiiensis]); Tucker 1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M.
phleoides subsp. hawaiiensis)
H
Wagner et al. 1990:1423 (K, Mo, L, M, H [as Mariscus phleoides subsp.
phleoides]); Wagner & Herbst 1995:18 (Ka); Tucker 1994:87/Wagner &
Herbst 1999:1900 (NOM:Syn. M. phleoides subsp. phleoides)
Wagner et al. 1990:1399 (K)
O Mo L
M
K
O Mo L
M
K
O Mo
M
H
H
Wagner et al. 1990:1425 (Ni, O, Mo, L, M, H [as Pycreus polystachyos
subsp. holosericeus], K, O, Mo, M, H [as subsp. polystachyos]); Tucker
1994:72/Herbst & Wagner 1999:20/Wagner & Herbst 1999:1901 (NOM:Syn.
P. polystachyos subspp. holocericeus & polystachyos); Wood & LeGrande
2006:23 (Le)
Wagner et al. 1990:1399 (Ni, K, O, L, M, H); Hughes 1995:4 (Mo)
Wagner et al. 1990:1424 (K, O, Mo, M [as Mariscus sandwicensis]); Tucker
1994:87/Wagner & Herbst 1999:1901 (NOM:Syn. M. sandwicensis)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 191
Scientific Name
Monocot
Cyperaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
H
Citation
Cyperus sanguinolentus Vahl
nat
Cyperus sphacelatus Rottb.
nat
Cyperus trachysanthos Hook. & Arn.
end
Cyperus trinervis R.Br.
nat
H
Wagner et al. 1990:1400 (H)
Cyperus virens Michx.
nat
H
Wagner et al. 1990:1400 (H)
Eleocharis calva Torr.
ind?
Eleocharis geniculata (L.) Roem. & Schult.
nat
K
O Mo L
M
Eleocharis obtusa (Willd.) Schult.
ind
K
O Mo
M
Eleocharis olivacea Torr.
nat
Eleocharis radicans (Poir.) Kunth
nat
K
Eleocharis schaffneri Boeck.
nat
K
Fimbristylis aestivalis (Retz.) Vahl
nat
K
Fimbristylis cymosa R.Br. ssp. spathacea (Roth)
T.Koyama
ind
Ni
K
O
Fimbristylis cymosa R.Br. ssp. umbellato-capitata
(Hillebr.) T.Koyama
ind
Le Ni
K
Fimbristylis dichotoma (L.) Vahl
ind
K
Mo
Ni
K
Ni
Oppenheimer 2008:27 (NEWNAT/Mo)
O Mo L
Wagner et al. 1990:1399 (Ni, K, O, Mo, L)
O
Ka
O
Wagner et al. 1990:1427 (H [as Pycreus sanguinolentus]); Tucker
1994:66/Herbst & Wagner 1999:20/Wagner & Herbst 1999:1901 (NOM:Syn.
P. sanguinolentus); Frohlich & Lau 2008:5 (O)
M
Wagner et al. 1990:1402 (Ni, O, Ka)
Wagner et al. 1990:1402 (K, O, Mo); Oppenheimer 2003:10 (WM); Imada et
al. 2008:12 (L)
H
Wagner et al. 1990:1403 (K, O, Mo, M, H)
H
Strong & Wagner 1997:43 (NEWNAT/H, KEY, DESCR)
H
Wagner et al. 1990:1403 (K, O, M); Imada et al. 2000:12 (H)
Strong & Wagner 1997:43 (NEWNAT/K, KEY, DESCR)
H
Wagner et al. 1990:1404 (K); Oppenheimer & Bartlett 2002:7 (H)
M
H
Wagner et al. 1990:1405 (Ni, K, O, WM, H)
O Mo L
M
H
Wagner et al. 1990:1405 (Ni, K, O, Mo, L, M, H); Wood & LeGrande
2006:23 (Le)
O Mo L
M
H
Wagner et al. 1990:1405 (K, O, Mo, M, H); Oppenheimer 2007:23 (L)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 192
Scientific Name
Monocot
Cyperaceae
Status
Kl Le Ni K
O Mo L
Fimbristylis ferruginea (L.) Vahl
nat
Fimbristylis hawaiiensis Hillebr.
end
Fimbristylis littoralis Gaudich.
nat
Fimbristylis schoenoides (Retz.) Vahl
nat
Gahnia aspera (R.Br.) Spreng. ssp. globosa (H.Mann)
J.Kern
end
K
O
Gahnia beecheyi H.Mann
end
K
O Mo L
Gahnia lacera (A.Rich.) Steud.
nat
Gahnia vitiensis Rendle ssp. kauaiensis (Benl)
T.Koyama
end
Hypolytrum dissitiflorum Steud.
M Ka H
O
Citation
Strong & Wagner 1997:44 (NEWNAT/O, KEY, DESCR)
K
M
H
Wagner et al. 1990:1406 (H)
H
Strong & Wagner 1997:45 (NEWNAT/K [as Fimbristylis miliacea], KEY,
DESCR); Imada et al. 2000:12 (H); Oppenheimer 2003 (EM):11; Strong
2004:1069/Imada 2007:35 (NOM:Syn. F. miliacea)
H
Wagner et al. 1990:1406 (H)
Wagner et al. 1990:1408 (O); Lorence et al. 1995:34 (K)
M
H
L
Wagner et al. 1990:1409 (K, O, L, M, H); Hughes 1995:4 (Mo)
Wagner et al. 1990:1410 (L [as Gahnia lanaiensis]); Koyama et al. 2010:30
(NOM:Syn. G. lanaiensis, status changed from "end" to "nat")
K
Wagner et al. 1990:1410 (K)
?
Wagner et al. 1990:1382 [reported by Hillebrand in 1888; no vouchers seen]
Kyllinga brevifolia Rottb.
nat
K
O Mo L
M
H
Wagner et al. 1990:1411 (K, O, Mo, L, M, H)
Kyllinga nemoralis (J.R.Forst. & G.Forst.) Dandy ex
Hutch. & Dalziel
nat
K
O Mo L
M
H
Wagner et al. 1990:1413 (K, O, M, H); Oppenheimer 2003:11 (Mo, L)
Machaerina angustifolia (Gaudich.) T.Koyama
ind
K
O Mo L
M
H
Wagner et al. 1990:1414 (K, O, Mo, L, M, H)
Machaerina mariscoides (Gaudich.) J.Kern ssp. meyenii
(Kunth) T.Koyama
end
K
O Mo L
M
H
Wagner et al. 1990:1414 (K, O, Mo, L, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 193
Scientific Name
Monocot
Cyperaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Morelotia gahniiformis Gaudich.
end
K
Oreobolus furcatus H.Mann
end
K
O Mo
M
Rhynchospora caduca Elliott
nat
K
O Mo
M
H
Wagner et al. 1990:1428 (M, H); Wagner & Herbst 1995:18 (K); Wagner et
al. 1997:56 (O); Wysong et al. 2007:4 (Mo)
Rhynchospora chinensis Nees & Meyen ssp. spiciformis
(Hillebr.) T.Koyama
ind
K
O Mo
M
H
Wagner et al. 1990:1428 (K, Mo, M, H); Kennedy et al. 2010:21 (O)
Rhynchospora globularis (Chapm.) Small
nat
H
Wagner et al. 1990:1429 (H)
Rhynchospora radicans (Schltdl. & Cham.) H.Pfeiff.
ssp. microcephala (Bertero ex Spreng.) W.W.Thomas
nat
M
H
Strong & Wagner 1997:46 (NEWNAT/H, KEY, DESCR); Herbst et al.
2004:6 (EM)
Rhynchospora rugosa (Vahl) Gale ssp. lavarum
(Gaudich.) T.Koyama
ind
K
O Mo
M
H
Wagner et al. 1990:1429 (K, O, Mo, M, H)
Rhynchospora sclerioides Hook. & Arn.
ind
K
O Mo L
M
H
Wagner et al. 1990:1430 (K, O, Mo, L, M, H)
Rhynchospora tenuis Willd. ex Link ssp. tenuis
nat
K
Mo
Schoenoplectiella juncoides (Roxb.) Lye
ind
K
Schoenoplectus californicus (C.A.Mey.) Palla
nat?
Schoenoplectus mucronatus (L.) Palla ssp. mucronatus
nat
Schoenoplectus tabernaemontani (C.C.Gmel.) Palla
ind
Ni
Ni
K
K
Mo L
O Mo L
O Mo
M
H
Citation
Wagner et al. 1990:1409 (Mo, L, M, H [as Gahnia gahniiformis]); Lorence et
al. 1995:34/Wagner & Herbst 2003:52 (K, NOM:Syn. G. gahniiformis, status
change from "ind" to "end")
Wagner et al. 1990:1424 (K, O, Mo, M)
Imada 2007:36 (NEWNAT/K, DESCR); Oppenheimer 2008:27 (Mo)
M
H
Wagner et al. 1990:1431 (K, H [as Schoenoplectus juncoides]); Lye
2003:25/Wagner et al. 2012:82 (COMBNOV:Syn. S. juncoides)
H
Wagner et al. 1990:1431 (Ni, K, O, Mo, L, M, H)
H
Strong & Wagner 1997:47 (NEWNAT/H, KEY, DESCR); Wagner & Herbst
2003:54 (Haw'n plants are subsp. mucronatus)
H
Wagner et al. 1990:1432 (Ni, K, O, Mo, H [as Schoenoplectus lacustris
subsp. validus]); Smith 1995:98/Kennedy et al. 2010:21 (NOM:Syn. S.
lacustris subsp. validus)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 194
Scientific Name
Monocot
Cyperaceae
Status
Schoenus apogon Roem. & Schult.
nat
Scleria testacea Nees
ind
Uncinia brevicaulis Thouars
ind
Uncinia uncinata (L.f.) Kük.
ind
Kl Le Ni K
O Mo L
M Ka H
Mo
M
H
Herbarium Pacificum Staff 1996:4 (NEWNAT/H); Strong & Wagner 1997:48
(H, KEY, DESCR)
H
Wagner et al. 1990:1434 (WM, H); Oppenheimer 2008:27 (Mo)
M
K
Mo
M
Citation
Wagner et al. 1990:1435 (EM)
H
Wagner et al. 1990:1436 (K, Mo, M, H)
Dioscoreaceae
Dioscorea alata L.
pol
Non-naturalized Polynesian introduction; Wagner et al. 1990:1437; Staples &
Herbst 2005:658 (KEY, DESCR)
Dioscorea bulbifera L.
pol
K
O Mo
M
H
Wagner et al. 1990:1437 (K, O, Mo, M, H)
Dioscorea pentaphylla L.
pol
K
O Mo
M
H
Wagner et al. 1990:1437 (K, O, Mo, M, H)
H
Wagner et al. 1990:1440 (H [as Eriocaulon sp. A]); Herbst & Wagner
1999:20 (NOM: Eriocaulon sp. A identified)
Eriocaulaceae
Eriocaulon scariosum Sm.
nat
Heliconiaceae
Heliconia bihai (L.) L.
nat
Heliconia latispatha Benth.
nat
Heliconia metallica Planch. & Linden ex Hook.
nat
M
K
O
M
Wagner et al. 1990:1441 (EM); Staples & Herbst 2005:662 (KEY), 663
(DESCR)
H
O
Wagner et al. 1990:1441 (EM, H); Lorence & Flynn 1997:10 (K); Staples &
Herbst 2005:661 (KEY), 665 (DESCR); Daehler & Baker 2006:8 (O)
Wagner et al. 1990:1441 (O); Staples & Herbst 2005:662 (KEY), 665
(DESCR)
Hydrocharitaceae
Egeria densa Planch.
nat
K
O Mo
M
H
Wagner et al. 1990:1443 (K, O, Mo, H); Oppenheimer 2003:14 (WM);
Staples & Herbst 2005:666 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 195
Scientific Name
Monocot
Hydrocharitaceae
Status
Kl Le Ni K
Halophila decipiens Ostenf.
nat
Halophila hawaiiana Doty & B.C.Stone
end
K
Vallisneria americana Michx.
nat
K
Vallisneria spiralis L.
nat
K?
O Mo L
M Ka H
O
H
O Mo
M
M
O
Citation
McDermid et al. 2002 (NEWNAT/O, H, DESCR)
Wagner et al. 1990:1443 (K, O, Mo, M); McDermid et al. 2003 (DESCR)
H?
Wagner et al. 1990:1442 (K?, H? [possibly naturalizing in waterways, e.g.,
Wailua River, Kaua‘i and Wailoa estuary, Hilo, Hawai‘i]); Staples et al.
2003:13 (NEWNAT/K, DESCR); Staples & Herbst 2005:667 (DESCR);
Oppenheimer 2008:29 (WM)
H?
Wagner et al. 1990:1442 (K?, H? [possibly naturalizing in waterways, e.g.,
Wailua River, Kaua‘i and Wailoa estuary, Hilo, Hawai‘i]); Staples et al.
2003:13 (NEWNAT/O, DESCR); Staples & Herbst 2005:667 (DESCR)
H
Wagner et al. 1990:1445 (H); Herbarium Pacificum Staff 1999:4
(NEWNAT/H [as Aristea cf. compressa]); Staples et al. 2003:13 (NOM: A.
cf. compressa misapplied)
H
Wagner et al. 1990:1446 (K, O, M, H); Staples & Herbst 2005:669 (KEY,
DESCR); Oppenheimer 2010:36 (Mo)
Iridaceae
Aristea gerrardii Weim.
nat
Crocosmia Xcrocosmiiflora (Lemoine) N.E.Br.
nat
Dietes iridioides (L.) Sweet
nat
Gladiolus dalenii Van Geel
nat
M
H
Starr et al. 2004:23 (NEWNAT/EM, H, DESCR); Oppenheimer 2007:25
(WM)
Sisyrinchium acre H.Mann
end
M
H
Wagner et al. 1990:1446 (EM, H)
Sisyrinchium exile E.P.Bicknell
nat
M
H
Wagner et al. 1990:1447 (EM, H); Herbarium Pacificum Staff 1999:4 (Mo)
Sisyrinchium rosulatum E.P.Bicknell
nat
H
Wagner et al. 1990:1447 (H)
Trimezia martinicensis (Jacq.) Herb.
nat
K
O Mo
M
O
Staples & Herbst 2005:669 (KEY), 670 (DESCR); Frohlich & Lau 2012:39
(NEWNAT/O, DESCR)
Mo
O
Wagner et al. 1990:1449 (O); Staples & Herbst 2005:669 (KEY), 672
(DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 196
Scientific Name
Monocot
Iridaceae
Status
Watsonia borbonica (Pourr.) Goldblatt
nat
Watsonia meriana (L.) Mill.
nat
Kl Le Ni K
O Mo L
M Ka H
K
Citation
M
Wagner et al. 1990:1449 (EM); Lorence et al. 1995:39 (K); Oppenheimer &
Bartlett 2002:9 (WM); Staples & Herbst 2005:668 (KEY), 673 (DESCR)
M
Oppenheimer 2004:12 (NEWNAT/EM)
Joinvilleaceae
Joinvillea ascendens Gaudich. ex Brongn. & Gris ssp.
ascendens
end
K
O Mo
M
H
Wagner et al. 1990:1450 (K, O, Mo, M, H)
M
H
Wagner et al. 1990:1452 (M, H)
Juncaceae
Juncus acuminatus Michx.
nat
Juncus bufonius L.
nat
K
O Mo
M
H
Wagner et al. 1990:1452 (K, Mo, M, H); Wagner & Herbst 1995:21 (O)
Juncus effusus L.
nat
K
O Mo
M
H
Wagner et al. 1990:1453 (Mo, M, H); Staples & Herbst 2005:674 (DESCR);
Imada 2007:37 (O); Frohlich & Lau 2012:40 (K)
Juncus ensifolius Wikstr.
nat
M
H
Wagner et al. 1990:1453 (M? [reported but unvouchered], H); Oppenheimer
2004:13 (EM)
Juncus planifolius R.Br.
nat
O Mo L
M
H
Wagner et al. 1990:1453 (K, O, Mo, M, H); Oppenheimer & Bartlett 2002:9
(L)
Juncus polyanthemos Buchenau
nat
O
M
Juncus tenuis Willd.
nat
K
Mo
M
Luzula hawaiiensis Buchenau var. glabrata (Hillebr.)
O.Deg. & I.Deg.
end
K
Mo
M
Luzula hawaiiensis Buchenau var. hawaiiensis Buchenau
end
K
Luzula hawaiiensis Buchenau var. oahuensis (O.Deg. &
Fosberg) O.Deg. & I.Deg.
end
K
L
O
M
Wagner et al. 1990:1454 (O, M)
H
Wagner et al. 1990:1454 (K, Mo, EM, H)
Wagner et al. 1990:1455 (K, Mo, M)
H
Wagner et al. 1990:1455 (K, L, M, H)
Wagner et al. 1990:1455 (O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 197
Scientific Name
Monocot
Lemnaceae
Status
Landoltia punctata (G.Mey.) Les & D.J.Crawford
nat
Lemna aequinoctialis Welw.
nat?
Lemna obscura (Austin) Daubs
nat
Spirodela polyrhiza (L.) Schleid.
nat?
Wolffia globosa (Roxb.) Hartog & Plas
nat
Kl Le Ni K
O Mo L
M Ka H
O
K
O Mo
M
O
K
O
M
Citation
H
Wagner et al. 1997:59 (NEWNAT/O, H [as Spirodela punctata], KEY);
Staples et al. 2002:11 (NOM:Syn. S. punctata, DESCR)
H
Wagner et al. 1990:1457 (K, O, Mo, M, H? [as Lemna perpusilla; reported
but unvouchered from H]); Wagner et al. 1997:58 (H, NOM: L. perpusilla
misapplied, KEY); Staples & Herbst 2005:675 (DESCR)
H
Wagner et al. 1997:58 (NEWNAT/O, H, KEY)
H
Wagner et al. 1990:1458 (O); Wagner et al. 1997:59 (M, H, KEY); Imada et
al. 2000:12 (K); Staples et al. 2002:11 (DESCR); Staples & Herbst 2005:675
(DESCR)
O Mo
Wagner et al. 1997:59 (NEWNAT/O, Mo, KEY)
Liliaceae
Asparagus asparagoides (Forssk.) Deflers
nat
Asparagus densiflorus (Kunth) Jessop
nat
Asparagus falcatus L.
M
K
?
O
L
M
Starr et al. 2002:20 (NEWNAT/EM, DESCR)
H
O?
Asparagus plumosus Baker
nat
K
Astelia argyrocoma A.Heller ex Skottsb.
end
K
Astelia menziesiana Sm.
end
K
Lorence & Flynn 1999:4 (NEWNAT/K); Oppenheimer & Bartlett 2000:6
(WM); Kraus 2003:76 (O); Oppenheimer 2003:14 (H); Staples & Herbst
2005:676, 681 (KEY), 682 (DESCR); Oppenheimer 2007:26 (L)
Frohlich & Lau 2010:11 (NEWNAT/O, DESCR); Wagner et al. 2012:84 (O
record adventive)
O Mo L
M
H
Lorence et al. 1995:40 (NEWNAT/K [as Asparagus setaceus], DESCR);
Imada et al. 2000:13 (O); Oppenheimer & Bartlett 2000:6 (WM);
Oppenheimer & Bartlett 2002:9 (EM); Starr et al. 2002:21 (H); Staples &
Herbst 2005:676, 681 (KEY), 682 (NOM: A. setaceus misapplied, DESCR);
Starr et al. 2006:36 (Mo); Oppenheimer 2008:29 (L)
Wagner et al. 1990:1460 (K)
O Mo L
M
H
Wagner et al. 1990:1460 (K, O, Mo, L, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 198
Scientific Name
Monocot
Liliaceae
Status
Kl Le Ni K
O Mo L
M Ka H
K
Citation
Astelia waialealae Wawra
end
Wagner et al. 1990:1461 (K)
Dianella caerulea Sims var. assera R.J.F.Hend.
nat
Dianella sandwicensis Hook. & Arn.
ind
K
O Mo L
M
H
Wagner et al. 1990:1462 (K, O, Mo, L, M, H)
Hippeastrum striatum (Lam.) H.E.Moore
nat
K
Mo L
M
H
Wagner et al. 1990:1463 (K, H [as Hippeastrum puniceum]); Herbst &
Wagner 1999:23 (NOM: H. puniceum misapplied); Staples & Herbst
2005:678 (KEY), 689 (DESCR); Oppenheimer & Bartlett 2002:9 (EM, WM);
Oppenheimer 2008:29 (Mo, L)
Molineria capitulata (Lour.) Herb.
nat
K
Zephyranthes citrina Baker
nat
K
Zephyranthes grandiflora Lindl.
nat
O
Lau & Frohlich 2012:18 (NEWNAT/O, DESCR)
Flynn & Lorence 2002:15 (NEWNAT/K, DESCR); Staples & Herbst
2005:678 (KEY), 690 (DESCR)
O
M
Lorence et al. 1995:40 (NEWNAT/K, DESCR); Staples et al. 2002:12 (O,
EM); Staples & Herbst 2005:678, 693 (KEY), 694 (DESCR)
L
Staples & Herbst 2005:678, 693 (KEY), 694 (DESCR); Oppenheimer 2011:8
(NEWNAT/L, DESCR)
Marantaceae
Calathea crotalifera S.Watson
nat
O
H
Wagner et al. 1990:1464 (O); Staples & Herbst 2005:697 (KEY), 700
(DESCR); Parker & Parsons 2010:42 (H)
Pleiostachya pruinosa (W.Bull ex Regel) K.Schum.
nat
O
Staples & Herbst 2005:697 (KEY), 706 (DESCR); Daehler & Baker 2006:8
(NEWNAT/O, DESCR)
Stromanthe tonckat (Aubl.) Eichler
nat
O
Daehler & Baker 2006:9 (NEWNAT/O, DESCR)
Musaceae
Musa Xparadisiaca L.
pol
K
O Mo L
M
H
Wagner et al. 1990:1465 (K, O, Mo, L, M, H); Staples & Herbst 2005:707
(KEY [as Musa acuminata x M. balbisiana]), 708 (DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 199
Scientific Name
Monocot
Musaceae
Status
Kl Le Ni K
K?
O Mo L
Musa troglodytarum L.
pol
O Mo?
Musa velutina H.Wendl. & Drude
nat
O
Najas guadalupensis (Spreng.) Magnus
nat
O
Najas marina L.
nat
O
M Ka H
M?
H?
Citation
Wagner et al. 1990:1466 (K?, O, Mo?, M?, H? [only coll. at BISH is from
1960 in Pälolo Valley, O‘ahu]); Staples & Herbst 2005:707 (KEY), 710
(DESCR)
Staples & Herbst 2005:707 (KEY), 710 (DESCR); Daehler & Baker 2006:9
(NEWNAT/O, DESCR)
Najadaceae
H
Wagner et al. 1990:1466; Herbst & Wagner 1996:11 (NEWNAT/H,
DESCR); Imada 2007:37 (O)
Wagner et al. 1990:1466; Shannon & Wagner 1996:14 (NEWNAT/O)
Orchidaceae
Anoectochilus sandvicensis Lindl.
end
K
O Mo L
M
H
Wagner et al. 1990:1469 (K, O, Mo, L, M, H)
Arundina graminifolia (D.Don) Hochr.
nat
K
O
M
H
Wagner et al. 1990:1471 (K, O, EM, H); Oppenheimer & Bartlett 2000:7
(WM); Staples & Herbst 2005:713 (KEY), 716 (DESCR)
Cymbidium dayanum Rchb.f.
nat
H
Ackerman et al. 2011:1 (NEWNAT/H, DESCR); Pratt & Bio 2012:78 (H
range extension)
Dendrobium antennatum Lindl.
nat
O
Staples & Herbst 2005:713, 714, 722 (KEY), 722 (DESCR); Frohlich & Lau
2010:13 (NEWNAT/O, DESCR)
Dendrobium crumenatum Sw.
nat
O
Frohlich & Lau 2010:13 (NEWNAT/O, DESCR)
?
O?
Ackerman et al. 2011:2 (NEWNAT/O, DESCR); Wagner et al. 2012:85 (O
record adventive)
Dendrobium mirbelianum Gaudich.
nat
O
Frohlich & Lau 2012:41 (NEWNAT/O, DESCR)
Dendrobium rhombeum Lindl.
nat
O
Frohlich & Lau 2012:42 (NEWNAT/O, DESCR)
Dendrobium hybrid
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 200
Scientific Name
Monocot
Orchidaceae
Status
Kl Le Ni K
L
M
H
Citation
nat
Epidendrum nocturnum Jacq.
nat
O
Habenaria rodeiensis Barb.Rodr.
nat
O
M
Liparis hawaiensis H.Mann
end
K
O Mo L
M
H
Wagner et al. 1990:1472 (K, O, Mo, L, M, H)
Phaius tankarvilleae (Banks ex L'Hér.) Blume
nat
K
O Mo L
M
H
Wagner et al. 1990:1473 (K, O, L, H); Oppenheimer et al. 1999:9 (WM);
Staples & Herbst 2005:712 (KEY), 732 (DESCR); Oppenheimer 2007:28
(Mo); Oppenheimer 2008:31 (EM)
Platanthera holochila (Hillebr.) Kraenzl.
end
K
O Mo
M
Polystachya concreta (Jacq.) Garay & Sweet
nat
Spathoglottis plicata Blume
nat
Vanda tricolor Lindl.
Zeuxine strateumatica (L.) Schltr.
O
M Ka H
Epidendrum Xobrienianum Rolfe
Vanilla planifolia Jacks.
K
O Mo L
Frohlich & Lau 2012:42 (NEWNAT/O, DESCR)
Wagner et al. 1990:1468 (M? [as Habenaria sp.; reported as naturalized in
1983 in a Kula pasture]); Herbst & Wagner 1999:24 (NEWNAT/EM [as H.
rodeiensis]); Oppenheimer 2006:12 (WM); Lau & Frohlich 2012:19 (O)
Wagner et al. 1990:1474 (K, O, Mo, M)
O
K
Wagner et al. 1990:1472 (K, O, L, M, H); Staples & Herbst 2005:713, 714
(KEY), 726 (DESCR)
Staples et al. 2003:17 (NEWNAT/O, DESCR)
O Mo L
M
nat
O
M
?
O?
H
Wagner et al. 1990:1476 (K, O, Mo, L, M, H); Staples & Herbst 2005:713
(KEY), 736 (DESCR)
Staples & Herbst 2005:713, 737 (KEY), 737 (DESCR); Oppenheimer
2006:12 (NEWNAT/WM); Frohlich & Lau 2010:14 (O)
Staples & Herbst 2005:713 (KEY), 737 (DESCR); Daehler & Baker 2006:17
(O? [adventive])
nat
M
Wagner et al. 1990:1468 (M?); Oppenheimer 2003:18 (NEWNAT/EM,
DESCR)
Pandanaceae
Freycinetia arborea Gaudich.
ind
K
O Mo L
M
H
Wagner et al. 1990:1478 (K, O, Mo, L, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 201
Scientific Name
Monocot
Pandanaceae
Pandanus tectorius Parkinson ex Z
Status
Kl Le Ni K
ind?
Ni
K
O Mo L
M Ka H
O Mo L
M
Citation
H
Wagner et al. 1990:1479 (Ni, K, O, Mo, L, M, H); Staples & Herbst
2005:739 (KEY, DESCR)
H?
Wagner et al. 1990:1491 (H? [single coll. in 1912 from Glenwood], DESCR);
Herbst & Clayton 1998:18 (KEY)
Poaceae
Agrostis canina L.
?
Agrostis capillaris L.
nat
Agrostis exarata Trin.
?
M
Herbst & Clayton 1998:18 (NEWNAT/EM, KEY)
H?
Agrostis hyemalis (Walter) Britton, Sterns & Poggenb.
nat
Agrostis sandwicensis Hillebr.
end
K
O?
M
H
Wagner et al. 1990:1492 (O? [reported by Hillebrand in 1888, but
unvouchered], M, H); Herbst & Clayton 1998:18 (K, KEY)
Agrostis stolonifera L.
nat
K
O Mo
M
H
Wagner et al. 1990:1494 (K, O, Mo, M, H); Herbst & Clayton 1998:18 (KEY)
Aira caryophyllea L.
nat
K
O Mo
M
H
Wagner et al. 1990:1495 (Mo, M, H); Lorence & Flynn 1997:11 (K); Wood
2006:18 (O)
Ammophila arenaria (L.) Link
O
Wagner et al. 1990:1491 (H? [as Agrostis exarata var. monolepis; single coll.
in 1916 from northeastern slope of Mauna Kea], DESCR); Herbst & Clayton
1998:18 (KEY); Snow 2008:38 (NOM:Syn. A. exarata var. monolepis)
Snow & Lau 2010:46 (NEWNAT/O)
?
Andropogon bicornis L.
nat
Andropogon glomeratus (Walter) Britton, Sterns &
Poggenb. var. pumilus (Vasey) L.H.Dewey
nat
Only in Northwestern Hawaiian Islands; see NWHI checklist
K
Snow & Lau 2010:46 (NEWNAT/K, NOM: Andropogon glomeratus, A.
virginicus, Schizachyrium scoparium misapplied, in part, KEY, DESCR)
O
H
Herbst & Clayton 1998:18 (NEWNAT/K [as Andropogon glomeratus],
NOM: Schizachyrium condensatum misapplied, in part); Snow & Lau
2010:47 (assign varietal name, KEY, add O (NOM: Schizachyrium
condensatum misapplied, in part), H, remove K (spms. reidentified as A.
bicornis)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 202
Scientific Name
Monocot
Poaceae
Status
Andropogon virginicus L. var. virginicus
nat
Anthephora hermaphrodita (L.) Kuntze
nat
Anthoxanthum odoratum L.
nat
Aristida adscensionis L.
nat
Arrhenatherum elatius (L.) P.Beauv. ex J.Presl & C.Presl
nat
Arthraxon hispidus (Thunb.) Makino
Kl Le Ni K
K
O Mo L
M Ka H
O Mo
M
O
K
nat
O Mo
M
H
Wagner et al. 1990:1498 (Mo, M, H); Lorence et al. 1995:44 (K); Snow &
Davidse 2011:17 (O)
H
Wagner et al. 1990:1482 (O?, Mo?, H? [adventive], DESCR); Herbst &
Clayton 1998:19 (O voucher unidentifiable to species) [note: BISH spms.
confirm naturalization on H]
H?
Wagner et al. 1990:1482 (H? [single coll. in 1936 from a Küka‘iau pasture,
Hämäkua coast], DESCR); Starr et al. 2003:29 (NEWNAT/EM)
H?
Wagner et al. 1990:1482 (H? [single coll. in 1972 at 27-mile marker in
Volcano], DESCR)
H
Wagner et al. 1990:1498 (K, O, M, H); Staples & Herbst 2005:742 (KEY,
DESCR); Kennedy et al. 2010:22 (L)
H
Wagner et al. 1990:1499 (O? [2 colls.: in 1916, Schofield Barracks; in 1967,
along outer slope of Diamond Head], DESCR); Staples et al. 2002:13
(NEWNAT/H) [note: BISH spm. confirms naturalization on O]
M
H
Wagner et al. 1990:1499 (K, O, L, M, H)
O
M
H
Wagner et al. 1990:1500 (O, M, H)
O Mo
M
H
Oppenheimer 2003:19 (NEWNAT/Mo, EM, WM, DESCR); Oppenheimer
2004:15 (O); Starr et al. 2004:26 (K, EM); Staples & Herbst 2005:741
(KEY), 742 (DESCR); Oppenheimer 2007:29 (H)
O? Mo?
M
Avena barbata Pott ex Link
nat
Avena fatua L.
nat
Avena sativa L.
nat
Axonopus compressus (Sw.) P.Beauv.
nat
K
O
L
M
O
K
K
Wagner et al. 1990:1497 (O, H [as Andropogon virginicus); Hughes 1995:8
(Mo); Oppenheimer et al. 1999:9 (WM); Herbarium Pacificum Staff 1999:7
(L); Imada et al. 2000:14 (K); Snow & Lau 2010:48 (NOM: varietal name
assigned to selected K & O spms., remove L [spm. reidentified as
Hyparrhenia hirta]), KEY)
Herbst & Clayton 1998:19 (NEWNAT/O)
?
Arundo donax L.
H
Citation
O
L
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 203
Scientific Name
Monocot
Poaceae
Status
Axonopus fissifolius (Raddi) Kuhlm.
nat
Bothriochloa barbinodis (Lag.) Herter
nat
Bothriochloa bladhii (Retz.) S.T.Blake
nat
Bothriochloa laguroides (DC.) Herter
nat
Bothriochloa pertusa (L.) A.Camus
nat
Bouteloua curtipendula (Michx.) Torr.
Kl Le Ni K
K
Ni
Ni
K
?
O Mo L
M Ka H
O Mo L
M
H
Wagner et al. 1990:1500 (K, O, L, M, H); Hughes 1995:8 (Mo); Staples &
Herbst 2005:741 (KEY), 743 (DESCR)
O Mo
M
H
Wagner et al. 1990:1502 (Ni, O, Mo, M); Herbst & Clayton 1998:19 (KEY);
Starr et al. 2006:38 (H)
O Mo
M Ka H?
Wagner et al. 1990:1502 (Mo?, H? [collected in pastures]); Herbarium
Pacificum Staff 1997:18 (NEWNAT/EM); Herbst & Clayton 1998:19 (KEY);
Starr et al. 2006:38 (Ka); Snow 2008:38 (O); Frohlich & Lau 2010:14 (O,
same as Snow 2008) [note: BISH spms. confirm naturalization on Mo]
M
Herbst & Clayton 1998:19 (NEWNAT/EM, KEY)
O Mo L
M Ka
H
O?
Briza maxima L.
nat
K
Briza minor L.
nat
K
Bromus catharticus Vahl
nat
Bromus diandrus Roth
nat
Citation
Wagner et al. 1990:1503 (Ni, K, O, Mo, L, M, Ka, H); Herbst & Clayton
1998:19 (KEY)
Wagner et al. 1990:1482 (O? [single coll. in 1916 from Tantalus], DESCR)
M
H
Wagner et al. 1990:1504 (K, M, H)
O Mo L
M
H
Wagner et al. 1990:1505 (K, O, Mo, L, M, H); Imada et al. 2000:14 (H)
K
O Mo L
M
H
Wagner et al. 1990:1508 (O, Mo, M, H [as Bromus willdenowii]); Lorence &
Flynn 1997:11 (K); Herbst & Clayton 1998:20 (NOM:Syn. B. willdenowii,
KEY); Oppenheimer 2008:31 (L)
K
L
M
H
Wagner et al. 1990:1508 (K, H [as Bromus rigidus]); Herbst & Clayton
1998:20 (KEY); Herbarium Pacificum Staff 1999:7 (EM); Oppenheimer
2008:32 (L); Snow 2008:38 (NOM:Syn. B. rigidus); Starr & Starr 2011:32
(ELEV record)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 204
Scientific Name
Monocot
Poaceae
Status
Bromus hordeaceus L.
nat
Bromus madritensis L.
Bromus rubens L.
Citation
H
Wagner et al. 1990:1505 (O?, M? [as Bromus molliformis; adventive,
DESCR], 1507 (K, O, M, H [as B. mollis]; Herbst & Clayton 1998:20
(NOM:Syn. B. mollis [=B. hordeaceus subsp. hordeaceus], B. molliformis
[=B. hordeaceus subsp. molliformis], KEY); Wysong et al. 2007:6 (Mo);
Snow 2008:39 (NOM: no subspp. recognized)
nat
Mo
M
H
Wagner et al. 1990:1505 (Mo?, M?, H? [adventive], DESCR); Herbst &
Clayton 1998:20 (KEY); Herbst & Wagner 1999:25 (NEWNAT/Mo. EM, H)
nat
Mo
H
Wagner et al. 1990:1507 (Mo?, H? [adventive], DESCR); Herbst & Clayton
1998:20 (KEY); Herbst & Wagner 1999:25 (NEWNAT/Mo, H)
H?
Wagner et al. 1990:1507 (H? [single coll. in 1943 from Kïpuka Puaulu],
DESCR); Herbst & Clayton 1998:20 (KEY)
Bromus squarrosus L. var. villosus J.G.Gmel.
?
Bromus tectorum L.
M Ka H
M
?
K
O Mo L
O Mo
Bromus secalinus L.
Bromus sterilis L.
Kl Le Ni K
M?
nat
Mo
M
?
M?
Calamagrostis expansa (Munro ex Hillebr.) Hitchc.
end
M
Calamagrostis hillebrandii (Munro ex Hillebr.) Hitchc.
end
Cenchrus agrimonioides Trin. var. agrimonioides
end
Cenchrus agrimonioides Trin. var. laysanensis F.Br.
end
Mo?
O
L
Wagner et al. 1990:1507 (EM? [reported by Hillebrand in 1888 from
‘Ulupalakua; voucher not seen], DESCR)
H?
Wagner et al. 1990:1507 (H? [known from 2 colls. from Hämäkua pastures in
1936 & 1938; not vouchered since], DESCR); Herbst & Clayton 1998:20
(KEY); Oppenheimer 2008:32 (NEWNAT/Mo, WM)
Wagner et al. 1990:1507 (EM? [adventive], DESCR); Herbst & Clayton
1998:20 (KEY)
H
Wagner et al. 1990:1509 (M); Herbst & Clayton 1998:21 (H)
M
Wagner et al. 1990:1509 (Mo? [reported by Hitchcock in 1922 from central
Moloka‘i; voucher not seen], WM)
M
Wagner et al. 1990:1512 (O, L, M); Herbst & Clayton 1998:21 (KEY)
Only in Northwestern Hawaiian Islands; see NWHI checklist
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 205
Scientific Name
Monocot
Poaceae
Cenchrus americanus (L.) Morrone
Cenchrus caninus (Reinw. ex Blume) Morrone
Status
Kl Le Ni K
O Mo L
M Ka H
nat
M
Oppenheimer 2007:29 (NEWNAT/WM [as Pennisetum glaucum], DESCR);
Chemisquy et al. 2010:127 (NOM:Syn. P. glaucum); Wagner et al. 2012:98
(use Panicum glaucum as replacement name for Pennisetum glaucum; not
followed here)
?
Cenchrus ciliaris L.
nat
Cenchrus clandestinus (Hochst. ex Chiov.) Morrone
nat
Cenchrus complanatus (Nees) Morrone
nat
Cenchrus echinatus L.
nat
Cenchrus longisetus M.C.Johnst.
nat
Citation
Wagner et al. 1990:1578 [as Pennisetum macrostachyum; cultivated and
occasionally escaping]; Herbst & Clayton 1998:32 (KEY); Staples & Herbst
2005:742 (KEY), 750 (DESCR, incl. var. atropurpureum Anonymous);
Chemisquy et al. 2010:127 (NOM:Syn. P. macrostachyum; var.
atropurpureum not placed)
Le
K
O Mo L
M Ka
H
Wagner et al. 1990:1512 (K, O, Mo, L, M, Ka, H); Herbst & Clayton
1998:21 (KEY); Wood & LeGrande 2006:25 (Le)
K
O Mo L
M
H
Wagner et al. 1990:1578 (O, L, M, H [as Pennisetum clandestinum]); Hughes
1995:9 (Mo); Wagner & Herbst 1995:24 (K); Herbst & Clayton 1998:32
(KEY); Chemisquy et al. 2010:127/Wagner et al. 2012:98 (COMBNOV:Syn.
P. clandestinum)
H
Herbst & Clayton 1998:32 (NEWNAT/O, L, H [as Pennisetum
complanatum], KEY); Chemisquy et al. 2010:127/Wagner et al. 2012:98
(COMBNOV:Syn. P. complanatum)
H
Wagner et al. 1990:1512 (Ni, K, O, Mo, L, M, Ka, H); Herbst & Clayton
1998:21 (KEY); Wood & LeGrande 2006:25 (Le)
H
Herbst & Clayton 1998:32, 33 (NEWNAT/H [as Pennisetum villosum],
KEY); Wagner et al. 2012:99 (NOM:Syn. P. villosum = Cenchrus villosus);
Wagner et al. 2005–: (NOM: C. villosus is illegal name, replaced by C.
longisetus)
O
Kl Le Ni
K
L
O Mo L
M Ka
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 206
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
O Mo L
M Ka H
L
Citation
Cenchrus polystachios (L.) Morrone
nat
K
O
M
H
Wagner et al. 1990:1579 (O, L, H [as Pennisetum polystachion]); Herbst &
Clayton 1998:32 (KEY); Starr et al. 2002:23 (EM); Starr et al. 2003:30 (EM,
WM); Starr et al. 2004:27 (K); Starr & Starr 2011:30 (WM again);
Chemisquy et al. 2010:129/Wagner et al. 2012:99 (COMBNOV:Syn. P.
polystachion)
Cenchrus purpureus (Schumach.) Morrone
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:1579 (K, O, L, M, H [as Pennisetum purpureum]); Herbst
& Clayton 1998:32 (KEY); Oppenheimer 2003:22 (Mo); Starr & Starr
2011:30 (Ka); Chemisquy et al. 2010:129/Wagner et al. 2012:99
(COMBNOV:Syn. P. purpureum)
Cenchrus setaceus (Forssk.) Morrone
nat
K
O
M Ka
H
Wagner et al. 1990:1581 (K, O, L, H [as Pennisetum setaceum]); Herbst &
Clayton 1998:32 (KEY); Starr et al. 2006:39 (EM, Ka); Chemisquy et al.
2010:129/Wagner et al. 2012:99 (COMBNOV:Syn. P. setaceum)
Cenchrus setigerus Vahl
nat
K
Cenchrus tribuloides L.
nat
Chloris barbata Sw.
nat
Chloris divaricata R.Br. var. cynodontioides (Balansa)
Lazarides
nat
Chloris divaricata R.Br. var. divaricata
nat
Ni
L
Chloris gayana Kunth
nat
Ni
O Mo L
M
H
Wagner et al. 1990:1515 (Ni, O, L, M, H); Hughes 1995:8 (Mo)
Chloris radiata (L.) Sw.
nat
O Mo L
M
H
Wagner et al. 1990:1515 (K, O, L, M, H); Oppenheimer 2003:20 (Mo);
Wood & LeGrande 2006:25 (Le? [observed but unvouchered])
L
Mo?
Wagner et al. 1990:1511 (Mo? [single adventive coll. in 1976 from Kaluako‘i
District], DESCR); Herbst & Clayton 1998:21 (KEY); Starr et al. 2004:26
(NEWNAT/K)
Ka
Kl Le Ni
Le?
Lorence et al. 1995:44 (NEWNAT/Ka); Herbst & Clayton 1998:21 (KEY)
K
O Mo L
M Ka
K
O
M
Wagner et al. 1990:1515 (K, O); Oppenheimer 2003:20 (WM)
M Ka
Wagner et al. 1990:1515 (Ni, L, M, Ka)
K
H
Wagner et al. 1990:1514 (Ni, K, O, Mo, L, M, H); Herbst & Clayton 1998:21
(Ka); Wood & LeGrande 2006:25 (Le)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 207
Scientific Name
Monocot
Poaceae
Status
Chloris truncata R.Br.
nat
Chloris virgata Sw.
nat
Chrysopogon aciculatus (Retz.) Trin.
Kl Le Ni K
O Mo L
M Ka H
Citation
O
M? Ka
K
O Mo L
M Ka
H
Wagner et al. 1990:1515 (K, Mo, L, M, Ka, H); Herbst & Clayton 1998:21
(O); Imada et al. 2000:14 (O); Wood & LeGrande 2006:25 (Le)
ind?
K
O Mo L
M
H
Wagner et al. 1990:1516 (K, O, Mo, L, H); Oppenheimer et al. 1999:9 (WM)
Coix lachryma-jobi L.
nat
K
O Mo
M
H
Wagner et al. 1990:1517 (K, O, Mo, M, H)
Cortaderia jubata (Lemoine ex Carrière) Stapf
nat
Cortaderia selloana (Schult. & Schult.f.) Asch. &
Graebn.
nat
Cymbopogon refractus (R.Br.) A.Camus
nat
Cynodon aethiopicus Clayton & J.R.Harlan
nat
Cynodon dactylon (L.) Pers.
nat
Cynodon nlemfuensis Vanderyst
nat
Dactylis glomerata L.
nat
Le
Ni
K
K
K
Wagner et al. 1990:1513 (O? [adventive], EM? [reported by Whitney in 1939
from ‘Ulupalakua, but unvouchered], Ka? [adventive], DESCR); Herbst &
Wagner 1999:25 (NEWNAT/O, Ka)
M
Loope & Medeiros 1992: 7 (NEWNAT/EM); Meidell et al. 1998:8 (WM);
Staples & Herbst 2005:742 (KEY), 744 (DESCR)
O
M?
Loope & Medeiros 1992: 7 (EM? [reported but unvouchered]); Staples &
Herbst 2005:742 (KEY), 744 (DESCR); Frohlich & Lau 2010:15
(NEWNAT/O, DESCR)
O Mo L
M
H
Wagner et al. 1990:1519 (K, O, Mo, M, H); Herbst & Clayton 1998:21 (Ni,
L)
O
M
H
Herbst & Clayton 1998:22 (NEWNAT/O, H, KEY); Oppenheimer 2004:15
(WM)
O Mo L
M Ka
H
Wagner et al. 1990:1520 (K, O, L, M, Ka, H); Herbst & Clayton 1998:22
(Mo, KEY); Staples & Herbst 2005:741 (KEY), 745 (DESCR)
Mo
M
H
Wagner et al. 1990:1520 (Mo?, H? [adventive], DESCR); Herbst & Clayton
1998:22 (KEY); Herbst & Wagner 1999:25 (NEWNAT/Mo, H);
Oppenheimer 2003:20 (WM)
O Mo
M
H
Wagner et al. 1990:1521 (K, O, Mo, M, H); Starr & Starr 2011:32 (ELEV
record)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 208
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
K
O Mo L
M Ka H
O Mo L
M Ka
H
Citation
Dactyloctenium aegyptium (L.) Willd.
nat
Deschampsia caespitosa (L.) P.Beauv. ssp. beringensis
(Hultén) W.E.Lawr.
nat
Deschampsia nubigena Hillebr.
end
K
Mo
M
Dichanthelium cynodon (Reichardt) C.A.Clark & Gould
end
K
O Mo
M
Dichanthelium hillebrandianum (Hitchc.) C.A.Clark &
Gould
end
K
O Mo
M
Dichanthelium isachnoides (Munro ex Hillebr.)
C.A.Clark & Gould
end
K
Mo
M
Wagner et al. 1990:1526 (K, Mo, M); Wagner & Herbst 2003:60 (maintain
Dichanthelium as distinct from Panicum); Clayton & Snow 2010:55 (treated
as Panicum isachnoides)
Dichanthelium koolauense (H.St.John & Hosaka)
C.A.Clark & Gould
end
O
M
Wagner et al. 1990:1526 (O); Herbst & Clayton 1998:30 (WM [as Panicum
koolauense); Wagner & Herbst 2003:60 (maintain Dichanthelium as distinct
from Panicum); Clayton & Snow 2010:55 (treated as Panicum koolauense)
Dichanthium annulatum (Forssk.) Stapf
nat
O Mo
M
Wagner et al. 1990:1528 (O); Hughes 1995:8 (Mo); Oppenheimer & Bartlett
2002:10 (WM)
Dichanthium aristatum (Poir.) C.E.Hubb.
nat
Wagner et al. 1990:1528 (K, O, L, M); Herbst & Clayton 1998:22 (Ka)
Dichanthium sericeum (R.Br.) A.Camus
nat
Dichanthium tenue (R.Br.) A.Camus
nat
M
K
Wagner et al. 1990:1521 (O, Mo, M, Ka, H); Lorence & Flynn 1997:11 (K);
Oppenheimer 2007:29 (L)
Snow & Davidse 2011:17 (NEWNAT/EM, DESCR)
H
Wagner et al. 1990:1524 (K, Mo, M, H)
Wagner et al. 1990:1525 (K, Mo, M); Wagner & Herbst 2003:60 (maintain
Dichanthelium as distinct from Panicum); Clayton & Snow 2010:55 (treated
as Panicum cynodon); Wood 2012:97 (O)
H
Wagner et al. 1990:1526 (K, Mo, M, H); Wagner & Herbst 2003:60
(maintain Dichanthelium as distinct from Panicum); Clayton & Snow
2010:55 (treated as Panicum hillebrandianum); Wood 2012:97 (O)
O
L
M Ka
Ni
O
L
M Ka
H
Wagner et al. 1990:1529 (Ni, O, L, M, Ka, H)
Ni?
O
M
H?
Wagner et al. 1990:1528 (Ni?, H? [adventive, although no vouchers from
Hawai‘i Island at BISH], DESCR); Herbst & Clayton 1998:22 (NEWNAT/O,
EM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 209
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
Dichelachne crinita (L.f.) Hook.f.
nat
Dichelachne micrantha (Cav.) Domin
nat
K
Digitaria abyssinica (Hochst. ex A.Rich.) Stapf
nat
K
Digitaria bicornis (Lam.) Roem. & Schult.
nat
Digitaria ciliaris (Retz.) Koeler
nat
Digitaria divaricatissima (R.Br.) Hughes
nat
Digitaria eriantha Steud.
nat
Digitaria fuscescens (J.Presl) Henrard
nat
Digitaria horizontalis Willd.
nat
Digitaria insularis (L.) Mez ex Ekman
nat
Digitaria radicosa (J.Presl) Miq.
nat
O Mo L
M Ka H
M
Kl Le Ni
K
Wagner et al. 1990:1482 (L? [as Dichelachne sciurea; single coll. in 1938
from Maunalei Pipeline trail], DESCR); Herbst & Clayton 1998:22
(NOM:Syn. D. sciurea); Snow & Davidse 2011:18 (NEWNAT/K, NOM:
Garnotia acutigluma misapplied, in part)
M
Herbst & Clayton 1998:23 (NEWNAT/K, M, KEY)
M
Snow & Lau 2010:50 (NEWNAT/WM, DESCR)
M Ka
L
K
Wagner et al. 1990:1482 (EM? [single coll. in 1982 from Polipoli], DESCR);
Starr et al. 2003:29 (NEWNAT/EM)
L?
O Mo L
O?
H
Wagner et al. 1990:1530 (Ni, K, O, Mo, L, M, Ka, H); Herbst & Clayton
1998:23 (KEY); Wood & LeGrande 2006:26 (Le)
H
Wagner et al. 1990:1530 (H? [collected twice, last in 1953 from Parker
Ranch], DESCR); Herbst et al. 2004:9 (NEWNAT/L, H)
M Ka? H
H
O
Le
L?
K
O Mo L
K
O?
Citation
Wagner et al. 1990:1530 (Ka?, H? [as Digitaria pentzii; pasture grass],
DESCR); Herbst & Clayton 1998:23 (NEWNAT/O?, M?, H, NOM:Syn. D.
pentzii, KEY); Staples et al. 2003:17 (K); Oppenheimer 2004:15 (EM)
Wagner et al. 1990:1531 (H); Herbst & Clayton 1998:23 (KEY)
Wagner et al. 1990:1530 (O?, L? [adventive], DESCR); Herbst & Clayton
1998:23 (KEY); Herbst & Wagner 1999:26 (NEWNAT/O, L? [voucher from
1944 not seen])
M Ka
H
Wagner et al. 1990:1531 (K, O, Mo, L, M, Ka, H); Herbst & Clayton
1998:23 (KEY); Wood & LeGrande 2006:26 (Le)
H
Wagner et al. 1990:1530 (O? [adventive], DESCR); Herbst & Clayton
1998:23 (K, KEY); Staples et al. 2003:18 (H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 210
Scientific Name
Monocot
Poaceae
Status
Digitaria setigera Roth
ind
Digitaria violascens Link
Kl Le Ni K
M Ka H
Citation
K
O Mo L
M
H
Wagner et al. 1990:1531 (Ni, K, O, Mo, L, M, H); Herbst & Clayton 1998:23
(KEY)
nat
K
O Mo L
M
H
Wagner et al. 1990:1532 (K, O, Mo, L, M, H); Herbst & Clayton 1998:23
(KEY)
Diplachne fusca (L.) Stapf ssp. uninervia (J.Presl)
P.M.Peterson & N.Snow
nat
K
O Mo L
M
H
Wagner et al. 1990:1558 (K, O, Mo, H [as Leptochloa uninervia]); Warren &
Herbst 1994:3 (Ka); Herbst & Clayton 1998:29 (remove Ka [spms.
reidentified as Eragrostis curvula], add M); Imada et al. 2000:14 (NOM:Syn.
L. uninervia = L. fusca subsp. uninervia); Oppenheimer 2003:21 (L);
Peterson et al. 2012:1327/Wagner et al. 2012:96 (COMBNOV:Syn. L. fusca
subsp. uninervia)
Disakisperma dubia (Kunth) P.M.Peterson & N.Snow
nat
Dissochondrus biflorus (Hillebr.) Kuntze ex Hack.
end
Distichlis spicata (L.) Greene
nat
Echinochloa colona (L.) Link
nat
Echinochloa crusgalli (L.) P.Beauv.
nat
Echinochloa esculenta (A.Braun) H.Scholz
Ni
O Mo L
Mo
K
Kl
?
Ni
Snow & Davidse 2011:18 (NEWNAT/Mo [as Leptochloa dubia]); Peterson et
al. 2012:1327/Wagner et al. 2012:96 (COMBNOV:Syn. L. dubia)
O Mo L
M
O?
M
K
O Mo L
M Ka
H
Wagner et al. 1990:1535 (Ni, K, O, Mo, L, M, Ka, H); Herbst & Clayton
1998:24 (KEY)
K
O Mo L
M
H
Wagner et al. 1990:1534 (O? [as Echinochloa walteri; single coll. in 1936
from Mäkaha Valley], 1535 (K, O, Mo, L, M, H [as E. crus-galli]); Herbst &
Clayton 1998:24 (NOM: E. walteri misapplied, KEY)
M?
H
Wagner et al. 1990:1534 (K, O, Mo, L, M, H)
Wagner et al. 1990:1482 (O? [single coll. in 1977 from Ala Moana Park], M?
[3 colls. in 1980 near Kanahä Pond], DESCR); Herbarium Pacificum Staff
1999:7 (NEWNAT/M)
Herbst & Clayton 1998:24 (EM? [as Echinochloa utilis; volunteer in
experimental grass plot in ‘Ulupalakua in 1937], KEY); Snow 2008:39
(NOM:Syn. E. utilis)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 211
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Echinochloa oryzoides (Ard.) Fritsch
?
O?
Wagner et al. 1990:1534 (O? [as Echinochloa stagnina; adventive in wet
sites], DESCR); Herbst & Clayton 1998:24 (O? [2 colls.: Honouliuli rice field
in 1928, Kapahulu nursery in 1930], NOM: E. stagnina misapplied, KEY)
Echinochloa picta (J.König) P.W.Michael
?
O?
Wagner et al. 1990:1534 (O? [as Echinochloa stagnina; adventive in wet
sites], DESCR); Herbst & Clayton 1998:24 (O? [single coll. in 1927 from
Pearl Harbor rice paddy], NOM: E. stagnina misapplied, KEY)
Ehrharta calycina Sm.
nat
M?
Ehrharta erecta Lam.
nat
M
Ehrharta stipoides Labill.
nat
K
O Mo
M Ka
H
Wagner et al. 1990:1536 (O, M, H); Lorence et al. 1995:45 (K); Herbst &
Clayton 1998:25 (Ka); Herbst & Wagner 1999:26 (KEY); Wysong et al.
2007:6 (Mo)
Eleusine indica (L.) Gaertn.
nat
K
O Mo L
M
H
Wagner et al. 1990:1537 (K, O, Mo, L, M, H)
Enneapogon desvauxii P.Beauv.
?
Entolasia marginata (R.Br.) Hughes
nat
Eragrostis amabilis (L.) Wight & Arn.
nat
Eragrostis atropioides Hillebr.
end
Eragrostis brownii (Kunth) Nees ex Steud.
nat
H
Herbst & Clayton 1998:25 (NEWNAT/EM); Herbst & Wagner 1999:26
(KEY); Starr et al. 2008:47 (EM)
M?
O
Le Ni
K
K
Herbst & Wagner 1999:26 (NEWNAT/EM?, H, KEY)
Wagner et al. 1990:1482 (EM? [single coll. in 1937 in ‘Ulupalakua pasture],
DESCR)
H
Flynn & Lorence 1998:5 (NEWNAT/H); Frohlich & Lau 2012:44 (O)
O Mo L
M Ka
H
Wagner et al. 1990:1545 (Ni, O, M, Ka, H [as Eragrostis tenella]); Lorence et
al. 1995:45 (K, NOM:Syn. E. tenella); Hughes 1995:8 (Mo); Herbst &
Clayton 1998:25 (KEY); Oppenheimer 2003:20 (L); Wood & LeGrande
2006:26 (Le)
O Mo L
M
H
Wagner et al. 1990:1540 (O, Mo, L, M, H); Herbst & Clayton 1998:26 (KEY)
Mo
M
H
Wagner et al. 1990:1540 (Mo, M, H); Herbst & Clayton 1998:26, 27
(NOM:Syn. Eragrostis molokaiensis, KEY); Lorence & Flynn 1999:5 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 212
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
K
O Mo L
Eragrostis cilianensis (All.) Vignolo ex Janch.
nat
Eragrostis ciliaris (L.) R.Br.
nat
O
Eragrostis curvula (Schrad.) Nees
nat
O
Eragrostis deflexa Hitchc.
end
Mo L
Eragrostis dielsii Pilg. ex Diels & Pritz.
nat
Mo
Eragrostis elongata (Willd.) J.Jacq.
nat
Eragrostis fosbergii Whitney
end
Eragrostis grandis Hillebr.
end
Eragrostis leptophylla Hitchc.
end
Eragrostis leptostachya (R.Br.) Steud.
nat
Eragrostis mauiensis Hitchc.
end
Eragrostis monticola (Gaudich.) Hillebr.
end
Eragrostis parviflora (R.Br.) Trin.
nat
K
M Ka H
O Mo
M Ka
H
Wagner et al. 1990:1541 (K, O, Mo, M, Ka, H); Herbst & Clayton 1998:25
(KEY)
Wagner et al. 1990:1537 (O? [adventive], DESCR); Herbst & Clayton
1998:25 (KEY); Herbst & Wagner 1999:26 (NEWNAT/O)
M Ka
M
Herbst & Clayton 1998:26 (NEWNAT/O, Ka, KEY), 29 (NOM: Leptochloa
uninervia misapplied, in part); Oppenheimer 2003:20 (EM)
H
Wagner et al. 1990:1541 (L, H); Herbst & Clayton 1998:26 (WM, KEY);
Snow & Lau 2010:50 (Mo)
Snow & Lau 2010:50 (NEWNAT/Mo, KEY)
O Mo
H?
O
K
Citation
Wagner et al. 1990:1538 (K?, H? [adventive], DESCR); Herbst & Clayton
1998:26 (KEY); Lorence et al. 1995:45 (NEWNAT/K); Oppenheimer
2003:21 (Mo); Herbst et al. 2004:9 (O)
Wagner et al. 1990:1541 (O); Herbst & Clayton 1998:26 (KEY)
O Mo L
M Ka
H
Wagner et al. 1990:1542 (K, O, Mo, L, M, H); Herbst & Clayton 1998:26, 27
(Ka, KEY)
L
M Ka
H
Wagner et al. 1990:1542 (L, M, H); Herbst & Clayton 1998:26, 27 (Ka, KEY)
Mo
M
Wagner et al. 1990:1542 (Mo [as Eragrostis hosakai]); Herbst & Clayton
1998:26, 27 (Mo, NOM:Syn.E. hosakai, KEY, status change from "end" to
"nat"); Staples et al. 2002:14 (WM)
L
M
Wagner et al. 1990:1543 (L, WM); Herbst & Clayton 1998:26 (KEY)
Mo L
M
K
H
Wagner et al. 1990:1543 (Mo, L, M, H); Herbst & Clayton 1998:26 (KEY)
Flynn & Lorence 1998:5 NEWNAT/K); Herbst & Clayton 1998:25 (KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 213
Scientific Name
Monocot
Poaceae
Eragrostis paupera Jedwabn.
Status
Kl Le Ni K
ind
O Mo L
M Ka H
O
Citation
Wagner et al. 1990:1543 (O); Herbst & Clayton 1998:25 (KEY); Snow &
Lau 2010:50 (KEY)
Eragrostis pectinacea (Michx.) Nees var. miserrima
(E.Fourn.) Reeder
?
K?
Snow & Lau 2010:51 (K? [single coll. in 1936 from Hanapëpë)
Eragrostis pectinacea (Michx.) Nees var. pectinacea
nat
K
O Mo L
Eragrostis pilosa (L.) P.Beauv.
nat
K
O
Eragrostis sessilispica Buckley
?
M Ka
H
L
Wagner et al. 1990:1545 (O, L, M, H); Lorence et al. 1995:45 (K); Herbst &
Clayton 1998:25, 27 (Mo, KEY); Oppenheimer 2006:13 (Ka); Snow & Lau
2010:51 (typical variety acknowledged with addition of var. miserrima)
Wagner et al. 1990:1538 (Mo?, M?, H? [adventive], DESCR); Lorence et al.
1995:45 (NEWNAT/K); Herbst & Clayton 1998:25, 27 (add L, remove Mo
[spm. reidentified as Eragrostis monticola], M [spm. reidentified as E.
pectinacea], H [spm. reidentified as E. leptophylla], KEY; Snow & Lau
2010:51 (O)
O?
Snow 2008:41 (O? [single coll. in 1938 in Wahiawä research plot])
Eragrostis superba Peyr.
nat
Eragrostis tenuifolia (A.Rich.) Hochst. ex Steud.
nat
Eragrostis trichophora Coss. & Durieu
nat
M
Eragrostis unioloides (Retz.) Nees ex Steud.
nat
M
H
Wagner et al. 1990:1538 (H? [adventive], DESCR); Herbst & Clayton
1998:25 (KEY); Herbst & Wagner 1999:26 (NEWNAT/H); Oppenheimer
2008:32 (EM)
Eragrostis variabilis (Gaudich.) Steud.
end
M Ka
H
Wagner et al. 1990:1545 (Ni, K, O, Mo, L, M, Ka, H); Herbst & Clayton
1998:26 (KEY); Wood & LeGrande 2006:26 (Le? [observed but
unvouchered]); Snow & Lau 2010:50 (NOM:Syn. Eragrostis hobdyi)
Eriochloa procera (Retz.) C.E.Hubb.
nat
K
Le? Ni
K
O
L
O Mo L
Mo
M
H
Herbst & Clayton 1998:25, 28 (NEWNAT/H, KEY); Starr & Starr 2011:30
(EM)
M
H
Herbst & Clayton 1998:26, 28 (NEWNAT/O, H, KEY); Oppenheimer
2004:15 (EM); Oppenheimer 2008:32 (L); Snow 2008:39 (K)
Snow & Lau 2010:51 (NEWNAT/M, DESCR)
Oppenheimer 2008:32 (NEWNAT/Mo, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 214
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
Eriochloa punctata (L.) Desv. ex Ham.
nat
Eustachys petraea (Sw.) Desv.
nat
Festuca aloha Catalán, Soreng & P.M.Peterson
end
K
Festuca bromoides L.
nat
K
Festuca hawaiiensis Hitchc.
end?
Festuca molokaiensis Soreng, P.M.Peterson & Catalán
end
Festuca myuros L.
nat
Festuca rubra L.
nat
Garnotia acutigluma (Steud.) Ohwi
nat?
Gastridium ventricosum (Gouan) Schinz & Thell.
nat
Glyceria notata Chevall.
nat
Hackelochloa granularis (L.) Kuntze
nat
Ni
O Mo L
M Ka H
O Mo
M
Citation
Wagner et al. 1990:1546 (Ni, O); Oppenheimer 2003:21 (Mo, WM)
Only in Northwestern Hawaiian Islands; see NWHI checklist
Catalán et al. 2009:51 (SPNOV/K, DESCR, KEY)
O Mo L
M
H
Wagner et al. 1990:1603 (K, O, Mo, L, M, H [as Vulpia bromoides]); Starr &
Starr 2011:32 (ELEV record); Wagner et al. 2012:101 (NOM:Syn. V.
bromoides)
M?
H
Wagner et al. 1990:1547 (EM? [reported by Hillebrand in 1888, presumably
from ‘Ulupalakua, but spm. likely destroyed], H); Catalán et al. 2009:57
(KEY)
Mo
K
O Mo L
Mo
K
Catalán et al. 2009:54 (SPNOV/Mo, DESCR, KEY)
M
H
Wagner et al. 1990:1603 (K, O, Mo, L, M, H [as Vulpia myuros]); Starr &
Starr 2011:32 (ELEV); Wagner et al. 2012:101 (NOM:Syn. V. myuros)
M
H
Wagner et al. 1990:1547 (M, H); Wagner et al. 1997:60 (K); Oppenheimer
2008:33 (Mo); Starr & Starr 2011:32 (ELEV); Oppenheimer 2011:9 (remove
K [=Festuca aloha]. Mo [=F. molokaiensis]); Catalán et al. 2009:57 (KEY)
M
H
Wagner et al. 1990:1548 (Mo, M, H); Lorence & Flynn 1999:5 (K); Snow &
Davidse 2011:18 (remove K [spm. reidentified as Dichelachne micrantha])
M
H
Wagner et al. 1990:1550 (K, M, H)
M
H
Wagner et al. 1990:1482 (EM? [single coll. in 1916 in Haleakalä Crater], H?
[2 colls., last in 1953, from North Kohala pasture] [as Glyceria fluitans],
DESCR); Herbst & Wagner 1999:27 (NEWNAT/EM, H); Snow 2008:39
(EM, H, NOM: G. fluitans misapplied)
H
Wagner et al. 1990:1483 (H? [single coll. in 1941 from Ka‘ü]); Herbst &
Wagner 1999:27 (NEWNAT/H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 215
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
O Mo L
M Ka H
M
Citation
Hemarthria altissima (Poir.) Stapf & C.E.Hubb.
nat
Heteropogon contortus (L.) P.Beauv. ex Roem. &
Schult.
ind?
Holcus lanatus L.
nat
Hordeum brachyantherum Nevski
nat
Hordeum murinum L. ssp. leporinum (Link) Arcang.
nat
Hordeum vulgare L.
nat
Hyparrhenia dregeana (Nees) Stent
nat
L
Hyparrhenia hirta (L.) Stapf
nat
Mo L
Hyparrhenia rufa (Nees) Stapf
nat
K
O Mo
M
H
Wagner et al. 1990:1554 (K, O, Mo, M, H); Herbst & Clayton 1998:28 (KEY)
Isachne distichophylla Munro ex Hillebr.
end
K
O Mo L
M
H
Wagner et al. 1990:1555 (K, O, Mo, L, M, H)
Isachne pallens Hillebr.
end
K
O
Ischaemum byrone (Trin.) Hitchc.
end
K
O Mo
Le Ni
Starr et al. 2004:26 (NEWNAT/EM, DESCR)
K
O Mo L
M Ka
H
Wagner et al. 1990:1550 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:26 (Le)
K
O Mo
M
H
Wagner et al. 1990:1551 (K, O, Mo, M, H); Oppenheimer et al. 1999:10
(WM)
H
Herbst & Clayton 1998:28 (NEWNAT/H, KEY)
M
H
Wagner et al. 1990:1552 (L, M, H [as Hordeum leporinum], O?, M?, H? [as
H. murinum; adventive]); Herbst & Clayton 1998:28 (NOM:Syn. H.
leporinum, KEY) [Note: no O‘ahu spms. at BISH]
M
H
Wagner et al. 1990:1552 (K?, O?, M?, H? [cultivated, occasional escape],
DESCR); Herbst & Clayton 1998:28 (KEY); Herbst & Wagner 1999:27
(NEWNAT/K, O, EM, H)
O?
K
L
O
Herbst & Wagner 1996:11 (NEWNAT/L, DESCR); Herbst & Clayton
1998:28 (KEY)
Wagner et al. 1990:1554 (Mo?, L? [adventive], DESCR); Wagner & Herbst
1995:24 (NEWNAT/Mo, L); Herbst & Clayton 1998:28 (KEY)
Wagner et al. 1990:1555 (K, O)
M
H
Wagner et al. 1990:1556 (Mo, M, H); Lorence et al. 1995:46 (K); Herbst &
Clayton 1998:29 (KEY); Imada et al. 2000:15 (O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 216
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Ischaemum ciliare Retz.
nat
M
Wagner et al. 1990:1556 (M? [as Ischaemum indicum; adventive]); Herbst &
Clayton 1998:29 (NEWNAT/EM, KEY); Sun & Phillips 2008:611/Snow &
Lau 2010:52 (NOM: I. indicum misapplied)
Ischaemum timorense Kunth
nat
M
Herbst & Wagner 1996:11 (NEWNAT/EM, DESCR); Herbst & Clayton
1998:29 (KEY)
Ixophorus unisetus (J.Presl) Schltdl.
?
Koeleria macrantha (Ledeb.) Schult.
nat
Lachnagrostis filiformis (G.Forst.) Trin.
ind
Lamarckia aurea (L.) Moench
Leptochloa panicea (Retz.) Ohwi ssp. brachiata (Steud.)
N.Snow
K? O?
K
L?
O Mo L
?
?
Lepturus repens (G.Forst.) R.Br.
ind
Leymus triticoides (Buckley) Pilg.
nat
M Ka
H
Wagner et al. 1990:1557 (H [as Koeleria nitida]); Herbst & Clayton 1998:29
(NOM:Syn. K. nitida)
H
Wagner et al. 1990:1492 (K, O, Mo, L, M, H [as Agrostis avenacea]); Herbst
& Clayton 1998:18 (KEY); Imada et al. 2000:14 (Ka); Snow 2008:39
(NOM:Syn. A. avenacea)
M?
nat
Leptochloa virgata (L.) P.Beauv.
Wagner et al. 1990:1483 (K?, O?, L? [vouchered but status unknown],
DESCR); Herbst & Wagner 1999:27 (K? [single coll. in 1941 from Kekaha],
O? [last coll. in 1943 from Mänoa Valley], L? [single coll. from Kö‘ele, no
date])
Wagner et al. 1990:1483 (EM? [single coll. in 1937 from Makawao pasture],
DESCR)
O
Frohlich & Lau 2012:44 (NEWNAT/O, DESCR)
O? Mo?
Wagner et al. 1990:1558 (O? [single coll. in 1911 from Honolulu], Mo?
[single coll. in 1916 from Kamoku])
Le?
Wood & LeGrande 2006:26 (Le? [observed but unvouchered])
H
Wagner et al. 1990:1482 (H? [as Elymus triticoides; single coll. in 1936 in
Humu‘ula pasture]); Herbst & Clayton 1998:29 (NEWNAT/H); Soreng et al.
2003:425 (NOM:Syn. E. triticoides)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 217
Scientific Name
Monocot
Poaceae
Status
Lolium multiflorum Lam.
nat
Lolium perenne L.
Kl Le Ni K
M Ka H
Citation
O Mo
M
H
Wagner et al. 1990:1561 (K, O, M, H); Herbst & Clayton 1998:29 (Mo);
Herbst & Wagner 1999:28 (KEY)
nat
O
M
H
Wagner et al. 1990:1561 (H); Starr et al. 2003:30 (EM); Frohlich & Lau
2008:8 (O); Herbst & Wagner 1999:28 (KEY)
Lolium temulentum L.
nat
O
Melinis minutiflora P.Beauv.
nat
Melinis nerviglumis (Franch.) Zizka
nat
Miscanthus floridulus (Labill.) Warb. ex K.Schum. &
Lauterb.
nat
Muhlenbergia mexicana (L.) Trin.
nat
Muhlenbergia microsperma (DC.) Trin.
nat
Nassella cernua (Stebbins & Á.Löve) Barkworth
nat
Oplismenus compositus (L.) P.Beauv.
K
?
Melinis repens (Willd.) Zizka
Oplismenus burmannii (Retz.) P.Beauv.
K
O Mo L
Wagner et al. 1990:1560 (O? [as Lolium tementulum (spelling error);
adventive], DESCR); Herbst & Wagner 1999:28 (NEWNAT/O, KEY)
O Mo L
M Ka
H
O?
Ni
K
Snow & Davidse 2011:20 (O? [adventive], DESCR)
O Mo L
M Ka
H
O
M
Herbst & Clayton 1998:30 (NEWNAT/EM)
M
Wagner et al. 1990:1562 (O, WM)
H
K?
nat
K
Wagner et al. 1990:1588 (Ni, K, O, Mo, L, M, Ka, H [as Rhynchelytrum
repens]); Herbst & Clayton 1998:29 (NOM:Syn. R. repens, KEY)
Frohlich & Lau 2012:45 (NEWNAT/O, DESCR)
O
?
Wagner et al. 1990:1562 (K, O, Mo, L, M, Ka, H); Herbst & Clayton
1998:29 (KEY)
Wagner et al. 1990:1599 (H [as Stipa cernua]); Barkworth 1990:609/Snow
2008:39 (NOM:Syn. S. cernua)
Wagner et al. 1990:1563 (K? [2 colls.: in 1916 near Wailua Falls; in 1957
from Hanakoa], DESCR)
O Mo
M
H
Wagner et al. 1990:1563 (K, O, Mo, H); Oppenheimer 2003:21 (WM);
Oppenheimer 2008:33 (EM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 218
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Oplismenus hirtellus (L.) P.Beauv. ssp. hirtellus
nat
K
O Mo L
M
H
Wagner et al. 1990:1565 (K, O, Mo, L, M, H [as Oplismenus hirtellus]);
Staples & Herbst 2005:741 (KEY), 750 (DESCR [as O. hirtellus]); Snow &
Lau 2010:52 (designated as subsp. hirtellus)
Oplismenus hirtellus (L.) P.Beauv. ssp. undulatifolius
U.Scholz
nat
K
O Mo
M
H
Snow & Lau 2010:52 (NEWNAT/K, O, Mo, EM, H, DESCR)
Panicum A
nat
Mo
Panicum antidotale Retz.
nat
O Mo
M
Panicum beecheyi Hook. & Arn.
end
O
M
Wagner et al. 1990:1567 (K, O, M)
M?
Wagner et al. 1990:1567 (Mo, EM); Herbst & Clayton 1998:30 (remove Mo
[=Panicum sp. A], single EM coll. in 1939 grown in Haleakalä branch
experimental station, current status unknown)
Panicum coloratum L.
K
?
Herbst & Clayton 1998:30 (NEWNAT/Mo)
H
Panicum dichotomiflorum Michx.
nat
O
Panicum fauriei Hitchc. var. carteri (Hosaka) Davidse
end
O Mo L
M
Panicum fauriei Hitchc. var. fauriei
end
O Mo L
M Ka
H
Wagner et al. 1990:1568 (O, Mo, M, Ka, H); Herbst & Clayton 1998:30 (L)
Panicum fauriei Hitchc. var. latius (H.St.John) Davidse
end
K
O Mo L
M Ka
H
Wagner et al. 1990:1568 (K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:26 (Le); Imada 2007:38 (Ni)
Panicum konaense Whitney & Hosaka
end
K
Mo
M
H
Wagner et al. 1990:1568 (Mo, M, H); Lorence et al. 1995:46 (K)
Panicum lineale H.St.John
end
K
Panicum longivaginatum H.St.John
end
Panicum miliaceum L.
nat
Le Ni
K
H
Wagner et al. 1990:1567 (O, Mo, H); Starr et al. 2003:30 (EM, WM); Starr et
al. 2004:27 (EM)
Wagner et al. 1990:1568 (O, H)
Wagner et al. 1990:1568 (O, Mo, M); Herbst & Clayton 1998:30 (L)
Wagner et al. 1990:1569 (K)
O
M
H
St. John 1987c:370 (SPNOV/H, DESCR); Herbst & Clayton 1998:31
(acceptance of species)
H
Wagner et al. 1990:1570 (K, O, Mo, L, M, H); Lorence & Flynn 1997:11 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 219
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
Panicum nephelophilum Gaudich.
end
Panicum niihauense H.St.John
end
Ni
Panicum pellitum Trin.
end
Le Ni
Panicum ramosius Hitchc.
end
Panicum repens L.
nat
Panicum tenuifolium Hook. & Arn.
end
Panicum torridum Gaudich.
end
Panicum virgatum L.
K
O Mo L
M Ka H
O Mo L
M
K
Wagner et al. 1990:1570 (K, O, Mo, L, M, H)
Wagner et al. 1990:1570 (Ni); Lorence et al. 1995:46 (K)
O
L
M
Mo L
O
Kl Le Ni
H
Citation
H
Ka
Wagner et al. 1990:1571 (Ni, L, M, H); Herbst & Clayton 1998:31 (O);
Wood & LeGrande 2006:27 (Le)
Wagner et al. 1990:1571 (Mo, L, Ka)
L
M
H
Wagner et al. 1990:1571 (O, L, H); Starr et al. 2002:23 (EM)
K
O Mo L
M
H
Wagner et al. 1990:1573 (O, Mo, L, M, H); Herbst & Clayton 1998:31 (K)
K
O Mo L
M Ka
H
Wagner et al. 1990:1573 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:27 (Le)
?
O?
Wagner et al. 1990:1574 (O); Snow & Lau 2010:52 (O? [2 colls. in 1938 &
1940 from Poamoho experimental station])
Panicum xerophilum (Hillebr.) Hitchc.
end
Ni
O Mo L
M Ka
H
Wagner et al. 1990:1574 (Ni, O, Mo, L, M, Ka, H)
Paspalidium distans (Trin.) Hughes
nat
Ni
Paspalum arundinaceum Poir.
nat
Paspalum conjugatum P.J.Bergius
nat
K
O Mo L
M
H
Wagner et al. 1990:1575 (K, O, Mo, L, M, H); Herbst & Clayton 1998:32
(KEY); Staples & Herbst 2005:741 (KEY), 746 (DESCR); Snow & Lau
2010:55 (KEY)
Paspalum dilatatum Poir.
nat
K
O Mo L
M
H
Wagner et al. 1990:1576 (K, O, Mo, L, M, H); Herbst & Clayton 1998:32
(KEY); Snow & Lau 2010:55 (KEY)
Herbst & Clayton 1998:31 (NEWNAT/Ni)
M
Snow & Lau 2010:53, 56 (NEWNAT/EM [as Paspalum aff. laxum], KEY);
Snow & Davidse 2011:18 (NOM: P. aff. laxum misapplied )
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 220
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
O Mo L
Paspalum distichum L.
nat
K
O
Paspalum fimbriatum Kunth
nat
K
O Mo L
Paspalum longifolium Roxb.
nat
K
Paspalum macrophyllum Kunth
nat
Paspalum malacophyllum Trin.
nat
Paspalum mandiocanum Trin. var. mandiocanum
nat
Paspalum notatum Flüggé
nat
Paspalum paniculatum L.
nat
Paspalum pilosum Lam.
nat
Paspalum scrobiculatum L.
ind?
Paspalum setaceum Michx.
nat
M Ka H
Wagner et al. 1990:1575 (K?, O? [adventive], DESCR); Herbst & Clayton
1998:32 (KEY); Herbst & Wagner 1999:28 (NEWNAT/K, O); Staples &
Herbst 2005:741 (KEY), 746 (DESCR); Snow & Lau 2010:55 (KEY)
M
Wagner et al. 1990:1576 (K, O); Herbst & Clayton 1998:31 (KEY); Starr et
al. 2002:23 (EM); Oppenheimer 2003:21 (Mo, WM); Oppenheimer 2008:33
(L); Snow & Lau 2010:55 (KEY)
M
Staples et al. 2003:18 (NEWNAT/K, DESCR); Oppenheimer 2004:15 (EM);
Snow & Lau 2010:55 (KEY)
H
Herbst & Clayton 1998:32 (NEWNAT/H, KEY); Snow & Lau 2010:56
(KEY)
M
Oppenheimer 2004:16 (NEWNAT/WM); Snow & Lau 2010:55 (KEY)
O Mo
M
Snow & Lau 2010:53, 55 (NEWNAT/O, EM [as Paspalum aff. thunbergii],
DESCR, KEY); Snow & Davidse 2011:18 (Mo, NOM: P. aff. thunbergii
misapplied)
Mo
M
Lorence & Flynn 1999:6 (NEWNAT/K); Oppenheimer 2007:29 (EM, WM);
Oppenheimer 2008:33 (Mo); Snow & Lau 2010:55 (KEY)
K
O
M
H
Mo
K
Citation
O Mo
Wagner et al. 1990:1575 (O?, H? [adventive], DESCR); Herbst & Clayton
1998:32 (KEY); Herbst & Wagner 1999:28 (NEWNAT/O, H); Oppenheimer
2004:16 (WM); Oppenheimer 2007:29 (EM); Snow & Lau 2010:53 (remove
EM, spm. reidentified as P. aff. laxum [= P. arundinaceum], 56 (KEY)
Snow & Lau 2010:53, 56 (NEWNAT/Mo, DESCR, KEY)
M
M
H
Wagner et al. 1990:1576 (K, O, Mo, M, H); Herbst & Clayton 1998:32
(KEY); Snow & Lau 2010:55 (KEY)
Oppenheimer 2007:29 (NEWNAT/EM); Snow & Lau 2010:56 (KEY)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 221
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Paspalum unispicatum (Scribn. & Merr.) Nash
nat
K
Paspalum urvillei Steud.
nat
K
O Mo L
M
H
Wagner et al. 1990:1577 (K, O, Mo, L, M, H); Herbst & Clayton 1998:32
(KEY); Snow & Lau 2010:55 (KEY)
Paspalum vaginatum Sw.
nat
K
O Mo
M
H
Wagner et al. 1990:1577 (K, O, H); Herbst & Clayton 1998:32 (KEY);
Herbarium Pacificum Staff 1999:7 (M); Oppenheimer 2003:22 (Mo); Staples
& Herbst 2005:741 (KEY), 746 (DESCR, treated as syn. of Paspalum
distichum); Snow & Lau 2010:55 (KEY)
Paspalum virgatum L.
nat
Phalaris aquatica L.
nat
Phalaris canariensis L.
nat
Phalaris paradoxa L.
?
Phleum pratense L.
nat
Phragmites australis (Cav.) Trin. ex Steud.
O
?
Phalaris minor Retz.
?
Snow & Davidse 2011:19 (NEWNAT/K, DESCR)
Snow & Lau 2010:54, 55 (NEWNAT/O, DESCR, KEY)
O?
M
H
Wagner et al. 1990:1483 (M?, H? [as Phalaris tuberosa var. stenoptera;
escape from cultivation], DESCR); Herbst & Clayton 1998:33
(NEWNAT/EM, H, NOM: P. tuberosa var. stenoptera misapplied, KEY)
M?
H?
Wagner et al. 1990:1483 (O? [single coll. in 1958 as volunteer in Nu‘uanu
garden], M? [single coll. in 1933 in Makawao cultivated field], H? [single
coll. in 1933 in Parker Ranch pasture], DESCR); Herbst & Clayton 1998:33
(KEY)
H
Wagner et al. 1990:1483 (O?, Mo?, H? [escape], DESCR); Herbst & Clayton
1998: 33 (KEY); Herbst & Wagner 1999:28 (NEWNAT/O, Mo, H)
H?
Wagner et al. 1990:1483 (WM? [single coll. in 1937 from Waiakoa], H?
[single coll. in 1903 from Honoka‘a], DESCR); Herbst & Clayton 1998:33
(KEY)
O Mo
M?
K
O
M?
O?
Wagner et al. 1990:1483 (K?, O?, M? [escaped pasture grass], DESCR);
Herbst & Wagner 1999:29 (NEWNAT/K, O, EM? [single coll. in 1941 from
Makawao experimental station])
Wagner et al. 1990:1483 (O? [as Phragmites karka; 2 colls. in 1956 & 1959
near Hickam Field; not vouchered since]); Herbst & Clayton 1998:33 (NOM:
P. karka misapplied)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 222
Scientific Name
Monocot
Poaceae
Status
Phyllostachys aurea Carrière ex Rivière & C.Rivière
nat
Phyllostachys nigra (Lodd. ex Lindl.) Munro var.
henionis (Mitford) Stapf ex Rendle
nat
Piptatherum miliaceum (L.) Coss.
nat
Poa annua L.
nat
Poa compressa L.
Kl Le Ni K
O Mo L
M Ka H
O
K
K
Citation
Staples et al. 2002:14 (NEWNAT/O, DESCR); Staples & Herbst 2005:742
(KEY), 747 (DESCR)
O Mo L
O Mo L
M
M
?
Wagner et al. 1990:1582 (O, Mo, M); Staples & Herbst 2005:742 (KEY), 747
(DESCR); Oppenheimer 2011:9 (K, L)
H
Dougill & Banko 2002 (NEWNAT/H [as Oryzopsis miliacea]); Snow
2008:40 (NOM:Syn. O. miliacea)
H
Wagner et al. 1990:1583 (K, O, Mo, L, M, H); Starr & Starr 2011:32 (ELEV
record)
H?
Wagner et al. 1990:1583 (H? [single coll. in 1916 from Humu‘ula sheep
station], DESCR)
Poa mannii Munro ex Hillebr.
end
K
Wagner et al. 1990:1584 (K); Lorence et al. 1995:46 (REDISCOVER/K)
Poa pratensis L.
nat
Poa sandvicensis (Reichardt) Hitchc.
end
K
Wagner et al. 1990:1584 (K)
Poa siphonoglossa Hack.
end
K
Wagner et al. 1990:1585 (K)
Polypogon fugax Nees ex Steud.
nat
Polypogon interruptus Kunth
nat
K
O
M
H
Wagner et al. 1990:1585 (K, O, M, H); Herbst & Clayton 1998:34 (KEY)
Polypogon monspeliensis (L.) Desf.
nat
K
O
M
H
Wagner et al. 1990:1586 (K, O, M, H); Herbst & Clayton 1998:34 (KEY)
Polypogon viridis (Gouan) Breistr.
nat
K
H
Wagner et al. 1990:1494 (K, H [as Agrostis semiverticillata]); Herbst &
Clayton 1998:18, 33, 34 (NOM:Syn. A. semiverticillata, KEY)
O
M
H
O
Wagner et al. 1990:1584 (O, M, H)
Herbst & Clayton 1998:34 (NEWNAT/O, KEY) [Note: collected once at
Nu‘uanu Pali in 1916)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 223
Scientific Name
Monocot
Poaceae
Status
Rytidosperma biannulare (Zotov) Connor & Edgar
nat
Rytidosperma caespitosum (Gaudich.) Connor & Edgar
Kl Le Ni K
O Mo L
M Ka H
M
Wagner et al. 1990:1522 (Mo? [reported by Whitney in 1939 as introduced in
1903], EM? [adventive] [as Danthonia semiannularis], DESCR); Herbst &
Clayton 1998:34 (NOM:Syn. D. semiannularis = Rytidosperma semiannulare,
KEY); Starr et al. 2004:27 (EM, WM [as R. semiannulare]); Snow 2008:40
(NEWNAT/Mo, M? [voucher not seen] [as R. biannulare], DESCR); Snow &
Lau 2010:56 (native range correction for R. biannulare); Darbyshire et al.
2010:665 (NOM: R. semiannulare misapplied)
nat
M
Darbyshire et al. 2010:667 (NEWNAT/EM)
Rytidosperma penicillatum (Labill.) Connor & Edgar
nat
M
Saccharum spontaneum L.
nat
Sacciolepis indica (L.) Chase
nat
Schedonnardus paniculatus (Nutt.) Trel.
Mo
Citation
O Mo
K
?
Schedonorus arundinaceus (Schreb.) Dumort.
nat
Schedonorus pratensis (Huds.) P.Beauv.
nat
Schizachyrium condensatum (Kunth) Nees
nat
H
O Mo L
Wagner et al. 1990:1589 (O, Mo)
M
H
O?
K
Wagner et al. 1990:1522 (H [as Danthonia pilosa]); Herbst & Clayton
1998:34 (NOM:Syn. D. pilosa = Rytidosperma pilosum, KEY); Herbst &
Wagner 1999:29 (EM [as R. pilosum]); Darbyshire et al. 2010:670 (NOM: R.
pilosum misapplied)
Wagner et al. 1990:1589 (K, O, Mo, M, H); Herbarium Pacificum Staff
1999:8 (L)
Wagner et al. 1990:1483 (O? [single coll. in 1939 at Poamoho experimental
station], DESCR)
O
M
H?
M
K
Wagner et al. 1990:1547 (H? [as Festuca arundinacea; adventive], DESCR);
Wagner et al. 1997:60 (NEWNAT/K); Oppenheimer 2003:21 (EM); Herbst et
al. 2004:10 (O); Snow 2008:40 (NOM:Syn. F. arundinacea, DESCR);
Catalán et al. 2009:57 (KEY [as F. arundinacea])
Snow 2008:40 (NEWNAT/EM, DESCR); Catalán et al. 2009:57 (KEY [as
Festuca pratensis])
H
Wagner et al. 1990:1590 (H); Lorence et al. 1995:47 (K); Herbst & Clayton
1998:18 (remove K, spm. reidentified as Andropogon glomeratus); Herbst et
al. 2004:10 (O); Snow & Lau 2010:48 (remove O, spm. reidentified as A.
glomeratus var. pumilus); Frohlich & Lau 2012:45 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 224
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Schizachyrium scoparium (Michx.) Nash
nat
K
O
M?
Schizostachyum glaucifolium (Rupr.) Munro
pol?
K
O Mo
M
H
Wagner et al. 1990:1591 (K, O, Mo, M, H); Staples & Herbst 2005:742
(KEY), 751 (DESCR)
Setaria palmifolia (J.König) Stapf
nat
K
O Mo L
M
H
Wagner et al. 1990:1592 (O, L, M, H); Staples et al. 2003:18 (K); Staples &
Herbst 2005:742 (KEY), 748 (DESCR); Oppenheimer 2006:13 (Mo)
Setaria parviflora (Poir.) Kerguélen
nat
K
O Mo L
M
H
Wagner et al. 1990:1592 (K, O, Mo, L, M, H [as Setaria gracilis]); Herbst &
Clayton 1998:35 (NOM:Syn. S. gracilis); Imada et al. 2000:15 (K, earlier
coll.)
Setaria sphacelata (Schumach.) M.B.Moss ex Stapf &
C.E.Hubb.
nat
K
O
M
H
Starr et al. 2002:23 (NEWNAT/EM, DESCR); Herbst et al. 2004:10 (K, O, H)
Setaria verticillata (L.) P.Beauv.
nat
K
O Mo L
M Ka
H
Wagner et al. 1990:1593 (Ni, K, O, Mo, L, M, Ka, H); Wood & LeGrande
2006:27 (Le)
Sorghum Xdrummondii (Nees ex Steud.) Millsp. &
Chase
nat
H?
Herbst & Clayton 1998:35 (NEWNAT/O, L, H? [single cultivated coll. in
1924 from Kohala], KEY)
Sorghum arundinaceum (Desv.) Stapf
nat
H
Snow & Lau 2010:56 (NEWNAT/H, DESCR)
Sorghum bicolor (L.) Moench
nat
K
O
M
H
Wagner et al. 1990:1594 (K, O, H); Herbst & Clayton 1998:35 (KEY);
Oppenheimer 2003:22 (WM); Starr et al. 2006:39 (EM)
Sorghum halepense (L.) Pers.
nat
K
O
M
H
Wagner et al. 1990:1594 (K, O, L, M, H [as Sorghum halpense]); Herbst &
Clayton 1998:35 (KEY, spelling change)
H?
Wagner et al. 1990:1483 (H? [collected twice, most recently in 1936 from
Kapäpala pasture], DESCR)
Sphenopholis obtusata (Michx.) Scribn.
Kl Le Ni
O
L
L
?
Lorence et al. 1995:47 (NEWNAT/K, DESCR); Herbst & Clayton 1998:35
(O, EM? [single coll. in 1939 from Haleakalä experimental station]); Snow &
Lau 2010:49
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 225
Scientific Name
Monocot
Poaceae
Status
Kl Le Ni K
O Mo L
Sporobolus africanus (Poir.) Robyns & Tournay
nat
K
O Mo L
Sporobolus diandrus (Retz.) P.Beauv.
nat
K
O
Sporobolus elongatus R.Br.
nat
Sporobolus indicus (L.) R.Br.
nat
Sporobolus piliferus (Trin.) Kunth
nat
Sporobolus pyramidatus (Lam.) Hitchc.
nat
Sporobolus virginicus (L.) Kunth
ind
Stenotaphrum secundatum (Walter) Kuntze
Themeda villosa (Poir) A.Camus
Thysanolaena latifolia (Roxb. ex Hornem.) Honda
K
M Ka
Wagner et al. 1990:1596 (K, O, Mo, L, M, H); Herbst & Clayton 1998:36
(Ka, KEY, NOM: Sporobolus indicus misapplied, in part)
L
H
Wagner et al. 1990:1596 (O, L, H [as Sporobolus diander]); Lorence & Flynn
1997:11 (K); Herbst & Clayton 1998:36 (KEY); Peterson et al. 2001:206
(epithet spelling change)
O
L
H
Herbst & Clayton 1998:36 (NEWNAT/L [as Sporobolus elongatus], KEY;
NEWNAT/H [as S. pyramidalis], KEY); Snow 2008:41 (O [NOM: S.
pyramidatus misapplied, in part], H [NOM: S. pyramidalis misapplied])
O
L
H
Wagner et al. 1990:1597 (K, O, L, M, H); Warren 1993 (Ka); Herbst &
Clayton 1998:36 (remove M, Ka, spms. reidentified as Sporobolus africanus,
KEY)
O
Herbst & Clayton 1998:35, 36 (NEWNAT/H, KEY); Snow 2008:41 (O);
Snow & Lau 2010:56 (remove H, spm. reidentified as Sporobolus
pyramidatus)
O Mo
K
nat
nat
Ni
Citation
H
K
?
M Ka H
Ka
H
Wagner et al. 1990:1596 (O? [adventive]); Wagner & Herbst 1995:24
(NEWNAT/O, DESCR); Herbst & Clayton 1998:35 (KEY); Starr et al.
2006:40 (Mo); Wood 2006:18 (K); Snow & Lau 2010:56 (H, NOM:
Sporobolus piliferus misapplied, in part); Starr & Starr 2011:31 (Ka)
O Mo L
M Ka
H
Wagner et al. 1990:1597 (Ni, K, O, Mo, L, M, Ka, H); Herbst & Clayton
1998:35 (KEY)
K
O Mo L
M
H
Wagner et al. 1990:1598 (K, O, Mo, L, H); Oppenheimer & Bartlett 2000:7
(EM); Oppenheimer & Bartlett 2002:11 (WM); Staples & Herbst 2005:741
(KEY), 748 (DESCR)
K
O
H
Wagner et al. 1990:1599 (K, O, H)
O?
Snow & Lau 2010:56 (O? [perhaps planted and spreading vegetatively on
Lä‘ie Trail], NOM:Syn. Thysanolaena maxima)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 226
Scientific Name
Monocot
Poaceae
Status
Tragus berteronianus Schult.
nat
Trisetum flavescens (L.) P.Beauv.
Kl Le Ni K
O Mo L
M Ka H
M Ka
Wagner et al. 1990:1601 (Mo, M, Ka); Snow & Davidse 2011:20 (O)
nat
M
Herbst & Clayton 1998:36 (NEWNAT/EM)
Trisetum glomeratum (Kunth) Trin.
end
M
Trisetum inaequale Whitney
end
Triticum aestivum L.
nat
Urochloa brizantha (Hochst. ex A.Rich.) R.D.Webster
nat
Urochloa decumbens (Stapf) R.D.Webster
nat
K
Mo?
?
K
O? Mo?
M?
K
O Mo L
M Ka
Urochloa distachya (L.) T.Q.Nguyen
Urochloa maxima (Jacq.) R.D.Webster
nat
O Mo
Citation
K
L
M
O Mo L
M
H
Wagner et al. 1990:1602 (L, M); Herbst & Clayton 1998:37 (NOM: Trisetum
glomeratum misapplied, in part); Oppenheimer 2008:33 (REDISCOVER/L)
H
M
Ni
Wagner et al. 1990:1602 (L, M, H); Herbst & Clayton 1998:37 (remove L, all
spms. reidentified as Trisetum inaequale)
Wagner et al. 1990:1484 (K?, O?, Mo?, L?, M?, H? [cultivated and escaping],
DESCR); Wagner & Herbst 2003:57 (NEWNAT/K, O, Mo, L, M, H)
Herbst & Clayton 1998:19 (KEY [as Brachiaria brizantha]); Starr et al.
2006:39 (NEWNAT/Ka, DESCR); Oppenheimer 2008:31 (EM); Snow & Lau
2010:49 (remove Ka [spm. reidentified as B. decumbens], retain separation of
Brachiaria from Urochloa); Zuloaga et al. 2003:630/Clayton & Snow
2010:60/Wagner et al. 2012:88 (NOM:Syn. B. brizantha)
Ka
H
Snow & Lau 2010:49 ([as Brachiaria decumbens] NEWNAT/K, Mo? [only
coll. in 1993 as vigorous contaminant in Hawai‘i Plant Materials Center field
in Ho‘olehua], Ka, H, DESCR, NOM: B. brizantha misapplied, in part; retain
separation of Brachiaria from Urochloa); Zuloaga et al. 2003:630/Clayton &
Snow 2010:60/Wagner et al. 2012:89 (NOM:Syn. B. decumbens)
Wagner et al. 1990:1503 (K? [as Brachiaria distachya; single coll. in 1946],
O?, Mo?, M? [as B. subquadripara; escaping and perhaps naturalized]);
Herbst & Clayton 1998:20 (KEY); Lorence et al. 1995:44 (NEWNAT/K [as
B. subquadripara); Snow & Lau 2010:49 (retain separation of Brachiaria from
Urochloa); Zuloaga et al. 2003:631/Wagner & Herbst 2003:58/Clayton &
Snow 2010:60 (NOM:Syn. B. distachya, B. subquadripara)
H
Wagner et al. 1990:1569 (Ni, K, O, Mo, L, M, Ka, H [as Panicum
maximum]); Zuloaga et al. 2003:632/Wagner et al. 2012:97 (NOM:Syn. P.
maximum)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 227
Scientific Name
Monocot
Poaceae
Status
Urochloa mollis (Sw.) Morrone & Zuloaga
nat
Urochloa mutica (Forssk.) T.Q.Nguyen
nat
Urochloa plantaginea (Link) R.D.Webster
Kl Le Ni K
O Mo L
M Ka H
O
K
Citation
Staples et al. 2002:13 (NEWNAT/O [as Brachiaria mollis], DESCR); Snow
& Lau 2010:49 (retain separation of Brachiaria from Urochloa); Zuloaga et
al. 2003:632/Wagner & Herbst 2003:58/Clayton & Snow 2010:61
(NOM:Syn. B. mollis)
O Mo L
M
H
Wagner et al. 1990:1504 (K, O, L, M, H [as Brachiaria mutica]); Herbst &
Clayton 1998:20 (KEY); Oppenheimer 2003:20 (Mo); Snow & Lau 2010:49
(retain separation of Brachiaria from Urochloa); Zuloaga et al.
2003:632/Wagner & Herbst 2003:58/Clayton & Snow 2010:61 (NOM:Syn.
B. mutica)
nat
O Mo
M
H
Wagner & Herbst 1995:23 (NEWNAT/O, Mo [as Brachiaria plantaginea],
DESCR); Herbst & Clayton 1998:20 (KEY); Oppenheimer & Bartlett
2002:10 (WM); Starr et al. 2004:26 (EM); Snow & Lau 2010:49 (H, retain
separation of Brachiaria from Urochloa); Zuloaga et al. 2003:633/Wagner &
Herbst 2003:58/Clayton & Snow 2010:61 (NOM:Syn. B. plantaginea)
Urochloa reptans (L.) Stapf
nat
O
Zizania latifolia (Griseb.) Turcz. ex Stapf
nat
K
Zoysia matrella (L.) Merr. var. pacifica Goudsw.
nat
K
Eichhornia crassipes (Mart.) Solms
nat
K
O
Monochoria vaginalis (Burm.f.) C.Presl
nat
K
O Mo
Herbst & Clayton 1998:20 (NEWNAT/O [as Brachiaria reptans], KEY);
Snow & Lau 2010:49 (retain separation of Brachiaria from Urochloa);
Zuloaga et al. 2003:633/Wagner & Herbst 2003:58/Clayton & Snow 2010:61
(NOM:Syn. B. reptans)
O?
H?
Wagner et al. 1990:1484 (O?, H? [cultivated and at least persisting in wetland
habitats], DESCR); Lorence & Flynn 1999:6 (NEWNAT/K)
Flynn & Lorence 1998:5 (NEWNAT/K); Staples & Herbst 2005:741 (KEY),
750 (DESCR)
Pontederiaceae
M
H
Wagner et al. 1990:1604 (K, O, M, H); Staples & Herbst 2005:751 (DESCR)
H
Wagner et al. 1990:1606 (K, O); Wagner & Herbst 1995:23 (H); Staples &
Herbst 2005:752 (DESCR); Oppenheimer 2011:9 (Mo)
Potamogetonaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 228
Scientific Name
Monocot
Potamogetonaceae
Status
Kl Le Ni K
K
O Mo L
Potamogeton foliosus Raf.
ind?
Potamogeton nodosus Poir.
ind?
Potamogeton tricarinatus A.Benn.
nat
Stuckenia pectinata (L.) Börner
nat
Ni
K
ind
Ni
K
O Mo L
end
K
pol
K
M Ka H
H
Citation
O
M
Wagner et al. 1990:1607 (K, O, M, H); Herbst 1998:4 (DESCR)
O
M
Wagner et al. 1990:1607 (O, M); Herbst 1998:4 (DESCR)
M
Wagner et al. 2012:102 (NEWNAT/EM)
H
Wagner et al. 1990:1608 (Ni, K [as Potamogeton pectinatus]); Wagner &
Herbst 1995:25 (H); Les & Haynes 1997:390/Herbst 1998:4
(COMBNOV:Syn: P. pectinatus = Coleogeton pectinatus); Haynes et al.
1998:241/Herbst & Wagner 1999:30 (NOM:Syn. C. pectinatus)
M
H
Wagner et al. 1990:1609 (Ni, K, O, Mo, L, M, H)
O Mo L
M
H
Wagner et al. 1990:1610 (K, O, Mo, L, M, H)
O Mo
M
H
Wagner et al. 1990:1612 (K, O, Mo, M, H); Staples & Herbst 2005:756
(DESCR)
Ruppiaceae
Ruppia maritima L.
Smilacaceae
Smilax melastomifolia Sm.
Taccaceae
Tacca leontopetaloides (L.) Kuntze
Typhaceae
Typha domingensis Pers.
nat
O
Typha latifolia L.
nat
K
Xyris complanata R.Br.
nat
K
Xyris platylepis Chapm.
nat
Wagner et al. 1990:1614 (O)
O
H
Wagner et al. 1990:1614 (K, O, H? [unvouchered]); Staples et al. 2003:19 (H)
H
Wagner et al. 1990:1615 (K, H)
H
Wagner et al. 1990:1615 (H)
Xyridaceae
Zingiberaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 229
Scientific Name
Monocot
Zingiberaceae
Status
Kl Le Ni K
O Mo L
M Ka H
O
Citation
Alpinia mutica Roxb.
nat
Wagner et al. 1990:1617 (O); Staples & Herbst 2005:759, 760 (KEY), 760
(DESCR)
Alpinia purpurata (Vieill.) K.Schum.
nat
K
Mo
Alpinia zerumbet (Pers.) B.L.Burtt & R.M.Sm.
nat
K
Mo
M
Curcuma longa L.
pol
K
Mo
M
Etlingera cevuga (Seem.) R.M.Sm.
nat
O
Hedychium coronarium J.König
nat
O Mo L
M
H
Wagner et al. 1990:1622 (O, Mo, L, M, H); Staples & Herbst 2005:758, 765
(KEY), 765 (DESCR)
Hedychium flavescens Carey ex Roscoe
nat
K
O Mo L
M
H
Wagner et al. 1990:1622 (K, O, Mo, L, M, H); Staples & Herbst 2005:758,
765 (KEY), 766 (DESCR)
Hedychium gardnerianum Sheppard ex Ker Gawl.
nat
K
O
L
M
H
Wagner et al. 1990:1623 (K, L, EM, H); Staples & Herbst 2005:758, 765
(KEY), 766 (DESCR); Staples et al. 2006:9 (O)
Zingiber zerumbet (L.) Sm.
pol
K
O Mo L
M
H
Wagner et al. 1990:1623 (K, O, Mo, L, M); Oppenheimer 2003:26 (H);
Staples & Herbst 2005:758, 768 (KEY), 769 (DESCR)
Asplenium Xsphenocookii W.H.Wagner
end
K
Asplenium Xflagrum W.H.Wagner & D.D.Palmer
end
K
Wagner et al. 1990:1618 (K, Mo); Staples & Herbst 2005:759 (KEY), 761
(DESCR)
Wagner et al. 1990:1617 [persisting after cultivation], DESCR; Flynn &
Lorence 2002:16 (NEWNAT/K, DESCR); Staples & Herbst 2005:759, 760
(KEY), 761 (DESCR); Oppenheimer 2008:35 (WM); Oppenheimer 2010:38
(Mo)
H
Wagner et al. 1990:1620 (Mo, M, H); Staples & Herbst 2005:758, 762
(KEY), 763 (DESCR); Wood 2012:100 (K)
Wagner et al. 1990:1621 (O)
Pteridophyte
Aspleniaceae
Palmer 2003:76 (K [as Asplenium polyodon x A. sphenotomum])
M
Palmer 2003:76 (K [as Asplenium hobdyi x A. normale]); Oppenheimer
2008:23 (EM)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 230
Scientific Name
Pteridophyte
Aspleniaceae
Status
Kl Le Ni K
O Mo L
M Ka H
K
Citation
Asplenium Xkokeense W.H.Wagner
end
Palmer 2003:76 (K [as Asplenium aethiopicum x A. polyodon])
Asplenium Xwaikamoi W.H.Wagner
end
Asplenium Xjoellaui N.Snow
end
Asplenium acuminatum Hook. & Arn.
end
K
O Mo L
M
H
Palmer 2003:55 (K, O, Mo, L, M, H)
Asplenium adiantum-nigrum L.
ind
K
O Mo L
M
H
Palmer 2003:55 (K, O, Mo, L, M, H)
Asplenium aethiopicum (Burm.f.) Bech.
ind
K
O Mo L
M
H
Palmer 2003:55 (K, O, Mo, L, M, H)
Asplenium contiguum Kaulf. var. contiguum
end
K
O Mo L
M
H
Palmer 2003:58 (K, O, Mo, L, M, H)
Asplenium contiguum Kaulf. var. hirtulum C.Chr.
end
K
Asplenium dielerectum Viane
end
K
Asplenium dielfalcatum Viane
end
Asplenium diellaciniatum Viane
end
K
Palmer 2003:117 (syn. of Diellia erecta [= Asplenium dielerectum]; Wagner
et al. 2012:103 (RESURRECT/K, NOM:Syn. Diellia laciniata [D. Lorence, in
prep.)
Asplenium dielmannii Viane
end
K
Palmer 2003:120 (K [as Diellia mannii]); Aguraiuja & Wood 2003:134
(REDISCOVER/K); Wood 2007:13 (REDISCOVER/K); Snow et al. 2011:12
(NOM:Syn. D. mannii)
Asplenium dielpallidum N.Snow
end
K
Palmer 2003:120 (K [as Diellia pallida]); Snow et al. 2011:12 (NOM:Syn. D.
pallida)
M
Palmer 2003:76 (EM [as Asplenium acuminatum x A. aethiopicum])
O
Palmer 2003:122 (O [as Diellia falcata x D. unisora = D. xlauii]); Snow et al.
2011:12 (COMBNOV [as Asplenium xlaui], Syn. Diellia xlauii); Snow
2011:161 (NOM: Asplenium xlaui, illegal name)
M
O Mo L
O
M
Palmer 2003:58 (K, M)
H
Palmer 2003:117 (K, O, Mo, L, M, H [as Diellia erecta]); Wood 2006:16
(REDISCOVER/K [f. alexandri]); Snow et al. 2011:12 (NOM:Syn. D. erecta;
status of fma. alexandri & pumila still under study)
Palmer 2003:119 (O [as Diellia falcata]); Snow et al. 2011:12 (NOM:Syn. D.
falcata)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 231
Scientific Name
Pteridophyte
Aspleniaceae
Status
Kl Le Ni K
Asplenium excisum C.Presl
ind
Asplenium haleakalense W.H.Wagner
end
Asplenium hobdyi W.H.Wagner
end
K
Asplenium horridum Kaulf. var. glabratum (W.J.Rob.)
D.D.Palmer
end
Asplenium horridum Kaulf. var. horridum
M Ka H
H
Palmer 2003:58 (K, O, Mo, L, M, H)
M
H
Palmer 2003:60 (EM); Kennedy et al. 2010:20 (H)
Mo
M
H
Palmer 2003:60 (K, Mo, M, H)
K
O Mo L
M
H
Palmer 2003:62 (K, O, Mo, L, M, H) [Note: Wagner et al. 2005– call this
Asplenium glabratum W.J.Rob. [Syn. A. horridum var. glabratum]
ind
K
O Mo L
M
H
Palmer 2003:62 (K, O, Mo, L, M, H) [Note: Wagner et al. 2005– call this
Asplenium caudatum G.Forst. [Syn. A. horridum]
Asplenium insiticium Brack.
ind
K
O Mo
M
H
Palmer 2003:63 (K, O, Mo, L, M, H)
Asplenium kaulfussii Schltdl. f. bipinnatum (Hillebr.)
D.D.Palmer
end
O
Palmer 2003:65 (O)
Asplenium kaulfussii Schltdl. f. dareoides (Hillebr.)
W.H.Wagner
end
O
Palmer 2003:65 (O)
Asplenium kaulfussii Schltdl. f. gemmiparum (Hillebr.)
D.D.Palmer
end
K
O
L
M
Asplenium kaulfussii Schltdl. f. kaulfussii
end
K
O Mo L
M
Asplenium leucostegioides Baker
end
Asplenium lobulatum Mett.
ind
K
O Mo L
M
H
Palmer 2003:66 (K, O, Mo, L, M, H)
end?
K
O Mo L
M
H
Palmer 2003:66 (K, O, Mo, L, M, H) [Note: Wagner et al. 2005– calls this
"end']
M
H
Palmer 2003:68 (M, H)
Asplenium monanthes L.
O Mo L
Citation
M
Asplenium macraei Hook. & Grev.
K
O Mo L
Palmer 2003:65 (K, O, L, M)
H
M
ind
Palmer 2003:66 (K, O, Mo, L, M, H)
Palmer 2003:119 (EM [as Diellia leucostegioides]); Snow et al. 2011:12
(NOM:Syn. D. leucostegioides)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 232
Scientific Name
Pteridophyte
Aspleniaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Asplenium nidus L.
ind
K
O Mo L
M
H
Palmer 2003:70 (K, O, Mo, L, M, H); Staples & Herbst 2005:28 (DESCR)
Asplenium normale D.Don
ind
K
O Mo L
M
H
Palmer 2003:70 (K, O, Mo, L, M, H)
Asplenium peruvianum Desv. var. insulare
(C.V.Morton) D.D.Palmer
end
M
H
Palmer 2003:70 (M, H)
Asplenium polyodon G.Forst.
ind
K
M
H
Palmer 2003:71 (K, O, Mo, L, M, H)
Asplenium schizophyllum C.Chr.
end
K
H
Palmer 2003:73 (K, H)
Asplenium sphenotomum Hillebr.
end
K
M
H
Palmer 2003:73 (K, O, Mo, L, M, H)
Asplenium trichomanes L. ssp. densum (Brack.)
W.H.Wagner
end
M
H
Palmer 2003:74 (EM, H)
Asplenium unilaterale Lam.
ind
M
H
Palmer 2003:74 (K, O, Mo, L, M, H)
Asplenium unisorum (W.H.Wagner) Viane
end
K
O Mo L
O Mo L
O Mo L
O
Palmer 2003:122 (O [as Diellia unisora]); Snow et al. 2011:12 (NOM:Syn. D.
unisora)
Athyriaceae
Athyrium microphyllum (Sm.) Alston
end
K
Cystopteris douglasii Hook.
end
Cystopteris sandwicensis Brack.
end
K
Deparia cataracticola M.Kato
end
K
Deparia fenzliana (Luerss.) M.Kato
end
K
Deparia kaalaana (Copel.) M.Kato
end
K
O Mo L
O
L
M
H
Palmer 2003:76 (K, O, Mo, L, M, H)
M
H
Palmer 2003:108 (EM, H)
M
Palmer 2003:108 (K, O, L, WM)
Kato 2001:3 (SPNOV/K, DESCR, KEY)
O Mo L
M
H
Kato 2001:7 (KEY); Palmer 2003:109 (K, O, Mo, L, M, H)
M
H
Kato 2001:7 (KEY); Palmer 2003:109 (K, EM, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 233
Scientific Name
Pteridophyte
Athyriaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Deparia marginalis (Hillebr.) M.Kato
end
K
Mo L
M
H
Kato 2001:7 (KEY); Palmer 2003:111 (K, Mo, L, M, H)
Deparia petersenii (Kunze) M.Kato
nat
K
O Mo L
M
H
Kato 2001:7 (KEY); Palmer 2003:111 (K, O, Mo, L, M, H)
Deparia prolifera (Kaulf.) Hook. & Grev.
end
K
O Mo L
M
H
Kato 2001:7 (KEY); Palmer 2003:111 (K, O, Mo, L, M, H)
Diplazium arnottii Brack.
end
K
O Mo L
M
H
Palmer 2003:123 (K, O, Mo, L, M, H)
Diplazium esculentum (Retz.) Sw.
nat
K
O Mo L
M
H
Palmer 2003:125 (K, O, L, M, H); Staples & Herbst 2005:29 (DESCR);
Oppenheimer 2007:21 (Mo)
Diplazium molokaiense W.J.Rob.
end
K
O Mo L
M
Diplazium sandwichianum (C.Presl) Diels
end
K
O Mo L
M
H
Palmer 2003:125 (K, O, Mo, L, M, H)
nat
K
O Mo L
M
H
Palmer 2003:78 (K, O, Mo, L, M, H); Wilson 2003:5 (H); Staples & Herbst
2005:30 (DESCR)
Blechnum appendiculatum Willd.
nat
K
O Mo L
M
H
Palmer 2003:79 (K, O, Mo, L, M, H); Staples & Herbst 2005:31 (DESCR [as
Blechnum occidentale])
Blechnum orientale L.
nat
Doodia kunthiana Gaudich.
end
K
O Mo L
M
H
Palmer 2003:129 (K, O, Mo, L, M, H); Christenhusz et al. 2011:20
(NOMNOV: proposed transfer to Blechnum as B. norfolkense); Christenhusz
2011:1772 (proposal to conserve NOMNOV B. hawaiianum over B.
norfolkense)
Doodia lyonii O.Deg.
end
K
O
M
H
Palmer 2003:131 (K, O, M, H); Christenhusz et al. 2011:20 (COMBNOV:
proposed transfer to Blechnum as B. lyonii)
Sadleria cyatheoides Kaulf.
end
K
O Mo L
M
H
Palmer 2003:231 (K, O, Mo, L, M, H); Staples & Herbst 2005:31 (DESCR)
Palmer 2003:125 (K, O, Mo, L, M); Wood 2007:14 (REDISCOVER/WM)
Azollaceae
Azolla filiculoides Lam.
Blechnaceae
O
Lau & Frohlich 2012:10 (NEWNAT/O, DESCR)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 234
Scientific Name
Pteridophyte
Blechnaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Sadleria pallida Hook. & Arn.
end
K
O Mo L
M
H
Palmer 2003:233 (K, O, Mo, L, M, H)
Sadleria souleyetiana (Gaudich.) T.Moore
end
K
O Mo L
M
H
Palmer 2003:233 (K, O, Mo, L, M, H)
Sadleria squarrosa (Gaudich.) T.Moore
end
K
O Mo L
M
H
Palmer 2003:234 (K, O, Mo, L, M, H)
Sadleria unisora (Baker) W.J.Rob.
end
K
Palmer 2003:234 (K)
Sadleria wagneriana D.D.Palmer & Flynn
end
K
Palmer 2003:234 (K)
end
K
O Mo L
M
H
Palmer 2003:238 (K, O, Mo, L, M, H [as Sceptridium subbifoliatum]); Snow
et al. 2011:14 (NOM:Syn. S. subbifoliatum) [Note: many workers continue to
recognize Sceptridium as a good genus]
nat
K
O
M
H
Palmer 2003:243 (K, O, M, H); Staples & Herbst 2005:32 (DESCR [as
Cyathea cooperi]); Oppenheimer 2008:26 (L)
Botrychiaceae
Botrychium subbifoliatum Brack.
Cyatheaceae
Sphaeropteris cooperi (Hook. ex F.Muell.) R.M.Tryon
L
Davalliaceae
Davallia fejeensis Hook.
nat
O
Staples & Herbst 2005:33 (DESCR); Frohlich & Lau 2008:5 (NEWNAT/O,
DESCR)
Dennstaedtiaceae
Hypolepis hawaiiensis Brownsey var. hawaiiensis
end
K
O Mo L
Hypolepis hawaiiensis Brownsey var. mauiensis
(Hillebr.) D.D.Palmer
end
Microlepia Xadulterina W.H.Wagner
end
K
O
Microlepia speluncae (L.) T.Moore
ind
K
O Mo
M
H
M
M
Palmer 2003:168 (K, O, Mo, L, M, H)
Palmer 2003:168 (WM); Oppenheimer 2008:27 (EM)
H
Palmer 2003:186 (K, O, H [as Microlepia speluncae x M. strigosa])
H
Palmer 2003:184 (K, O, H); Oppenheimer 2004:11 (WM); Oppenheimer
2008:28 (Mo)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 235
Scientific Name
Pteridophyte
Dennstaedtiaceae
Status
Kl Le Ni K
Microlepia strigosa (Thunb.) C.Presl var. mauiensis
(W.H.Wagner) D.D.Palmer
end
Microlepia strigosa (Thunb.) C.Presl var. strigosa
ind
K
Pteridium aquilinum (L.) Kuhn ssp. decompositum
(Gaudich.) Lamoureux ex J.A.Thomson
end
K
O Mo L
M Ka H
Citation
M
H
Palmer 2003:186 (M, H)
O Mo L
M
H
Palmer 2003:186 (K, O, Mo, L, M, H); Staples & Herbst 2005:34 (DESCR
[as Microlepia strigosa])
O Mo L
M
H
Palmer 2003:224 (K, O, Mo, L, M, H [as var. decompositum]); Thomson
2004:800 (NOM:Syn. Pteridium aquilinum var. decompositum)
Dicksoniaceae
Cibotium Xheleniae D.D.Palmer
end
O
Palmer 2003:98 (O [as Cibotium chamissoi x C. menziesii])
Cibotium chamissoi Kaulf.
end
O Mo L
M
H
Palmer 2003:95 (O, Mo, L, M, H); Staples & Herbst 2005:36 (DESCR)
Cibotium glaucum (Sm.) Hook. & Arn.
end
K
O Mo L
M
H
Palmer 2003:95 (K, O, Mo, L, M, H); Staples & Herbst 2005:36 (DESCR)
Cibotium menziesii Hook.
end
K
O Mo L
M
H
Palmer 2003:96 (K, O, Mo, L, M, H); Staples & Herbst 2005:36 (DESCR)
Cibotium nealiae O.Deg.
end
K
Dicksonia fibrosa Colenso
nat
Palmer 2003:96 (K)
H
Lorence & Flynn 2006:1 (NEWNAT/H, DESCR, KEY)
Dryopteridaceae
Arachniodes insularis W.H.Wagner
end
K
O Mo
M
H
Palmer 2003:49 (K, O, Mo, M, H)
Ctenitis latifrons (Brack.) Copel.
end
K
O Mo L
M
H
Palmer 2003:100 (K, O, Mo, L, M, H)
Ctenitis squamigera (Hook. & Arn.) Copel.
end
K
O Mo L
M
Cyrtomium caryotideum (Wall.) C.Presl
ind
K
O Mo L
M
H
Palmer 2003:104 (K, O, Mo, L, M, H)
Cyrtomium falcatum (L.f.) C.Presl
nat
K
O Mo L
M
H
Palmer 2003:106 (K, O, Mo, L, M, H); Staples & Herbst 2005:37 (DESCR)
Dryopteris crinalis (Hook. & Arn.) C.Chr. var. crinalis
end
K
O Mo L
M
H
Palmer 2003:139 (K, O, Mo, L, M, H)
Palmer 2003:100 (K, O, Mo, L, M); Wood 2012:94 (REDISCOVER/K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 236
Scientific Name
Pteridophyte
Dryopteridaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Dryopteris crinalis (Hook. & Arn.) C.Chr. var.
podosorus (W.H.Wagner & Flynn) D.D.Palmer
end
K
Palmer 2003:139 (K)
Dryopteris fusco-atra (Hillebr.) W.J.Rob. var. fusco-atra
end
K
Dryopteris fusco-atra (Hillebr.) W.J.Rob. var.
lamoureuxii Fraser-Jenk.
end
Dryopteris glabra (Brack.) Kuntze var. alboviridis
(W.H.Wagner) D.D.Palmer
end
K
Palmer 2003:141 (K)
Dryopteris glabra (Brack.) Kuntze var. flynnii
D.D.Palmer
end
K
Palmer 2003:143 (K)
Dryopteris glabra (Brack.) Kuntze var. glabra
end
K
Dryopteris glabra (Brack.) Kuntze var. hobdyana
(W.H.Wagner) D.D.Palmer
end
Dryopteris glabra (Brack.) Kuntze var. nuda (Underw.)
Fraser-Jenk.
end
K
Dryopteris glabra (Brack.) Kuntze var. pusilla (Hillebr.)
Fraser-Jenk.
end
K
Dryopteris glabra (Brack.) Kuntze var. soripes (Hillebr.)
Herat ex Fraser-Jenk.
end
K
O Mo
M
H
Palmer 2003:146 (K, O, Mo, M, H)
Dryopteris hawaiiensis (Hillebr.) W.J.Rob.
end
K
O Mo
M
H
Palmer 2003:146 (K, O, Mo, M, H)
Dryopteris mauiensis C.Chr.
end
K
O Mo L
M
H
Palmer 2003:146 (K, O, Mo, L, M, H)
Dryopteris rubiginosa (Brack.) Kuntze
end
K
O Mo L
M
H
Palmer 2003:194 (K, O, Mo, L, M, H [as Nothoperanema rubiginosa]); Smith
et al. 2006:717/Snow et al. 2011:13 (NOM:Syn. Nothoperanema rubiginosa)
O Mo L
M
H
M
O Mo L
O Mo
M
Palmer 2003:140 (K, O, Mo, L, M, H)
Palmer 2003:140 (EM); Oppenheimer 2011:7 (WM)
H
Palmer 2003:143 (K, O, Mo, L, M, H)
M
Palmer 2003:143 (EM)
M
Palmer 2003:144 (K, O, Mo, M)
Palmer 2003:144 (K)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 237
Scientific Name
Pteridophyte
Dryopteridaceae
Status
Dryopteris sandwicensis (Hook. & Arn.) C.Chr.
end
Dryopteris subbipinnata W.H.Wagner & Hobdy
Kl Le Ni K
K
O Mo L
M Ka H
O Mo L
Citation
M
H
Palmer 2003:147 (K, O, Mo, L, M, H)
end
M
H
Palmer 2003:147 (EM, WM, H)
Dryopteris tetrapinnata W.H.Wagner & Hobdy
end
M
Dryopteris unidentata (Hook. & Arn.) C.Chr. var.
paleacea (Hillebr.) Herat ex Fraser-Jenk.
end
K
O Mo
M
H
Palmer 2003:150 (K, O, Mo, M, H)
Dryopteris unidentata (Hook. & Arn.) C.Chr. var.
unidentata
end
K
O Mo L
M
H
Palmer 2003:151 (K, O, Mo, L, M, H)
Dryopteris wallichiana (Spreng.) Hyl.
ind
K
O Mo
M
H
Palmer 2003:151 (K, O, Mo, M, H)
Polystichum bonseyi W.H.Wagner & Hobdy
end
M
H
Palmer 2003:218 (EM, H)
Polystichum haleakalense Brack.
end
M
H
Palmer 2003:220 (EM, H)
Polystichum hillebrandii Carruth.
end
M
H
Palmer 2003:220 (M, H)
Tectaria gaudichaudii (Mett.) Maxon
end
K
O Mo L
M
H
Palmer 2003:249 (K, O, Mo, L, M, H)
Tectaria incisa Cav.
nat
K
O
M
H
Palmer 2003:249 (K, O, H); Imada 2007:36 (EM)
Elaphoglossum aemulum (Kaulf.) Brack.
end
K
O Mo L
M
H
Palmer 2003:154 (K, O, Mo, L, M, H)
Elaphoglossum alatum Gaudich.
end
Elaphoglossum crassicaule Copel.
end
K
Elaphoglossum crassifolium (Gaudich.) W.R.Anderson
& Crosby
end
K
Palmer 2003:149 (EM)
Elaphoglossaceae
O
Palmer 2003:154 (O)
Palmer 2003:155 (K)
O Mo L
M
H
Palmer 2003:156 (K, O, Mo, L, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 238
Scientific Name
Pteridophyte
Elaphoglossaceae
Status
Kl Le Ni K
Elaphoglossum fauriei Copel.
end
Elaphoglossum paleaceum (Hook. & Grev.) Sledge
ind
Elaphoglossum parvisquameum Skottsb.
end
Elaphoglossum pellucidum Gaudich.
end
K
Elaphoglossum wawrae (Luerss.) C.Chr.
end
Dicranopteris linearis (Burm.f.) Underw. f. emarginata
(Brack.) W.H.Wagner
O Mo L
M Ka H
O Mo
K
Citation
Palmer 2003:156 (O, Mo)
O Mo L
M
H
Palmer 2003:158 (K, O, Mo, M, H); Oppenheimer 2010:35 (L)
Mo L
M
H
Palmer 2003:158 (Mo, L, M, H)
O Mo
M
H
Palmer 2003:158 (K, O, Mo, M, H)
K
O Mo
M
H
Palmer 2003:160 (K, O, Mo, M, H)
end
K
O Mo L
M
H
Palmer 2003:115 (K, O, Mo, L, M, H)
Dicranopteris linearis (Burm.f.) Underw. f. linearis
ind
K
O Mo L
M
H
Palmer 2003:115 (K, O, Mo, L, M, H)
Diplopterygium pinnatum (Kunze) Nakai
end
K
O Mo L
M
H
Palmer 2003:127 (K, O, Mo, L, M, H)
Sticherus owhyhensis (Hook.) Ching
end
K
O Mo L
M
H
Palmer 2003:247 (K, O, Mo, L, M, H)
Gleicheniaceae
Grammitidaceae
Adenophorus Xbishopii (W.H.Wagner) xxxxx
end
Adenophorus Xabbottiae W.H.Wagner
end
Adenophorus Xcarsonii Ranker
end
Adenophorus abietinus (D.C.Eaton) K.A.Wilson
end
K
Adenophorus epigaeus (L.E.Bishop) W.H.Wagner
end
K
Adenophorus haalilioanus (Brack.) K.A.Wilson
end
K
K
O
Palmer 2003:41 (O [as Adenophorus oahuensis x A. pinnatifidus])
O
Palmer 2003:36 (K, O [as Adenophorus hymenophylloides x A.
tamariscinus])
H
O
L
M
Palmer 2003:36 (H [as Adenophorus hymenophylloides x A. tripinnatifidus])
Palmer 2003:33 (K, O, L, EM, WM)
Palmer 2003:33 (K)
O
Palmer 2003:38 (K, O)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 239
Scientific Name
Pteridophyte
Grammitidaceae
Status
Kl Le Ni K
K
O Mo L
M Ka H
O Mo L
M
H
Citation
Adenophorus hymenophylloides (Kaulf.) Hook. & Grev.
end
Adenophorus oahuensis (Copel.) L.E.Bishop
end
Adenophorus periens L.E.Bishop
end
K
O Mo L
M
H
Palmer 2003:39 (K, O, Mo, L, M, H)
Adenophorus pinnatifidus Gaudich. var. pinnatifidus
end
K
O Mo L
M
H
Palmer 2003:41 (K, O, Mo, L, M, H)
Adenophorus pinnatifidus Gaudich. var. rockii (Copel.)
D.D.Palmer
end
K
O Mo
M
Adenophorus tamariscinus (Kaulf.) Hook. & Grev. var.
montanus (Hillebr.) L.E.Bishop
end
Mo
M
H
Palmer 2003:35 (Mo, M, H)
Adenophorus tamariscinus (Kaulf.) Hook. & Grev. var.
tamariscinus
end
K
O Mo L
M
H
Palmer 2003:35 (K, O, Mo, L, M, H)
Adenophorus tenellus (Kaulf.) Ranker
end
K
O Mo L
M
H
Palmer 2003:165 (K, O, Mo, L, M, H [as Grammitis tenella]); Ranker
2008:171/Snow et al. 2011:14 (NOM:Syn. G. tenella)
Adenophorus tripinnatifidus Gaudich.
end
K
O Mo L
M
H
Palmer 2003:35 (K, O, Mo, L, M, H)
Oreogrammitis baldwinii (Baker) Parris
end
K
Oreogrammitis forbesiana (W.H.Wagner) Parris
end
K
Mo
M
Oreogrammitis hookeri (Brack.) Parris
ind
K
O Mo L
M
H
Palmer 2003:165 (K, O, Mo, L, M, H); Parris 2007:260 (NOM:Syn.
Grammitis hookeri)
end
K
O Mo L
M
H
Palmer 2003:81 (K, O, Mo, L, M, H)
O
Palmer 2003:34 (K, O, Mo, L, M, H)
Palmer 2003:39 (O)
Palmer 2003:41 (K, O, Mo, M)
Palmer 2003:163 (K); Parris 2007:256 (NOM:Syn. Grammitis baldwinii)
Palmer 2003:163 (K, Mo, M); Parris 2007:260 (NOM:Syn. Grammitis
forbesiana)
Hymenophyllaceae
Callistopteris baldwinii (D.C.Eaton) Copel.
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 240
Scientific Name
Pteridophyte
Hymenophyllaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Crepidomanes draytonianum (Brack.) Ebihara & K.
Iwats.
end
K
O Mo L
M
H
Palmer 2003:253 (K, O, Mo, L, M, H [as Vandenboschia draytoniana]);
Ebihara et al. 2006:238 (COMBNOV:Syn. V. draytoniana)/Snow et al.
2011:13
Crepidomanes minutum (Blume) K.Iwats.
ind
K
O Mo L
M
H
Palmer 2003:162 (K, O, Mo, L, M, H [as Gonocormus minutus]); Ebihara et
al. 2006:238/Snow et al. 2011:13 (NOM:Syn. G. minutus)
Crepidomanes proliferum (Blume) Bostock
ind
M
H
Palmer 2003:162 (EM, H [as Gonocormus prolifer]); Ebihara et al.
2006:238/Snow et al. 2011:13 (NOM:Syn. G. prolifer)
Hymenophyllum lanceolatum Hook. & Arn.
end
O Mo L
M
H
Palmer 2003:242 (K, O, Mo, L, M, H [as Sphaerocionium lanceolatum]);
Ebihara et al. 2006:230/Snow et al. 2011:13 (NOM:Syn. S. lanceolatum)
Hymenophyllum obtusum Hook. & Arn.
end
O Mo L
M
H
Palmer 2003:243 (O, Mo, L, M, H [as Sphaerocionium obtusum]); Ebihara et
al. 2006:230/Snow et al. 2011:13 (NOM:Syn. S. obtusum)
Hymenophyllum recurvum Gaudich.
end
K
O Mo L
M
H
Palmer 2003:182 (K, O, Mo, L, M, H [as Mecodium recurvum]); Ebihara et
al. 2006:231/Snow et al. 2011:13 (NOM:Syn. M. recurvum)
Vandenboschia cyrtotheca (Hillebr.) Copel.
end
K
O Mo L
M
H
Palmer 2003:251 (K, O, Mo, L, M, H)
Vandenboschia davallioides (Gaudich.) Copel.
end
K
O Mo L
M
H
Palmer 2003:253 (K, O, Mo, L, M, H)
Vandenboschia tubiflora F.S.Wagner
end
K
K
Palmer 2003:253 (K)
Isoetaceae
Isoëtes hawaiiensis W.C.Taylor & W.H.Wagner
end
M
H
Palmer 2003:266 (WM, H)
Lindsaeaceae
Lindsaea ensifolia Sw.
nat
K
O Mo
M
H
Palmer 2003:174 (K, O, M, H); Oppenheimer 2008:29 (Mo)
Lindsaea repens (Bory) Thwaites var. macraeana (Hook.
& Arn.) C.Chr.
end
K
O Mo L
M
H
Palmer 2003:174 (K, O, Mo, L, M, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 241
Scientific Name
Pteridophyte
Lindsaeaceae
Status
Kl Le Ni K
Sphenomeris chinensis (L.) Maxon
ind
K
x_Lindsaeosoria flynnii W.H.Wagner
end
K
O Mo L
M Ka H
O Mo L
M
H
Citation
Palmer 2003:245 (K, O, Mo, L, M, H); Staples & Herbst 2005:35 (DESCR
[as Odontosoria chinensis])
Palmer 2003:174 (K [as Lindsaea ensifolia x Sphenomeris chinensis])
Lycopodiaceae
Huperzia Xkoolauensis (W.H.Wagner) Kartesz
end
O
Huperzia Xsulcinervia (Spring) Trevis.
end
O
Huperzia Xmedeirosii Beitel & W.H.Wagner
end
Huperzia Xcarlquistii Beitel & W.H.Wagner
end
Huperzia Xgillettii Beitel & W.H.Wagner
end
Huperzia erosa Beitel & W.H.Wagner
Palmer 2003:259 (O [as Huperzia nutans x H. phyllantha])
L
H
Palmer 2003:263 (O, L, H [as Huperzia erubescens x H. serrata])
M
Palmer 2003:265 (WM [as Huperzia haleakalae x H. subintegra])
O
M
Palmer 2003:265 (O, WM [as Huperzia erubescens x H. subintegra])
K
O Mo
M
H
Palmer 2003:265 (K, O, Mo, M, H [as Huperzia serrata x H. subintegra])
end
K
O Mo L
M
H
Palmer 2003:261 (K, O, Mo, L, M, H)
Huperzia erubescens (Brack.) Holub
ind
K
O Mo
M
H
Palmer 2003:262 (K, O, Mo, M, H)
Huperzia filiformis (Sw.) Holub
ind
K
O Mo L
M
H
Palmer 2003:265 (O, Mo, L, M, H); Wood 2012:95 (K)
Huperzia haleakalae (Brack.) Holub
ind
Huperzia mannii (Hillebr.) Kartesz & Gandhi
end
K
Huperzia nutans (Brack.) Rothm.
end
K
O
Huperzia phyllantha (Hook. & Arn.) Holub
ind
K
O Mo L
Huperzia serrata (Thunb. ex Murray) Trevis.
ind
K
O Mo L
Huperzia stemmermanniae (A.C.Medeiros,
W.H.Wagner & Hobdy) Kartesz
end
M
M
Palmer 2003:262 (EM)
H
Palmer 2003:256 (K, EM, WM, H)
Palmer 2003:257 (K, O)
M
M
H
Palmer 2003:257 (K, O, Mo, L, M, H)
H
Palmer 2003:262 (K, O, Mo, L, H)
H
Palmer 2003:257 (EM, H)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 242
Scientific Name
Pteridophyte
Lycopodiaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Huperzia subintegra (Hillebr.) Beitel & W.H.Wagner
end
K
O Mo
M
Palmer 2003:263 (K, O, Mo, M)
Lycopodiella cernua (L.) Pic.Serm.
ind
K
O Mo L
M
H
Palmer 2003:269 (K, O, Mo, L, M, H)
Lycopodium venustulum Gaudich. var. venustulum
ind
K
O Mo L
M
H
Palmer 2003:271 (K, O, Mo, L, M, H)
Lycopodium venustulum Gaudich. var. verticale
W.H.Wagner
end
H
Palmer 2003:271 (H)
Marattiaceae
Angiopteris evecta (G.Forst.) Hoffm.
nat
K
O Mo L
M
H
Palmer 2003:48 (O, L, M, H); Staples & Herbst 2005:39 (DESCR); Wood
2006:18 (K); Oppenheimer 2010:37 (Mo)
Marattia douglasii (C.Presl) Baker
end
K
O Mo L
M
H
Palmer 2003:178 (K, O, Mo, L, M, H)
Marsileaceae
Marsilea crenata C.Presl
nat
Marsilea villosa Kaulf.
end
Ni
O
Wilson 2003:5 (NEWNAT/O)
O Mo
Palmer 2003:180 (Ni, O, Mo); Staples & Herbst 2005:40 (DESCR)
Palmer 2003:193 (O [as Nephrolepis exaltata subsp. hawaiiensis x N.
multiflora])
Nephrolepidaceae
Nephrolepis Xmedlerae W.H.Wagner
end
O
Nephrolepis Xcopelandii W.H.Wagner
end
O
M
H
Palmer 2003:193 (O, H [as Nephrolepis cordifolia x N. multiflora]);
Oppenheimer 2004:14 (WM)
Nephrolepis brownii (Desv.) Hovenkamp & Miyam.
nat
K
O Mo L
M
H
Palmer 2003:193 (K, O, Mo, L, M, H [as Nephrolepis multiflora]); Staples &
Herbst 2005:40 (KEY [as N. multiflora]), 41 (DESCR); Wood & LeGrande
2006:24 (Le); Imada 2007:38 (Ni); Hovenkamp & Miyamoto 2005:293
(NOM:Syn. N. multiflora)
Nephrolepis cordifolia (L.) C.Presl
ind
K
O Mo L
M
H
Palmer 2003:189 (K, O, Mo, L, M, H); Staples & Herbst 2005:40 (DESCR,
KEY); Hovenkamp & Miyamoto 2005:294
Le Ni
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 243
Scientific Name
Pteridophyte
Nephrolepidaceae
Status
Nephrolepis exaltata (L.) Schott
ind
Nephrolepis falcata (Cav.) C.Chr. ‘Furcans’
Nephrolepis hirsutula (G.Forst.) C.Presl ‘Superba’
Kl Le Ni K
Ni
O Mo L
M Ka H
Citation
K
O Mo L
M
H
Palmer 2003:190 (Ni, K, O, Mo, L, M, H [as subsp. hawaiiensis]); Staples &
Herbst 2005:40 (KEY), 41 (DESCR); Hovenkamp & Miyamoto 2005:300
(NOM:Syn. Nephrolepis exaltata subsp. hawaiiensis, status change from
"end" to "ind")
nat
K
O Mo L
M
H
Palmer 2003:190 (K, O, Mo, L, M, H); Staples & Herbst 2005:40 (KEY), 41
(DESCR)
nat
K
Ophioglossum nudicaule L.f.
ind
K
O
Ophioglossum pendulum L.
ind
K
O Mo L
Ophioglossum petiolatum Hook.
ind
K
Ophioglossum polyphyllum A.Braun
ind
Lepisorus thunbergianus (Kaulf.) Ching
M
Palmer 2003:190 (K, EM)
Ophioglossaceae
H
Palmer 2003:198 (K, O, H)
M
H
Palmer 2003:196 (O, Mo, EM, WM, H [as Ophioderma pendulum subsp.
pendulum]; K, O, Mo, L, M, H [as Ophioderma pendulum subsp. falcatum);
Smith et al. 2006:709/Wagner et al. 2012:105 (NOM:Syn. Ophioderma
pendulum subsp. falcatum, no infraspecific taxa recognized)
O Mo L
M
H
Palmer 2003:199 (K, O, L, M, H); Oppenheimer 2010:37 (Mo)
K
O Mo L
M Ka
H
Palmer 2003:199 (K, O, Mo, L, M, Ka, H)
ind
K
O Mo L
M
H
Palmer 2003:172 (K, O, Mo, L, M, H)
Microsorum spectrum (Kaulf.) Copel. var.
pentadactylum (Hillebr.) D.D.Palmer
end
K
Microsorum spectrum (Kaulf.) Copel. var. spectrum
end
K
O Mo L
M
H
Palmer 2003:188 (K, O, Mo, L, M, H)
Phlebodium aureum (L.) J.Sm.
nat
K
O Mo L
M
H
Palmer 2003:203 (K, O, Mo, L, M, H); Staples & Herbst 2005:42 (KEY), 44
(DESCR)
Polypodiaceae
M?
Palmer 2003:188 (K, EM?)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 244
Scientific Name
Pteridophyte
Polypodiaceae
Status
Kl Le Ni K
K
O Mo L
M Ka H
O Mo L
M
H
Citation
Phymatosorus grossus (Langsd. & Fisch.) Brownlie
nat
Phymatosorus scolopendria (Burm.f.) Pic.Serm.
nat
Platycerium bifurcatum (Cav.) C.Chr.
nat
O
Platycerium superbum de Jonch. & Hennipman
nat
O
Polypodium pellucidum Kaulf. var. acuminatum
D.D.Palmer
end
K
Polypodium pellucidum Kaulf. var. pellucidum f.
opacum (Hillebr.) D.D.Palmer
end
K
Polypodium pellucidum Kaulf. var. pellucidum f.
pellucidum Kaulf.
end
K
Polypodium pellucidum Kaulf. var. vulcanicum Skottsb.
end
Pyrrosia piloselloides (L.) M.G.Price
nat
Stenogrammitis saffordii (Maxon) Labiak
end
K
O Mo L
M
Psilotum Xintermedium W.H.Wagner
end
K
O
L
M
Psilotum complanatum Sw.
ind
K
O Mo L
M
H
Palmer 2003:272 (K, O, Mo, L, M, H)
Psilotum nudum (L.) P.Beauv.
ind
K
O Mo L
M Ka
H
Palmer 2003:272 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst 2005:46
(DESCR)
M
M
Palmer 2003:205 (K, O, Mo, L, M, H); Staples & Herbst 2005:42 (KEY, as
Microsorum), 43 (DESCR [as M. scolopendria])
Oppenheimer 2006:13 (NEWNAT/WM)
H
Palmer 2003:210 (O, EM, H); Staples & Herbst 2005:42 (KEY), 45 (DESCR)
Palmer 2003:210 (O); Staples & Herbst 2005:42 (KEY), 45 (DESCR)
M
Palmer 2003:216 (K, EM)
O Mo L
M
Palmer 2003:216 (K, O, Mo, L); Oppenheimer 2004:16 (WM)
O Mo L
M
H
Palmer 2003:216 (K, O, Mo, L, M, H)
Mo
M
H
Palmer 2003:216 (Mo, M, H)
O
Lau & Frohlich 2012:21 (NEWNAT/O, DESCR)
H
Palmer 2003:169 (K, O, Mo, L, M, H [as Lellingeria saffordii]); Labiak
2011:147 (NOM:Syn. L. saffordii)
Psilotaceae
Ni
Palmer 2003:272 (K, O, L, M)
Pteridaceae
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 245
Scientific Name
Pteridophyte
Pteridaceae
Status
Adiantum ‘Edwinii'
nat
Adiantum capillus-veneris L.
ind
Adiantum hispidulum Sw.
Kl Le Ni K
O Mo L
O
M Ka H
Citation
L
M
K
O Mo L
M
H
Palmer 2003:43 (Ni, K, O, Mo, L, M, H); Staples & Herbst 2005:47 (KEY),
49 (DESCR)
nat
K
O Mo L
M Ka
H
Palmer 2003:43 (K, O, Mo, L, M, Ka, H); Staples & Herbst 2005:47 (KEY),
49 (DESCR)
Adiantum raddianum C.Presl
nat
K
O Mo L
M
H
Palmer 2003:45 (K, O, Mo, L, M, H); Staples & Herbst 2005:47 (KEY), 50
(DESCR)
Adiantum tenerum Sw.
nat
O
M
Ceratopteris gaudichaudii Brongn. var. vulgaris
Masuyama & Watano
nat
O
Ceratopteris thalictroides (L.) Brongn.
nat
K
O
Cheilanthes viridis (Forssk.) Sw.
nat
K
O
L
Coniogramme pilosa (Brack.) Hieron.
end
K
O Mo L
Doryopteris angelica K.R.Wood & W.H.Wagner
end
K
Doryopteris decipiens (Hook.) J.Sm.
end
Ni
Le Ni
K
Palmer 2003:43 (L, WM); Lau & Frohlich 2012:21 (O, DESCR)
Palmer 2003:45 (O, EM); Staples & Herbst 2005:47 (KEY), 50 (DESCR)
Masuyama & Watano 2010:79 (VARNOV/O, KEY, DESCR) [Note: a
presumably naturalized variety distributed from China, Japan, Korea, Nepal,
Taiwan, Guam, and Hawai‘i; 2 O‘ahu vouchers are cited, none from other
islands]
H
Palmer 2003:83 (K, O); Imada 2007:39 (H); Staples & Herbst 2005:42
(DESCR); Masuyama & Watano 2010:77 (KEY; Hawaiian Islands are
included in worldwide distribution, but no representative vouchers were
listed; thus, previously known island distribution maintained here)
M
H
Palmer 2003:85 (K, O, M, H); Oppenheimer 2007:30 (L)
M
H
Palmer 2003:98 (K, O, Mo, L, M, H)
Palmer 2003:132 (K); Yesilyurt 2005:550 (distribution listed as K, O, Mo, L,
KEY, DESCR)
O Mo L
M Ka
H
Palmer 2003:132 (Le, Ni, K, O, Mo, L, M, Ka, H); Yesilyurt 2005:552
(distribution listed as K, O, Mo, L, M, H, KEY, DESCR); Wood & LeGrande
2006:28 (Le)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 246
Scientific Name
Pteridophyte
Pteridaceae
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
Doryopteris decora Brack.
end
K
O Mo L
M Ka
H
Palmer 2003:133 (K, O, Mo, L, M, Ka, H); Yesilyurt 2005:553 (distribution
listed as K, O, Mo, L, M, H, KEY, DESCR), 557 (proposal to resurrect
Doryopteris tryonii (K) from synonymy of D. decora [Palmer 2003:133])
Doryopteris subdecipiens W.H.Wagner
end
K
O
M Ka
H
Palmer 2003:133 (K, O, L, M, Ka, H [as Doryopteris decipiens x D. decora]);
Yesilyurt 2005:553 (distribution listed as K, L, H, KEY, DESCR)
Doryopteris takeuchii (W.H.Wagner) W.H.Wagner
end
Pellaea ternifolia (Cav.) Link
ind
Pityrogramma Xmckenneyi W.H.Wagner
end
Pityrogramma austroamericana Domin
nat
Ni
K
O Mo L
M Ka
H
Palmer 2003:207 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst 2005:47
(KEY), 51 (DESCR)
Pityrogramma calomelanos (L.) Link
nat
Ni
K
O Mo L
M Ka
H
Palmer 2003:209 (Ni, K, O, Mo, L, M, Ka, H); Staples & Herbst 2005:47
(KEY), 51 (DESCR)
Pteris cretica L.
ind
K
O Mo L
M
H
Palmer 2003:225 (K, O, Mo, L, M, H); Staples & Herbst 2005:47 (KEY), 51
(DESCR)
Pteris excelsa Gaudich.
ind
K
O Mo L
M
H
Palmer 2003:227 (K, O, Mo, L, M, H)
Pteris hillebrandii Copel.
end
K
O Mo L
M
H
Palmer 2003:227 (K, O, Mo, L, M, H)
Pteris irregularis Kaulf.
end
K
O Mo L
M
H
Palmer 2003:227 (K, O, Mo, L, M, H)
Pteris lidgatei (Baker) Christ
end
O Mo
M
Palmer 2003:229 (O, Mo, WM); Oppenheimer 2008:33 (REDISCOVER/Mo)
Pteris tremula R.Br.
nat
M
Oppenheimer 2007:30 (NEWNAT/EM)
Pteris vittata L.
nat
L
O
Palmer 2003:133 (O); Yesilyurt 2005:555 (distribution listed as O, L, KEY,
DESCR)
M
H
O
K
Palmer 2003:201 (EM, H)
Palmer 2003:209 (O)
O Mo L
M
H
Palmer 2003:229 (K, O, M, H); Oppenheimer 2004:17 (L); Staples & Herbst
2005:47 (KEY), 52 (DESCR); Oppenheimer 2008:34 (Mo)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 247
Scientific Name
Pteridophyte
Salviniaceae
Salvinia molesta D.S.Mitch.
Status
Kl Le Ni K
O Mo L
M Ka H
Citation
nat
O
H
Palmer 2003:237 (O, H); Wilson 2003:6 (NEWNAT/O, H, DESCR); Staples
& Herbst 2005:52 (DESCR)
Lygodium japonicum (Thunb.) Sw.
nat
O
H
Palmer 2003:176 (H); Wilson 2003:5 (O); Frohlich & Lau 2012:47 (O range
extension)
Schizaea robusta Baker
end
K
O Mo L
M
H
Palmer 2003:241 (K, O, Mo, L, M, H)
Selaginella arbuscula (Kaulf.) Spring
end
K
O Mo L
M
H
Palmer 2003:274 (K, O, Mo, L, M, H)
Selaginella deflexa Brack.
end
K
O Mo
M
H
Palmer 2003:275 (K, O, Mo, M, H)
Selaginella kraussiana (Kunze) A.Braun
nat
O
M
H
Palmer 2003:275 (O, M, H); Staples & Herbst 2005:53 (DESCR)
Selaginella stellata Spring
nat
H
Palmer 2003:277 (H); Wilson 2003:6 (H, earlier coll.)
Selaginella umbrosa Lem. ex Hieron.
nat
H
Palmer 2003:277 (H); Wilson 2003:6 (NEWNAT/H); Staples & Herbst
2005:53 (DESCR)
Schizaeaceae
Selaginellaceae
Thelypteridaceae
Cyclosorus Xintermedius W.H.Shieh & J.L.Tsai
end
K
O
Cyclosorus Xpalmeri (W.H.Wagner) W.H.Wagner
end
K
O
Cyclosorus boydiae (D.C.Eaton) W.H.Wagner
end
Cyclosorus cyatheoides (Kaulf.) Farw.
end
K
M
Palmer 2003:91 (K, O, M [as Christella dentata x C. parasitica]); Snow et al.
2011:15 (NOM:Syn. C. xintermedia)
Palmer 2003:91 (K, O [as Christella cyatheoides x C. dentata]); Snow et al.
2011:15 (NOM:Syn. C. xpalmeri)
O
M
O Mo L
M
Palmer 2003:87 (O, EM [as Christella boydiae]); Snow et al. 2011:15
(NOM:Syn. C. boydiae)
H
Palmer 2003:88 (K, O, Mo, L, M, H [as Christella cyatheoides]); Snow et al.
2011:15 (NOM:Syn. C. cyatheoides)
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 248
Scientific Name
Pteridophyte
Thelypteridaceae
Status
Cyclosorus dentatus (Forssk.) Ching
nat
Cyclosorus hudsonianus (Brack.) Ching
Kl Le Ni K
Ni
O Mo L
M Ka H
Citation
K
O Mo L
M
H
Palmer 2003:88 (K, O, Mo, L, M, H [as Christella dentata]); Imada 2007:39
(Ni); Snow et al. 2011:15 (NOM:Syn. C. dentata)
end
K
O Mo L
M
H
Palmer 2003:212 (K, O, Mo, L, M, H [as Pneumatopteris hudsoniana]);
Palmer 2005:81 (KEY); Snow et al. 2011:15 (NOM:Syn. P. hudsoniana)
Cyclosorus interruptus (Willd.) H.Itô
ind
K
O Mo L
M
H
Palmer 2003:102 (K, O, Mo, L, M, H)
Cyclosorus parasiticus (L.) Farw.
nat
K
O Mo L
M
H
Palmer 2003:90 (K, O, Mo, L, M, H [as Christella parasitica]); Snow et al.
2011:15 (NOM:Syn. C. parasitica)
Cyclosorus pendens (D.D.Palmer) N.Snow
end
K
O Mo
M
H
Palmer 2005:81 (SPNOV/K, O, Mo, EM, WM, H [as Pneumatopteris
pendens], DESCR, KEY); Snow et al. 2011:15 (COMBNOV:Syn. P.
pendens); Wood 2012:100 (REDISCOVER/K)
Cyclosorus sandwicensis (Brack.) Copel.
end
K
O Mo L
M
H
Palmer 2003:214 (K, O, Mo, L, M, H [as Pneumatopteris sandwicensis]);
Palmer 2005:81 (KEY); Snow et al. 2011:15 (NOM:Syn. P. sandwicensis)
Cyclosorus wailele (Flynn) W.H.Wagner
end
K
Macrothelypteris torresiana (Gaudich.) Ching
nat
K
O
M
H
Palmer 2003:176 (K, O, M, H)
Pseudophegopteris keraudreniana (Gaudich.) Holttum
end
K
O Mo L
M
H
Palmer 2003:222 (K, O, Mo, L, M, H)
Thelypteris globulifera (Brack.) C.F.Reed
end
K
O Mo L
M
H
Palmer 2003:47 (K, O, Mo, L, M, H [as Amauropelta globulifera]); Snow et
al. 2011:15 (NOM:Syn. A. globulifera)
ind
K
O Mo L
M
H
Palmer 2003:166 (K, O, Mo, L, M, H)
Palmer 2003:90 (K [as Christella wailele]); Snow et al. 2011:15 (NOM:Syn.
C. wailele)
Vittariaceae
Haplopteris elongata (Sw.) E.H.Crane
Islands: Kl=Ka‘ula Rock; Le=Lehua; Ni=Ni‘ihau; K=Kaua‘i; O=O‘ahu; Mo=Moloka‘i; L=Läna‘i; M=Maui; Ka=Kaho‘olawe; H=Hawai‘i
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
A- 249
APPENDIX B. Northwestern Hawaiian Islands Native and Naturalized Vascular Plants Checklist
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Acanthaceae
Asystasia gangetica (L.) T.Anderson
nat
Mi
Wagner et al. 1990:168 (Mi); Staples & Herbst 2005:81 (KEY), 82 (DESCR)
Ruellia brittoniana Leonard
nat
Mi
Wagner et al. 1990:174 (Mi); Staples & Herbst 2005:80, 81, 91 (KEY), 91
(DESCR)
Sesuvium portulacastrum (L.) L.
ind
Mi PH Li
Tetragonia tetragonioides (Pall.) Kuntze
nat
Mi
Aizoaceae
La
Ne
Wagner et al. 1990:176 (PH, Li, La, Ne); Staples & Herbst 2005:96 (KEY), 99
(DESCR); Wagner et al. 2012:4 (Mi)
N
Wagner et al. 1990:178 (N); Starr & Martz 2000:10 (Mi); Staples & Herbst
2005:96 (KEY), 98 (DESCR)
Amaranthaceae
Achyranthes aspera L. var. aspera
nat
Achyranthes atollensis H.St.John
end
Amaranthus brownii Christoph. & Caum
end
Amaranthus lividus L. ssp. polygonoides (Moq.)
Probst
nat
Mi
Amaranthus spinosus L.
nat
Ku Mi
Amaranthus viridis L.
nat
Ku Mi
nat
Mi
Ku Mi PH
La
Wagner et al. 1990:180 (La)
La
Wagner et al. 1990:181 (Ku, Mi, PH, La)
N
Wagner et al. 1990:186 (N)
Wagner et al. 1990:188 (no NWHI records); Bruegmann 1999:1 (Mi)
Wagner et al. 1990:188 (Ku); Starr et al. 2002:17 (Mi)
La
Wagner et al. 1990:189 (Ku); Wagner & Herbst 1995:14 (Mi); Shannon &
Wagner 1996:13 (La)
Anacardiaceae
Schinus terebinthifolius Raddi
Wagner et al. 1990:198 (Mi); Staples & Herbst 2005:104 (KEY), 106 (DESCR)
Apiaceae
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 1
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Apiaceae
Anethum graveolens L.
nat
Mi
Wagner et al. 1990:200 (no NWHI records); Starr et al. 2002:17 (Mi); Staples &
Herbst 2005:112 (KEY), 113 (DESCR)
Ciclospermum leptophyllum (Pers.) Sprague ex
Britton & P.Wilson
nat
Ku Mi
Catharanthus roseus (L.) G.Don
nat
Mi
Wagner et al. 1990:215 (Mi); Staples & Herbst 2005:117 (KEY), 120 (DESCR)
Thevetia peruviana (Pers.) K.Schum.
nat
Mi
Wagner et al. 1990:215 (as Cascabela thevetia, no NWHI records); Starr et al.
2002:17 (Mi); Wagner & Herbst 2003:5 (NOM:Syn. C. thevetia); Staples &
Herbst 2005:116 (KEY), 127 (DESCR)
Bidens alba (L.) DC. var. radiata (Sch.Bip.)
Ballard ex Melchert
nat
Ku Mi
Bidens pilosa L.
nat
Mi
Wagner et al. 1990:279 (Mi)
Calyptocarpus vialis Less.
nat
Mi
Wagner et al. 1990:284 (no NWHI records); Bruegmann 1999:1 (Mi)
Conyza bonariensis (L.) Cronquist
nat
Ku Mi
Conyza canadensis (L.) Cronquist var. pusilla
(Nutt.) Cronquist
nat
Mi
Emilia fosbergii Nicolson
nat
Ku
Wagner et al. 1990:312 (Ku)
Flaveria trinervia (Spreng.) C.Mohr
nat
Ku
Wagner et al. 1990:317 (no NWHI records); Starr et al. 2010:62 (Ku)
Gaillardia pulchella Foug.
nat
Ku
Wagner et al. 1990:318 (Ku)
Wagner et al. 1990:201 (Mi); Starr et al. 2003:24 (Ku)
Apocynaceae
Asteraceae
Wagner et al. 1990:270 (Ku, Mi)
La
FF
Wagner et al. 1990:288 (Ku, Mi, La, FF)
Wagner et al. 1990:288 (no NWHI records); Wagner & Herbst 1995:16 (Mi)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 2
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Asteraceae
Melanthera integrifolia (Nutt.) W.L.Wagner &
H.Rob.
end
Ku
La
Wagner et al. 1990:336 (as Lipochaeta integrifolia, Ku, La); Wagner & Robinson
2002 (NOM:Syn. L. integrifolia); Staples & Herbst 2005:150 (KEY), 169
(DESCR)
Pluchea Xfosbergii Cooperr. & Galang
nat
Mi
Pluchea carolinensis (Jacq.) G.Don
nat
Ku Mi
Pluchea indica (L.) Less.
nat
Mi
Pseudognaphalium sandwicensium (Gaudich.)
Anderb. var. sandwicensium
end
Ku Mi
Sonchus oleraceus L.
nat
Ku Mi PH
Sphagneticola trilobata (L.) Pruski
nat
Mi
Wagner et al. 1990:373 (as Wedelia trilobata, Mi); Wagner et al. 1997:55
(NOM:Syn. W. trilobata); Staples & Herbst 2005:150 (KEY), 165 (DESCR)
Tridax procumbens L.
nat
Mi
Wagner et al. 1990:370 (Mi)
Verbesina encelioides (Cav.) Benth. & Hook.
nat
Ku Mi PH
Xanthium strumarium L. var. canadense (Mill.)
Torr. & A.Gray
nat
Mi
Wagner et al. 1990:376 (Mi)
nat
Mi
Wagner et al. 2012:17 (Mi)
Wagner et al. 1990:350 (Mi)
FF
Wagner et al. 1990:351 (as Pluchea symphytifolia, Ku, Mi, FF); Khan & Jarvis
1989:661/Wagner & Herbst 1995:16 (NOM: P. symphytifolia misapplied)
La
Wagner et al. 1990:351 (Mi, La)
Wagner et al. 1990:321 (as Gnaphalium sandwicensium var. sandwicensium, Ku,
Mi); Wagner et al. 1997:54 (NOM:Syn. G. sandwicensium var. sandwicensium);
Staples & Herbst 2005:149 (KEY), 170 (DESCR)
FF
N
Wagner et al. 1990:358 (Ku, Mi, PH, FF, N)
Wagner et al. 1990:372 (Ku, Mi); Starr et al. 2002:18 (PH)
Basellaceae
Basella alba L.
Boraginaceae
Heliotropium curassavicum L.
ind
La
FF
N
Wagner et al. 1990:395 (La, FF, N)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 3
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Boraginaceae
Heliotropium procumbens Mill. var. depressum
(Cham.) Fosberg
nat
Tournefortia argentea L.f.
nat
Mi
Ku Mi PH Li
La
FF
Wagner et al. 1990:396 (Mi, FF)
FF
Wagner et al. 1990:398 (Ku, Mi, PH, Li, La, FF); Staples & Herbst 2005:197
(KEY), 199 (DESCR)
Brassicaceae
Brassica nigra (L.) W.D.J.Koch
nat
Mi
Brassica rapa L.
nat
Capsella bursa-pastoris (L.) Medik.
nat
Mi
Coronopus didymus (L.) Sm.
nat
Mi PH
La
Wagner et al. 1990:403 (Mi, PH); Staples et al. 2003:9 (La)
Lepidium bidentatum Montin var. o-waihiense
(Cham. & Schltdl.) Fosberg
end
Ku Mi PH
La
Wagner et al. 1990:406 (Ku, Mi, PH, La)
Lepidium virginicum L.
nat
Ku Mi
Wagner et al. 1990:409 (Mi); Starr et al. 2003:26 (Ku)
Lobularia maritima (L.) Desv.
nat
Ku Mi
Wagner et al. 1990:409 (Ku, Mi); Staples & Herbst 2005:200 (KEY), 203
(DESCR)
Wagner et al. 1990:400 (no NWHI records); Starr et al. 2003:25 (Mi)
PH
Wagner et al. 1990:400 (as Brassica campestris, PH); Al-Shehbaz
1985:290/Wagner & Herbst 1995:17 (NOM:Syn. B. campestris); Staples & Herbst
2005:200 (KEY), 202 (DESCR)
Wagner et al. 1990:402 (as Capsella rubella, no NWHI records); Wagner &
Herbst 1995:17 (NOM:Syn. C. rubella); Herbst & Wagner 1996:10 (Mi)
Capparaceae
Capparis sandwichiana DC.
end
Mi PH
Sagina japonica (Sw.) Ohwi
nat
Mi
Schiedea verticillata F.Br.
end
La
Wagner et al. 1990:492 (Mi, PH, La); Staples & Herbst 2005:219 (DESCR)
Caryophyllaceae
Wagner et al. 1990:506 (no NWHI records); Starr et al. 2003:26 (Mi)
N
Wagner et al. 1990:520 (N); Wagner et al. 2005:48 (KEY, DESCR)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 4
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Caryophyllaceae
Spergularia marina (L.) Griseb.
nat
Ku Mi
FF
Stellaria media (L.) Vill.
nat
Ku Mi
Casuarina equisetifolia L.
nat
Ku Mi PH Li
Casuarina glauca Sieber ex Spreng.
nat
Wagner et al. 1990:526 (Ku, Mi, FF)
Wagner et al. 1990:526 (Ku); Starr et al. 2003:26 (Mi)
Casuarinaceae
La
FF
Wagner et al. 1990:528 (Ku, Mi, PH, Li, La, FF); Staples & Herbst 2005:228
(KEY), 229 (DESCR)
Mi
Wagner et al. 1990:529 (no NWHI records); Starr et al. 2002:19 (Mi); Staples &
Herbst 2005:228 (KEY), 230 (DESCR)
Chenopodiaceae
Atriplex suberecta I.Verd.
nat
Chenopodium murale L.
nat
Chenopodium oahuense (Meyen) Aellen
end
Salicornia virginica L.
nat
Ku Mi
Li
La
FF
Wagner et al. 1990:535 (FF)
FF
Wagner et al. 1990:538 (Ku, Mi, FF)
FF
Ne N
FF
Wagner et al. 1990:538 (Li, La, FF, Ne, N)
Wagner et al. 1990:540 (FF) [Note: single coll. in 1968; population apparently
destroyed in 1969 storm]
Convolvulaceae
Ipomoea aquatica Forssk.
nat
Mi
Wagner et al. 2012:29 (Mi)
Ipomoea batatas (L.) Lam.
pol
Mi
Wagner et al. 2012:29 (Mi)
Ipomoea indica (Burm.) Merr.
ind
Ku Mi
Li
La
Ipomoea pes-caprae (L.) R.Br. ssp. brasiliensis (L.)
Ooststr.
ind
Ku Mi
Li
La
Ipomoea triloba L.
nat
Mi
FF
N
Wagner et al. 1990:556 (Ku, Mi, Li, La, N); Staples & Herbst 2005:245, 247
(KEY), 249 (DESCR)
N
Wagner et al. 1990:559 (Mi, Li, La, FF, N); Starr et al. 2003:26 (Ku); Staples &
Herbst 2005:245, 247 (KEY), 249 (DESCR)
Wagner et al. 1990:559 (Mi)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 5
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Cucurbitaceae
Coccinia grandis (L.) Voigt
nat
Mi
Wagner et al. 1990:569 (no NWHI records); Staples & Herbst 2005:262 (KEY),
265 (DESCR); Starr et al. 2010:63 (Mi)
Momordica charantia L.
nat
Mi
Wagner et al. 1990:572 (no NWHI records); Starr & Martz 2000:10 (Mi); Staples
& Herbst 2005:262 (KEY), 268 (DESCR)
Sicyos maximowiczii Cogn.
end
Sicyos pachycarpus Hook. & Arn.
end
La
Sicyos semitonsus H.St.John
end
La
Ku
PH Li
La
Wagner et al. 1990:578 (Ku, PH, Li, La); Wagner & Shannon 1999:444
N
Wagner et al. 1990:579 (La, N); Wagner & Shannon 1999:445
Wagner et al. 1990:579 (La); Wagner & Shannon 1999:446
Euphorbiaceae
N
Wagner et al. 1990:606 (as Chamaesyce celastroides var. celastroides, N);
Govaerts et al. 2000:318 (NOM:Syn. C. celastroides var. celastroides); Steinmann
& Porter 2002/Yang & Berry 2011 (molecular supoort for nesting of Chamaesyce
within Euphorbia)
Euphorbia celastroides Boiss. var. celastroides
end
Euphorbia cyathophora Murray
nat
Mi
Wagner et al. 1990:618 (Mi); Staples & Herbst 2005:278, 285 (KEY), 288
(DESCR)
Euphorbia heterophylla L.
nat
Mi
Wagner et al. 1990:619 (Mi)
Euphorbia hirta L.
nat
Ku Mi
FF
Wagner et al. 1990:609 (as Chamaesyce hirta, Ku, Mi, FF); Govaerts et al.
2000:320 (NOM:Syn. C. hirta); Steinmann & Porter 2002/Yang & Berry 2011
(molecular supoort for nesting of Chamaesyce within Euphorbia)
Euphorbia hypericifolia L.
nat
Ku Mi
FF
Wagner et al. 1990:609 (as Chamaesyce hypericifolia, Ku, Mi, FF); Govaerts et
al. 2000:320 (NOM:Syn. C. hypericifolia); Steinmann & Porter 2002/Yang &
Berry 2011 (molecular supoort for nesting of Chamaesyce within Euphorbia)
Euphorbia hyssopifolia L.
nat
Mi
Wagner et al. 1990:610 (as Chamaesyce hyssopifolia, no NWHI records);
Bruegmann 1999:2 (Mi); Govaerts et al. 2000:320 (NOM:Syn. C. hyssopifolia);
Steinmann & Porter 2002/Yang & Berry 2011 (molecular supoort for nesting of
Chamaesyce within Euphorbia)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 6
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Euphorbiaceae
Euphorbia maculata L.
nat
Ku Mi
Wagner et al. 1990:610 (as Chamaesyce maculata, Ku, Mi); Govaerts et al.
2000:322 (NOM:Syn. C. maculata); Steinmann & Porter 2002/Yang & Berry
2011 (molecular supoort for nesting of Chamaesyce within Euphorbia)
Euphorbia peplus L.
nat
Ku Mi
Wagner et al. 1990:619 (Mi); Starr et al. 2003:27 (Ku)
Euphorbia prostrata Aiton
nat
Mi
Wagner et al. 1990:613 (as Chamaesyce prostrata, Mi); Govaerts et al. 2000:324
(NOM:Syn. C. prostrata); Steinmann & Porter 2002/Yang & Berry 2011
(molecular supoort for nesting of Chamaesyce within Euphorbia)
Albizia lebbeck (L.) Benth.
nat
Mi
Wagner et al. 1990:644 (Mi); Staples & Herbst 2005:298 (KEY), 330 (DESCR)
Caesalpinia bonduc (L.) Roxb.
ind
Mi
Crotalaria incana L.
nat
Mi
Wagner et al. 1990:660 (Mi)
Crotalaria pallida Aiton
nat
Mi
Wagner et al. 1990:661 (Mi)
Desmanthus pernambucanus (L.) Thell.
nat
Mi
Wagner et al. 1990:665 (as Desmanthus virgatus, Mi); Wagner & Herbst 1995:20
(NOM:Syn. D. virgatus)
Desmodium sandwicense E.Mey.
nat
Mi
Wagner et al. 1990:669 (Mi)
Indigofera spicata Forssk.
nat
Mi
Wagner et al. 1990:675 (as Indigofera spicata, no NWHI records); Herbst &
Wagner 1999:21 (NOM: I. spicata misapplied = I. hendecaphylla); Starr et al.
2010:64 (Mi); Wagner et al. 2012:39 (NOM: I. hendecaphylla misapplied)
Leucaena leucocephala (Lam.) de Wit
nat
Mi
Wagner et al. 1990:679 (Mi); Staples & Herbst 2005:298 (KEY), 333 (DESCR)
Medicago lupulina L.
nat
Mi
Wagner et al. 1990:684 (Mi)
Medicago polymorpha L.
nat
Mi
Wagner et al. 1990:684 (no NWHI records); Bruegmann 1999:2 (Mi)
Fabaceae
La
Wagner et al. 1990:647 (no NWHI records); Staples et al. 2003:11 (La); Starr et
al. 2010:64 (Mi)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 7
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Fabaceae
Medicago sativa L.
nat
Mi
Wagner et al. 1990:686 (Mi)
Melilotus alba Medik.
nat
Mi
Wagner et al. 1990:686 (Mi)
Melilotus indica (L.) All.
nat
Mi
Wagner et al. 1990:687 (Mi)
Prosopis pallida (Humb. & Bonpl. ex Willd.)
Kunth
nat
Mi
Wagner et al. 1990:692 (Mi); Staples & Herbst 2005:298 (KEY), 322 (DESCR)
Sesbania tomentosa Hook. & Arn.
end
Vachellia farnesiana (L.) Wight & Arn.
nat
Ne N
Mi
Wagner et al. 1990:704 (Ne, N); Staples & Herbst 2005:300 (KEY), 327 (DESCR)
Wagner et al. 1990:641 (as Acacia farnesiana, Mi); Mabberley 2008:890/Wagner
et al. 2012:36 (NOM:Syn. A. farnesiana)
Frankeniaceae
Frankenia salina (Molina) I.M.Johnst.
nat
FF
Wagner et al. 1990:724 (FF)
Gentianaceae
Centaurium erythraea Raf. ssp. erythraea
nat
Wagner et al. 1990:725 (Mi)
Mi
Goodeniaceae
Scaevola taccada (Gaertn.) Roxb.
FF
Wagner et al. 1990:788 (as Scaevola sericea, Ku, Mi, PH, Li, La, FF); Green
1991/Shannon et al. 1997 (NOM:Syn. S. sericea); Staples & Herbst 2005:348
(DESCR)
ind
Ku Mi PH Li
La
end
Li
La
Wagner et al. 1990:795 (Li, La)
La
Wagner et al. 1990:823 (Ku, Mi, La); Wagner 1999a:276
Hydrophyllaceae
Nama sandwicensis A.Gray
Lamiaceae
Phyllostegia variabilis Bitter
end
Ku Mi
Lauraceae
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 8
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Lauraceae
Cassytha filiformis L.
ind
Ku
Wagner et al. 1990:844 (no NWHI records); Starr et al. 2003:28 (Ku)
Malvaceae
Abutilon grandifolium (Willd.) Sweet
nat
Mi
Wagner et al. 1990:872 (Mi); Staples & Herbst 2005:383, 385 (KEY), 385
(DESCR)
Hibiscus tiliaceus L.
ind?
Mi
FF
Wagner et al. 1990:888 (Mi, FF); Fryxell 2001:258 (COMBNOV [Talipariti
tiliaceum]; Pfeil et al. 2002/Wagner & Herbst 2003:33 (Talipariti is nested within
Hibiscus and best maintained there); Staples & Herbst 2005:383, 387 (KEY), 390
(DESCR)
Malva parviflora L.
nat
Mi
FF
Wagner et al. 1990:893 (FF); Starr et al. 2004:24 (Mi)
Malvastrum coromandelianum (L.) Garcke ssp.
coromandelianum
nat
Mi
Sida fallax Walp.
ind
Mi
Sida rhombifolia L.
nat?
Mi
Wagner et al. 1990:898 (no NWHI records); Starr & Martz 2000:10 (Mi)
Ficus macrophylla Desf. ex Pers.
nat
Mi
Wagner et al. 1990:924 (no NWHI records); Staples & Herbst 2005:403, 407
(KEY), 409 (DESCR); Starr et al. 2010:64 (Mi)
Ficus microcarpa L.f.
nat
Mi
Wagner et al. 1990:924 (no NWHI records); Starr et al. 2002:21 (Mi); Staples &
Herbst 2005:403, 408 (KEY), 409 (DESCR)
Wagner et al. 1990:894 (Mi)
N
Wagner et al. 1990:897 (Mi, N); Staples & Herbst 2005:383 (KEY), 392 (DESCR)
Moraceae
Nyctaginaceae
Boerhavia acutifolia (Choisy) J.W.Moore
ind
Boerhavia coccinea Mill.
nat
PH
La
Wagner et al. 1990:978 (as Boerhavia glabrata, no NWHI records); Herbst &
Wagner 1999:24 (NOM: B. glabrata misapplied); Staples et al. 2003:15 (PH, La)
Ku
Wagner et al. 1990:978 (no NWHI records); Starr et al. 2003:29 (Ku)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 9
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Nyctaginaceae
Boerhavia herbstii Fosberg
end
PH Li
Boerhavia repens L.
ind
Ku Mi PH Li
Mirabilis jalapa L.
nat
Pisonia grandis R.Br.
ind
Wagner et al. 1990:980 (PH, Li)
La
FF
N
Mi
Wagner et al. 1990:981 (Ku, Mi, PH, Li, La, FF); Evenhuis & Eldredge 2004:55
(N)
Wagner et al. 1990:982 (Mi); Staples & Herbst 2005:434 (DESCR)
Li
Wagner et al. 1990:985 (Li); Staples & Herbst 2005:434 (DESCR)
Onagraceae
Oenothera laciniata Hill
nat
Ku Mi
Wagner et al. 1990:1000 (Mi); Wagner & Herbst 1995:22 (Ku)
Oxalis corniculata L.
pol?
Ku Mi
Wagner et al. 1990:1002 (Mi); Starr et al. 2003:29 (Ku); Staples & Herbst
2005:447 (KEY)
Oxalis debilis Kunth var. corymbosa (DC.)
Lourteig
nat
Mi
Plantago lanceolata L.
nat
Mi
Plantago major L.
nat
Mi
Wagner et al. 1990:1051 (Mi)
Antigonon leptopus Hook. & Arn.
nat
Mi
Wagner et al. 1990:1060 (no NWHI records); Starr et al. 2002:23 (Mi); Staples &
Herbst 2005:461 (KEY), 462 (DESCR)
Coccoloba uvifera (L.) L.
nat
Mi
Wagner et al. 1990:1059 (no NWHI records); Starr et al. 2002:24 (Mi); Staples &
Herbst 2005:461 (KEY), 462 (DESCR)
Rumex albescens Hillebr.
end
Oxalidaceae
Wagner et al. 1990:1002 (as Oxalis corymbosa, no NWHI records); Herbst &
Wagner 1999:25 (NOM:Syn. O. corymbosa = O. debilis); Staples & Herbst
2005:447 (KEY), 448 (DESCR, as var. corymbosa); Starr et al. 2010:65 (Mi)
Plantaginaceae
FF
Wagner et al. 1990:1051 (Mi, FF)
Polygonaceae
N
Wagner et al. 1990:1066 (N)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 10
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Portulacaceae
Portulaca lutea Sol. ex G.Forst.
ind
Mi
Portulaca oleracea L.
nat
Ku Mi PH
Portulaca villosa Cham.
end
Li
La GP FF
La
Ne N
FF
Wagner et al. 1990:1071 (Mi, Li, La, GP, FF, Ne, N)
N
Wagner et al. 1990:1072 (Mi, PH, La, FF, N); Staples & Herbst 2005:465 (KEY),
466 (DESCR); Kennedy et al. 2010:22 (Ku)
N
Wagner et al. 1990:1074 (N)
Primulaceae
Anagallis arvensis L.
nat
Ku Mi
Wagner et al. 1990:1077 (Mi); Starr et al. 2003:30 (Ku)
Santalaceae
Santalum ellipticum Gaudich.
end
La
Wagner et al. 1990:1220 (La)
Scrophulariaceae
Bacopa monnieri (L.) Wettst.
ind
Mi
Wagner et al. 1990:1237 (Mi); Staples & Herbst 2005:524 (KEY), 525 (DESCR)
Solanaceae
Nicotiana tabacum L.
nat
La
Solanum americanum Mill.
ind?
Ku Mi PH Li
La
N
Wagner et al. 1990:1268 (Ku, Mi, PH, Li, La, N); Staples & Herbst 2005:528,
536 (KEY), 539 (DESCR)
Solanum nelsonii Dunal
end
Ku Mi PH
La
N
Wagner et al. 1990:1273 (Ku, Mi, PH, La, N)
ind?
Mi
Wagner et al. 1990:1262 (La); Staples & Herbst 2005:528 (KEY), 540 (DESCR)
Sterculiaceae
Waltheria indica L.
Wagner et al. 1990:1280 (Mi); Saunders 2011:79 (proposed resurrection of
Waltheria pyrolifolia A.Gray (WM) from synonymy of W. indica); Wagner et al.
2012:71 (W. pyrifolia not recognized as distinct)
Urticaceae
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 11
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Dicot
Urticaceae
Pilea microphylla (L.) Liebm.
nat
Mi
Wagner et al. 1990:1306 (no NWHI records); Wagner & Herbst 1995:26 (Mi);
Shannon & Wagner 1996:14 (Mi again); Staples & Herbst 2005:553 (KEY), 554
(DESCR)
Lantana camara L.
nat
Mi
Wagner et al. 1990:1320 (Mi); Staples & Herbst 2005:556 (KEY), 564 (DESCR)
Phyla nodiflora (L.) Greene
nat
Mi
Starr et al. 2002:26 (Mi); Staples & Herbst 2005:555 (KEY), 566 (DESCR)
Stachytarpheta jamaicensis (L.) Vahl
nat
Mi
Wagner et al. 1990:1322 (Mi)
Verbena litoralis Kunth
nat
Mi
Wagner et al. 1990:1325 (Mi)
Verbenaceae
Zygophyllaceae
Tribulus cistoides L.
ind
Ku Mi PH Li
La
FF
N
Wagner et al. 1990:1343 (Ku, Mi, PH, Li, La, FF, N); Staples & Herbst 2005:576
(DESCR)
Monocot
Arecaceae
Cocos nucifera L.
pol
Pritchardia remota Becc.
end
La
Wagner et al. 1990:1362 (La); Staples & Herbst 2005:617 (KEY), 620 (DESCR)
N
Wagner et al. 1990:1373 (N); Hodel 2007:S-39 (Ni, NOM:Syn. Pritchardia
aylmer-robinsonii)
Commelinaceae
Commelina diffusa Burm.f.
nat
Mi
Wagner et al. 1990:1379 (Mi)
nat
Mi
Wagner et al. 1990:1395 (as Cyperus alternifolius ssp. flabelliformis, Mi); Tucker
1994:20/Strong & Wagner 1997:41 (NOM:Syn. C. alternifolius ssp.
flabelliformis); Staples & Herbst 2005:656 (KEY, DESCR)
Cyperaceae
Cyperus involucratus Rottb.
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 12
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Monocot
Cyperaceae
Cyperus javanicus Houtt.
ind
Mi
Wagner et al. 1990:1420 (as Mariscus javanicus, Mi); Tucker 1994:87/Wagner &
Herbst 1999:1900 (NOM:Syn. M. javanicus)
Cyperus laevigatus L.
ind
La
Wagner et al. 1990:1398 (La)
Cyperus pennatiformis Kük. var. bryanii Kük.
end
La
Wagner et al. 1990:1421 (as Mariscus pennatiformis subsp. bryanii, La); Tucker
1994:87/Wagner & Herbst 1999:1900 (NOM:Syn. M. pennatiformis subsp.
bryanii)
Cyperus polystachyos Rottb.
ind
Mi
Cyperus rotundus L.
nat
Ku Mi
Fimbristylis cymosa R.Br. ssp. spathacea (Roth)
T.Koyama
ind
Mi
La
Fimbristylis cymosa R.Br. ssp. umbellato-capitata
(Hillebr.) T.Koyama
ind
Ku Mi
La
Halophila decipiens Ostenf.
nat
Mi
McDermid et al. 2002 (NEWNAT/Mi, DESCR)
Halophila hawaiiana Doty & B.C.Stone
end
Mi PH
Wagner et al. 1990:1443 (no NWHI records); DeFelice 1999:3 (Mi); McDermid
et al. 2003 (DESCR); Kennedy et al. 2010:22 (PH)
?
Mi?
Wagner et al. 1990:1482 (Mi? [introduced to stabilize sand dunes, apparently
persisting vegetatively], DESCR)
Andropogon glomeratus (Walter) Britton, Sterns &
Poggenb. var. pumilus (Vasey) L.H.Dewey
nat
Mi
Snow & Lau 2010:47 (Mi; NOM: Andropogon virginicus misapplied, KEY)
Bothriochloa pertusa (L.) A.Camus
nat
Mi
Wagner et al. 1990:1503 (no NWHI records); Starr & Martz 2000:11 (Mi)
Wagner et al. 1990:1425 (as Pycreus polystachyos subsp. polystachyos, Mi);
Wagner & Herbst 1999:1901 (NOM:Syn. P. polystachyos subsp. polystachyos)
FF
Wagner et al. 1990:1399 (Ku, Mi); Strong & Wagner 1997:42 (FF)
FF
Wagner et al. 1990:1405 (Mi, La, FF)
Wagner et al. 1990:1405 (Ku, Mi, La)
Hydrocharitaceae
Poaceae
Ammophila arenaria (L.) Link
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 13
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Monocot
Poaceae
Bromus catharticus Vahl
nat
Mi
Wagner et al. 1990:1508 (as Bromus willdenowii, Mi); Herbst & Clayton 1998:20
(NOM:Syn. B. willdenowii)
Cenchrus agrimonioides Trin. var. laysanensis
F.Br.
end
Ku Mi
Cenchrus ciliaris L.
nat
Mi
Cenchrus echinatus L.
nat
Ku Mi PH Li
Chloris barbata Sw.
nat
Ku Mi
Chloris divaricata R.Br. var. divaricata
nat
Mi
Chloris virgata Sw.
nat
Ku Mi
Cynodon dactylon (L.) Pers.
nat
Ku Mi PH
Dactyloctenium aegyptium (L.) Willd.
nat
Mi
Digitaria ciliaris (Retz.) Koeler
nat
Ku Mi
Digitaria insularis (L.) Mez ex Ekman
nat
Mi
Wagner et al. 1990:1531 (Mi)
Diplachne fusca (L.) Stapf ssp. uninervia (J.Presl)
P.M.Peterson & N.Snow
nat
Mi
Wagner et al. 1990:1558 (as Leptochloa uninervia, no NWHI records); Starr &
Martz 2000:11 (Mi); Imada et al. 2000:14 (NOM:Syn. L. uninervia = L. fusca
subsp. uninervia); Peterson et al. 2012:1327/Wagner et al. 2012:96 (COMBNOV,
Syn. L. fusca subsp. uninervia)
Echinochloa crusgalli (L.) P.Beauv.
nat
Mi
Wagner et al. 1990:1534 (no NWHI records); Starr et al. 2002:22 (Mi)
La
Wagner et al. 1990:1512 (Ku, Mi, La); Herbst & Clayton 1998:21 (KEY)
Wagner et al. 1990:1512 (no NWHI records); Starr & Martz 2000:11 (Mi); Herbst
& Clayton 1998:21 (KEY)
La
FF
N
Wagner et al. 1990:1512 (Ku, Mi, Li, La, FF, N); Starr et al. 2002:22 (PH);
Herbst & Clayton 1998:21 (KEY)
Wagner et al. 1990:1514 (Ku, Mi)
Wagner et al. 1990:1515 (no NWHI records); Starr & Martz 2000:11 (Mi)
Wagner et al. 1990:1515 (Ku); Starr & Martz 2000:11 (Mi)
La
FF
Wagner et al. 1990:1520 (Ku, Mi, PH, La, FF); Staples & Herbst 2005:741
(KEY), 745 (DESCR)
Wagner et al. 1990:1521 (no NWHI records); Wagner et al. 1997:60 (Mi)
FF
Wagner et al. 1990:1530 (Ku, Mi, FF)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 14
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Monocot
Poaceae
Eleusine indica (L.) Gaertn.
nat
Ku Mi
Eragrostis amabilis (L.) Wight & Arn.
nat
Ku Mi
Eragrostis paupera Jedwabn.
ind
Ku Mi PH
Eragrostis variabilis (Gaudich.) Steud.
end
Ku Mi PH Li
Eriochloa procera (Retz.) C.E.Hubb.
nat
Mi
Eustachys petraea (Sw.) Desv.
nat
Mi
Festuca myuros L.
nat
Mi
Wagner et al. 1990:1603 (as Vulpia myuros, Mi); Wagner et al. 2012:101
(NOM:Syn. V. myuros)
Hordeum murinum L. ssp. leporinum (Link)
Arcang.
nat
Mi
Wagner et al. 1990:1552 (as Hordeum leporinum, Mi); Herbst & Clayton 1998:28
(NOM:Syn. H. leporinum)
Lepturus repens (G.Forst.) R.Br.
ind
Melinis repens (Willd.) Zizka
nat
Panicum torridum Gaudich.
end
Paspalum fimbriatum Kunth
nat
Paspalum setaceum Michx.
nat
Mi
Starr & Martz 2000:11 (Mi)
Paspalum urvillei Steud.
nat
Mi
Wagner et al. 1990:1577 (Mi)
Ku Mi PH Li
FF
Wagner et al. 1990:1537 (Ku, Mi, FF)
Wagner et al. 1990:1545 (as Eragrostis tenella, Mi); Lorence et al. 1995:45
(NOM:Syn. E. tenella); Starr et al. 2003:30 (Ku)
FF
Wagner et al. 1990:1543 (Ku, Mi, PH, FF)
La
N
Wagner et al. 1990:1545 (Ku, Mi, PH, Li, La, N); Snow & Lau 2010:50
(NOM:Syn. Eragrostis hobdyi)
Snow 2010:52 (Mi); Starr & Starr 2011:30 (Mi again)
FF
La
Wagner et al. 1990:1513 (as Chloris petraea, Mi?, FF? [adventive]); Wagner &
Herbst 1995:24 (Mi, FF; NOM:Syn. C. petraea)
FF
Wagner et al. 1990:1560 (Ku, Mi, PH, Li, La, FF)
Mi
Wagner et al. 1990:1588 (as Rhynchelytrum repens, Mi); Herbst & Clayton
1998:29 (NOM:Syn. R. repens, KEY)
Ne N
Ku
Wagner et al. 1990:1573 (Ne, N)
Wagner et al. 1990:1576 (Ku)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 15
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Monocot
Poaceae
Poa annua L.
nat
Ku Mi
Wagner et al. 1990:1583 (Ku, Mi)
Polypogon interruptus Kunth
nat
Ku Mi
Wagner et al. 1990:1585 (Mi); Wagner & Herbst 1995:24 (Ku)
Polypogon monspeliensis (L.) Desf.
nat
Mi
Setaria verticillata (L.) P.Beauv.
nat
Ku Mi PH
Sporobolus indicus (L.) R.Br.
nat
Mi
Wagner et al. 1990:1597 (Mi)
Sporobolus piliferus (Trin.) Kunth
nat
Mi
Starr et al. 2010:66 (Mi)
Sporobolus pyramidatus (Lam.) Hitchc.
nat
Ku Mi
La
Sporobolus virginicus (L.) Kunth
ind
Mi
La
Stenotaphrum secundatum (Walter) Kuntze
nat
Mi
Wagner et al. 1990:1598 (Mi); Staples & Herbst 2005:741 (KEY), 748 (DESCR)
Urochloa maxima (Jacq.) R.D.Webster
nat
Mi
Wagner et al. 1990:1569 (as Panicum maximum, no NWHI records); Starr et al.
2002:22 (Mi); Zuloaga et al. 2003:632/Wagner et al. 2012:97 (NOM:Syn. P.
maximum)
Urochloa mutica (Forssk.) T.Q.Nguyen
nat
Mi
Wagner et al. 1990:1504 (as Brachiaria mutica, Mi); Snow & Lau 2010:49 (retain
separation of Brachiaria from Urochloa); Zuloaga et al. 2003:632/Wagner &
Herbst 2003:58/Clayton & Snow 2010:61 (NOM:Syn. B. mutica)
Wagner et al. 1990:1586 (Mi)
FF
N
FF
Wagner et al. 1990:1593 (Ku, Mi, PH, FF, N)
Wagner et al. 1990:1596 (Ku?, FF? [adventive]); Wagner & Herbst 1995:24 (Ku,
La, FF); Starr et al. 2010:66 (Mi)
Wagner et al. 1990:1597 (Mi, La)
Pteridophyte
Nephrolepidaceae
Nephrolepis brownii (Desv.) Hovenkamp &
Miyam.
nat
N
Palmer 2003:193 (as Nephrolepis multiflora, N); Staples & Herbst 2005:40 (KEY,
as N. multiflora), 41 (DESCR); Hovenkamp & Miyamoto 2005:293 (NOM:Syn.
N. multiflora)
Psilotaceae
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 16
Scientific Name
Status Ku Mi PH Li La GP FF Ne N
Citation
Pteridophyte
Psilotaceae
Psilotum nudum (L.) P.Beauv.
ind
Mi
Palmer 2003:272; Staples & Herbst 2005:46 (DESCR); Kennedy et al. 2010:23
(Mi)
Islands: Ku=Kure; Mi=Midway; PH=Pearl Hermes; Li=Lisianski; La=Laysan; GP=Gardner Pinnacles; FF=French Frigate Shoals; Ne=Necker; N=Nihoa
Status: end=endemic; ind=indigenous; nat=naturalized; pol=Polynesian
B- 17
Synonymy List Arranged By Family
Appendix C. Synonymy list for Hawaiian native and naturalized vascular plant checklist, arranged alphabetically by major group and family
This appendix compiles scientific names that have changed for native and
naturalized vascular plant subsequent to publication of the Manual of the Flowering
Plants of Hawai‘i (Wagner et al. 1990) and Hawai‘i's Ferns and Fern Allies (Palmer
2003). The list includes both taxonomic and nomenclatural synonyms, misapplied
names, and corrected misspellings. It is arranged in a nested alphabetical sequence
by major group, family, and currently accepted scientific name.
The literature citation provides a source for the name change; while primary
sources were preferred, secondary sources are often cited. In some cases, both
Group
Family
primary and secondary sources are cited; these are denoted by a slash symbol
between the two citations. Occasionally, a name has been changed, and later
changed again [e.g., Blechum brownei (Acanthaceae) became a synonym of B.
pyramidatum, which later was synonymized into Ruellia blechum]. In such cases, the
connection between the oldest and currently accepted names may not be apparent,
and citations, separated by a semicolon, are provided for the name change to an
intermediate name, followed by one for the current disposition.
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Acanthaceae
Odontonema cuspidatum (Nees) Kuntze
Odontonema strictum sensu Wagner et al. (1990), non
(Nees) Kuntze
Dicot
Acanthaceae
Pseuderanthemum variabile (R.Br.) Radlk.
Pseuderanthemum fasciculatum sensu Herbarium
Pacificum Staff (1998), non (Oerst.) Leonard
Herbst & Wagner 1999:12
Lorence et al. 1995:20
Dicot
Acanthaceae
Ruellia blechum L.
Blechum brownei Juss.
Wagner & Herbst 1995:14; Tripp
et al. 2009:900/Wagner et al.
2012:3
Dicot
Acanthaceae
Ruellia blechum L.
Blechum pyramidatum (Lam.) Urb.
Tripp et al. 2009:900/Wagner et
al. 2012:3
Dicot
Acanthaceae
Ruellia brevifolia (Pohl) C.Ezcurra
Ruellia graecizans Backer
Ezcurra 1993:802/Herbst &
Wagner 1999:13
Dicot
Apocynaceae
Alyxia stellata (J.R.Forst. & G.Forst.) Roem. & Schult.
Alyxia oliviformis Gaudich.
Middleton 2002:56
Dicot
Apocynaceae
Thevetia peruviana (Pers.) K.Schum.
Cascabela peruviana (Pers.) Raf.
Wagner & Herbst 1999:1858
Dicot
Araliaceae
Polyscias bisattenuata (Sherff) Lowry & G.M.Plunkett
Tetraplasandra bisattenuata Sherff
Lowry & Plunkett 2010:73
Dicot
Araliaceae
Polyscias flynnii (Lowry & K.R.Wood) Lowry &
G.M.Plunkett
Tetraplasandra flynnii Lowry & K.R.Wood
Lowry & Plunkett 2010:74
Dicot
Araliaceae
Polyscias gymnocarpa (Hillebr.) Lowry &
G.M.Plunkett
Tetraplasandra gymnocarpa (Hillebr.) Sherff
Lowry & Plunkett 2010:74
Dicot
Araliaceae
Polyscias hawaiensis (A.Gray) Lowry & G.M.Plunkett
Tetraplasandra hawaiensis A.Gray
Lowry & Plunkett 2010:74
Dicot
Araliaceae
Polyscias kavaiensis (H.Mann) Lowry & G.M.Plunkett
Tetraplasandra kavaiensis (H.Mann) Sherff
Lowry & Plunkett 2010:74
Dicot
Araliaceae
Polyscias lydgatei (Hillebr.) Lowry & G.M.Plunkett
Tetraplasandra lydgatei (Hillebr.) Harms
Lowry & Plunkett 2010:74
Dicot
Araliaceae
Polyscias oahuensis (A.Gray) Lowry & G.M.Plunkett
Tetraplasandra oahuensis (A.Gray) Harms
Lowry & Plunkett 2010:74
Dicot
Araliaceae
Polyscias racemosa (C.N.Forbes) Lowry &
G.M.Plunkett
Munroidendron racemosum (C.N.Forbes) Sherff
Lowry & Plunkett 2010:74
C-1
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Araliaceae
Polyscias sandwicensis (A.Gray) Lowry &
G.M.Plunkett
Dicot
Araliaceae
Polyscias waialealae (Rock) Lowry & G.M.Plunkett
Tetraplasandra waialealae Rock
Lowry & Plunkett 2010:75
Dicot
Araliaceae
Polyscias waimeae (Wawra) Lowry & G.M.Plunkett
Tetraplasandra waimeae Wawra
Lowry & Plunkett 2010:75
Dicot
Araliaceae
Schefflera heptaphylla (L.) Frodin
Schefflera taiwaniana sensu Daehler & Baker (2006),
non (Nakai) Kaneh.
Frohlich & Lau 2012:28
Dicot
Asclepiadaceae
Calotropis procera (Aiton) W.T.Aiton
Calotropis gigantea sensu Starr et al. (2003, 2004), in
part, non (L.) W.T.Aiton
Starr & Starr 2012:87
Dicot
Asclepiadaceae
Stapelia gigantea N.E.Br.
Stapelia nobilis N.E.Br.
Staples & Herbst 2005:144
King & Robinson 1987:60/
Herbst & Wagner 1999:14
Reynoldsia sandwicensis A.Gray
Lowry & Plunkett 2010:74
Dicot
Asteraceae
Adenostemma viscosum J.R.Forst. & G.Forst.
Adenostemma lavenia sensu Wagner et al. (1990), non
(L.) Kuntze
Dicot
Asteraceae
Cyanthillium cinereum (L.) H.Rob.
Vernonia cinerea (L.) Less. var. parviflora (Reinw.)
DC.
Wagner et al. 1997:52
Dicot
Asteraceae
Delairea odorata Lem.
Senecio mikanioides Otto ex Walp.
Jeffrey 1986:933/Herbst &
Wagner 1999:14
Dicot
Asteraceae
Dubautia waianapanapaensis G.D.Carr
Dubautia dolosa (O.Deg. & Sherff) G.D.Carr
Carr 1995:1580/Wagner & Herbst
2003:7
Dicot
Asteraceae
Eclipta prostrata (L.) L.
Eclipta alba (L.) Hassk.
Wagner & Herbst 1995:16
Dicot
Asteraceae
Euchiton japonicus (Thunb.) A.Anderb.
Gnaphalium japonicum Thunb.
Wagner et al. 1997:54
Dicot
Asteraceae
Euchiton sphaericus (Willd.) A.Anderb.
Gnaphalium japonicum sensu Wagner et al. (1990),
non Euchiton japonicus (Thunb.) A.Anderb.
Wagner et al. 1997:54
Dicot
Asteraceae
Gamochaeta purpurea (L.) Cabrera
Gnaphalium purpureum L.
Wagner et al. 1997:54
Dicot
Asteraceae
Kalimeris indica (L.) Sch.Bip. subsp. indica
Boltonia indica (L.) Benth.
Gu & Hoch 1997:785/Herbst &
Wagner 1999:14
Dicot
Asteraceae
Keysseria erici (C.N.Forbes) Cabrera
Lagenifera [Lagenophora] erici C.N.Forbes
Nicolson 1996:341; Nesom
2000:58/Nesom 2001b:514
Dicot
Asteraceae
Keysseria helenae (C.N.Forbes & Lydgate) Cabrera
Lagenifera [Lagenophora] helenae C.N.Forbes &
Lydgate
Nicolson 1996:341; Nesom
2000:58/Nesom 2001b:514
Dicot
Asteraceae
Keysseria maviensis (H.Mann) Cabrera
Lagenifera [Lagenophora] maviensis H.Mann
Nicolson 1996:341; Nesom
2000:58/Nesom 2001b:514
Dicot
Asteraceae
Lactuca sativa L.
Lactuca serriola L.
Koopman et al. 2001
Dicot
Asteraceae
Leucanthemum vulgare Lam.
Chrysanthemum leucanthemum L.
Soreng & Cope 1991:162/Herbst
& Wagner 1999:15
Dicot
Asteraceae
Lipochaeta connata (Gaudich.) DC. subsp. acris
(Sherff) W.L.Wagner & H.Rob.
Lipochaeta connata (Gaudich.) DC. var. acris (Sherff)
R.C.Gardner
Wagner & Robinson 2002:545
C-2
Synonymy List Arranged By Family
Group
Dicot
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Asteraceae
Lipochaeta connata (Gaudich.) DC. subsp. connata
Lipochaeta connata (Gaudich.) DC. var. connata
Wagner & Robinson 2002:545
Dicot
Asteraceae
Lipochaeta lobata (Gaudich.) DC. subsp. leptophylla
(O.Deg. & Sherff) W.L.Wagner & H.Rob.
Lipochaeta lobata (Gaudich.) DC. var. leptophylla
O.Deg. & Sherff
Wagner & Robinson 2002:547
Dicot
Asteraceae
Lipochaeta lobata (Gaudich.) DC. subsp. lobata
Lipochaeta lobata (Gaudich.) DC. var. lobata
Wagner & Robinson 2002:546
Dicot
Asteraceae
Logfia gallica (L.) Coss. & Germ.
Filago gallica L.
Wagner et al. 1997:54
Dicot
Asteraceae
Melanthera bryanii (Sherff) W.L.Wagner & H.Rob.
Lipochaeta bryanii Sherff
Wagner & Robinson 2002:552
Dicot
Asteraceae
Melanthera fauriei (H.Lév.) W.L.Wagner & H.Rob.
Lipochaeta fauriei H.Lév.
Wagner & Robinson 2002:552
Dicot
Asteraceae
Melanthera integrifolia (Nutt.) W.L.Wagner & H.Rob.
Lipochaeta integrifolia (Nutt.) Gray
Wagner & Robinson 2002:552
0Dicot
Asteraceae
Melanthera kamolensis (O.Deg. & Sherff)
W.L.Wagner & H.Rob.
Lipochaeta kamolensis O.Deg. & Sherff
Wagner & Robinson 2002:552
Asteraceae
Melanthera lavarum (Gaudich.) W.L.Wagner &
H.Rob.
Lipochaeta lavarum (Gaudich.) DC.
Wagner & Robinson 2002:553
Dicot
Asteraceae
Melanthera micrantha (Nutt.) W.L.Wagner & H.Rob.
subsp. exigua (O.Deg. & Sherff) W.L.Wagner &
H.Rob.
Lipochaeta micrantha (Nutt.) A.Gray var. exigua
(O.Deg. & Sherff) R.C.Gardner
Wagner & Robinson 2002:554
Dicot
Asteraceae
Melanthera micrantha (Nutt.) W.L.Wagner & H.Rob.
subsp. micrantha
Lipochaeta micrantha (Nutt.) A.Gray var. micrantha
Wagner & Robinson 2002:554
Dicot
Asteraceae
Melanthera perdita (Sherff) W.L.Wagner & H.Rob.
Lipochaeta perdita Sherff
Wagner & Robinson 2002:554
Dicot
Asteraceae
Melanthera populifolia (Sherff) W.L.Wagner &
H.Rob.
Lipochaeta populifolia (Sherff) R.C.Gardner
Wagner & Robinson 2002:554
Dicot
Asteraceae
Melanthera remyi (A.Gray) W.L.Wagner & H.Rob.
Lipochaeta remyi A.Gray
Wagner & Robinson 2002:554
Dicot
Asteraceae
Melanthera subcordata (A.Gray) W.L.Wagner &
H.Rob.
Lipochaeta subcordata A.Gray
Wagner & Robinson 2002:554
Dicot
Asteraceae
Melanthera tenuifolia (A.Gray) W.L.Wagner &
H.Rob.
Lipochaeta tenuifolia A.Gray
Wagner & Robinson 2002:554
Dicot
Asteraceae
Melanthera tenuis (O.Deg. & Sherff) W.L.Wagner &
H.Rob.
Lipochaeta tenuis O.Deg. & Sherff
Wagner & Robinson 2002:555
Dicot
Asteraceae
Melanthera venosa (Sherff) W.L.Wagner & H.Rob.
Lipochaeta venosa Sherff
Wagner & Robinson 2002:555
Dicot
Asteraceae
Melanthera waimeaensis (H.St.John) W.L.Wagner &
H.Rob.
Lipochaeta waimeaensis H.St.John
Wagner & Robinson 2002:555
Dicot
Asteraceae
Pluchea carolinensis (Jacq.) G.Don
Pluchea symphytifolia sensu Wagner et al. (1990), non
(Mill.) Gillis
Wagner & Herbst 1995:16/ Khan
& Jarvis 1989
Dicot
Asteraceae
Pseudelephantopus spicatus (B.Juss. ex Aubl.)
C.F.Baker
Elephantopus spicatus Juss. ex Aubl.
Herbst & Wagner 1999:15
Dicot
C-3
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Asteraceae
Pseudognaphalium sandwicensium (Gaudich.)
A.Anderb. var. hawaiiense (O.Deg. & Sherff)
W.L.Wagner
Gnaphalium sandwicensium Gaudich. var. hawaiiense
O.Deg. & Sherff
Wagner et al. 1997:54
Asteraceae
Pseudognaphalium sandwicensium (Gaudich.)
A.Anderb. var. kilaueanum (O.Deg. & Sherff)
W.L.Wagner
Gnaphalium sandwicensium Gaudich. var. kilaueanum
O.Deg. & Sherff
Wagner et al. 1997:54
Asteraceae
Pseudognaphalium sandwicensium (Gaudich.)
A.Anderb. var. molokaiense (O.Deg. & Sherff)
W.L.Wagner
Gnaphalium sandwicensium Gaudich. var.
molokaiense O.Deg. & Sherff
Wagner et al. 1997:54
Dicot
Asteraceae
Pseudognaphalium sandwicensium (Gaudich.)
A.Anderb. var. sandwicensium
Gnaphalium sandwicensium Gaudich. var.
sandwicensium
Wagner et al. 1997:54
Dicot
Asteraceae
Pseudogynoxys chenopodioides (Kunth) Cabrera
Senecio confusus Britten
Staples & Herbst 2005:163
Dicot
Asteraceae
Sphagneticola trilobata (L.) Pruski
Wedelia trilobata (L.) A.S.Hitchc.
Wagner et al. 1997:55
Dicot
Asteraceae
Symphyotrichum expansum (Poepp. ex Spreng.)
G.L.Nesom
Aster subulatus Michx. var. sandwicensis (A.Gray)
A.G.Jones
Nesom 1994:292; Sundberg
2004:907; Nesom 2005:2133
Dicot
Asteraceae
Symphyotrichum expansum (Poepp. ex Spreng.)
G.L.Nesom
Symphyotrichum squamatum sensu Nesom (1994),
non (Spreng.) Nesom
Sundberg 2004:907; Nesom
2005:2133
Dicot
Asteraceae
Symphyotrichum expansum (Poepp. ex Spreng.)
G.L.Nesom
Symphyotrichum subulatum (Michx.) Nesom var.
parviflorum (Nees) Sundberg
Nesom 2005:2133
Dicot
Asteraceae
Tanacetum parthenium (L.) Sch.Bip.
Chrysanthemum parthenium (L.) Bernh.
Herbst & Wagner 1999:15
Dicot
Asteraceae
Zinnia palmeri A.Gray
Zinnia maritima Kunth [Note: not a syn., but Z.
palmeri sometimes considered a syn. of Z. maritima
Staples et al. 2002:6; Wagner &
Herbst 2003:11
Dicot
Balsaminaceae
Impatiens sodeni Engl. & Warb. ex Engl.
Impatiens oliveri C.Wright ex W.Wats.
Grey-Wilson 1982:20/Herbst &
Wagner 1999:15
Dicot
Balsaminaceae
Impatiens walleriana Hook.f
Impatiens wallerana Hook.f (spelling change)
Staples & Herbst 2005:172
Dicot
Begoniaceae
Begonia foliosa Kunth
Begonia foliosa Kunth var. miniata sensu Wagner et
al. (1990), non (Planch.) L.B.Sm. & B.G.Shubert
Imada 2007:35
Dicot
Begoniaceae
Begonia vitifolia Schott
Begonia reniformis sensu Wagner et al. (1990), non
Dryand.
Staples et al. 2002:6
Dicot
Brassicaceae
Brassica rapa L.
Brassica campestris L.
Al-Shehbaz 1985:290/Wagner &
Herbst 1995:17
Dicot
Brassicaceae
Capsella bursa-pastoris (L.) Medik.
Capsella rubella Reut.
Wagner & Herbst 1995:17
Lorence et al. 1995:26
Wagner & Herbst 1995:17
Dicot
Dicot
Dicot
Dicot
Brassicaceae
Hirschfeldia incana (L.) Lagr.-Foss.
Brassica nigra sensu Wagner et al. (1990), in part, non
(L.) W.D.J.Koch
Dicot
Brassicaceae
Lepidium africanum (Burm.f.) DC.
Lepidium hyssopifolium sensu Wagner et al. (1990),
non Desv.
C-4
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Brassicaceae
Nasturtium microphyllum Boenn. ex Rchb.
Rorippa microphylla (Boenn. ex Reichenb.) Hyl. ex
A.& D.Löve
Al-Shehbaz & Price
1998:124/Herbst & Wagner
1999:15
Dicot
Brassicaceae
Rorippa sarmentosa (G.Forst. ex DC.) J.F.Macbr.
Nasturtium sarmentosum (G.Forst. ex DC.) Schinz &
Guillaumin
Wagner & Herbst 1995:18
Dicot
Brassicaceae
Sinapis arvensis L.
Brassica campestris sensu Wagner et al. (1990), in
part, non L.
Herbarium Pacificum Staff
1998:9
Dicot
Cactaceae
Opuntia monacantha (Willd.) Haw.
Opuntia vulgaris sensu Wagner et al. (1990), non Mill.
Staples et al. 2002:6
Dicot
Cactaceae
Selenicereus macdonaldiae (Hook.) Britton & Rose
Selenicereus grandiflorus sensu Lorence et al. (1995),
non (L.) Britton & Rose
Herbst & Wagner 1999:16/
Staples & Herbst 2005:215
Dicot
Campanulaceae
Clermontia arborescens (H.Mann) Hillebr. subsp.
waikoluensis (H.St.John) Lammers
Clermontia waikoluensis H.St.John
Lammers 1991:25
Dicot
Campanulaceae
Clermontia peleana Rock subsp. peleana
Clermontia peleana Rock
Lammers 1991:34
Dicot
Campanulaceae
Clermontia peleana Rock subsp. singuliflora (Rock)
Lammers
Clermontia singuliflora (Rock) Rock
Lammers 1991:32
Lammers 1998:32/Herbst &
Wagner 1999:16
Lammers 1998:32/Herbst &
Wagner 1999:17
Dicot
Campanulaceae
Cyanea calycina (Cham.) Lammers
Cyanea lanceolata (Gaudich.) Lammers, Givnish &
Sytsma subsp. calycina (Cham.) Lammers, Givnish &
Sytsma
Dicot
Campanulaceae
Cyanea calycina (Cham.) Lammers
Rollandia lanceolata Gaudich. subsp. calycina (Cham.)
Lammers
Dicot
Campanulaceae
Cyanea coriacea (A.Gray) Hillebr.
Cyanea fauriei H.Lév.
Lammers 2004:96
Dicot
Campanulaceae
Cyanea crispa (Gaudich.) Lammers, Givnish & Sytsma
Rollandia crispa Gaudich.
Lammers 1993:439/Herbst &
Wagner 1999:17
Dicot
Campanulaceae
Cyanea cylindrocalyx (Rock) Lammers
Cyanea grimesiana Gaudich. subsp. cylindrocalyx
(Rock) Lammers
Lammers 1998:31/Herbst &
Wagner 1999:16
Dicot
Campanulaceae
Cyanea dunbariae Rock
Cyanea dunbarii Rock (spelling change)
Herbst & Wagner 1999:16
Dicot
Campanulaceae
Cyanea eleeleensis (H.St.John) Lammers
Delissea eleeleensis H.St.John
Lammers 1992:129/Herbst &
Wagner 1999:16
Dicot
Campanulaceae
Cyanea floribunda E.Wimm.
Cyanea degeneriana sensu Wagner et al. (1990), non
E.Wimm.
Wagner & Herbst 2003:15
Dicot
Campanulaceae
Cyanea habenata (H.St.John) Lammers
Delissea habenata H.St.John
Lammers 1998:33/Herbst &
Wagner 1999:16
Dicot
Campanulaceae
Cyanea hirtella (H.Mann) Hillebr.
Cyanea degeneriana E.Wimm.
Wagner & Herbst 2003:15
Dicot
Campanulaceae
Cyanea hirtella (H.Mann) Hillebr.
Cyanea sylvestris A.Heller
Wagner et al. 2005– website
Dicot
Campanulaceae
Cyanea humboldtiana (Gaudich.) Lammers, Givnish &
Sytsma
Rollandia humboldtiana Gaudich.
Lammers 1993:439/Herbst &
Wagner 1999:17
C-5
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Campanulaceae
Cyanea kahiliensis (H.St.John) Lammers
Cyanea spathulata (Hillebr.) A.Heller subsp.
longipetiolata Lammers
Dicot
Campanulaceae
Cyanea kahiliensis (H.St.John) Lammers
Delissea kahiliensis H.St.John
Lammers 1998:33/Herbst &
Wagner 1999:16
Dicot
Campanulaceae
Cyanea kolekoleensis (H.St.John) Lammers
Delissea kolekoleensis H.St.John
Lammers 1992:130/Herbst &
Wagner 1999:16
Dicot
Campanulaceae
Cyanea koolauensis Lammers, Givnish & Sytsma
Rollandia angustifolia (Hillebr.) Rock
Lammers 1993:439/Herbst &
Wagner 1999:17
Dicot
Campanulaceae
Cyanea lanceolata (Gaudich.) Lammers, Givnish &
Sytsma
Rollandia lanceolata Gaudich. subsp. lanceolata
Lammers 1993:439/Herbst &
Wagner 1999:17
Dicot
Campanulaceae
Cyanea lobata H.Mann subsp. baldwinii (C.N.Forbes
& G.C.Munro) H.St.John
Cyanea baldwinii C.N.Forbes & G.C.Munro
Lammers 1999:387
Dicot
Campanulaceae
Cyanea lobata H.Mann subsp. lobata
Cyanea lobata H.Mann
Lammers 1999:387
Dicot
Campanulaceae
Cyanea longiflora (Wawra) Lammers, Givnish &
Sytsma
Rollandia longiflora Wawra
Lammers 1993:439/Herbst &
Wagner 1999:17
Dicot
Campanulaceae
Cyanea mauiensis (Rock) Lammers
Cyanea grimesiana Gaudich. var. mauiensis Rock
Lammers 1998:31/Herbst &
Wagner 1999:17
Dicot
Campanulaceae
Cyanea munroi (Hosaka) Lammers
Cyanea grimesiana Gaudich. var. munroi Hosaka
Lammers 1998:31/Herbst &
Wagner 1999:17
Dicot
Campanulaceae
Cyanea parvifolia (C.N.Forbes) Lammers, Givnish &
Sytsma
Rollandia parvifolia C.N.Forbes
Lammers 1993:439/Herbst &
Wagner 1999:17
Dicot
Campanulaceae
Cyanea purpurellifolia (Rock) Lammers, Givnish &
Sytsma
Rollandia purpurellifolia Rock
Lammers 1993:439/Herbst &
Wagner 1999:17
Dicot
Campanulaceae
Cyanea rivularis Rock
Delissea rivularis (Rock) E.Wimm.
Lammers 2005:62
Dicot
Campanulaceae
Cyanea sessilifolia (O.Deg.) Lammers
Rollandia sessilifolia O.Deg.
Lammers 1998:32/Herbst &
Wagner 1999:18
Dicot
Campanulaceae
Cyanea spathulata (Hillebr.) A.Heller
Cyanea spathulata (Hillebr.) A.Heller subsp. spathulata
Lammers 2004:100
Dicot
Campanulaceae
Cyanea st.-johnii (Hosaka) Lammers, Givnish &
Sytsma
Rollandia st.-johnii Hosaka
Lammers 1993:440/Herbst &
Wagner 1999:18
Dicot
Campanulaceae
Delissea kauaiensis (Lammers) Lammers
Delissea niihauensis H.St.John subsp. kauaiensis
(Lammers) Lammers
Lammers 2005:19
Dicot
Campanulaceae
Delissea kauaiensis (Lammers) Lammers
Delissea undulata Gaudich. subsp. kauaiensis
Lammers
Lammers 2005:19
Dicot
Campanulaceae
Delissea lanaiensis (Rock) Lammers
Delissea sinuata Hillebr. subsp. lanaiensis (Rock)
Lammers
Lammers 1998:34/Herbst &
Wagner 1999:18
Dicot
Campanulaceae
Delissea niihauensis H.St.John
Delissea niihauensis H.St.John subsp. niihauensis
Lammers 2005:17
C-6
Lammers 1998:33/Herbst &
Wagner 1999:16
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Campanulaceae
Delissea niihauensis H.St.John
Delissea undulata Gaudich. subsp. niihauensis
(H.St.John) Lammers
Dicot
Campanulaceae
Delissea sinuata Hillebr.
Delissea sinuata Hillebr. subsp. sinuata
Lammers 1998:34
Dicot
Campanulaceae
Delissea subcordata Gaudich. subsp. obtusifolia
(Wawra) Lammers
Delissea subcordata Gaudich. var. obtusifolia Wawra
Lammers 2005:34
Dicot
Campanulaceae
Delissea subcordata Gaudich. subsp. subcordata
Delissea subcordata Gaudich.
Lammers 2005:33
Dicot
Campanulaceae
Delissea undulata Gaudich.
Delissea undulata Gaudich. subsp. undulata
Lammers 2005:21
Campanulaceae
Lobelia dunbariae Rock subsp. dunbariae
Lobelia dunbarii Rock subsp. dunbarii (spelling
change)
Wagner et al. 2005– website
Dicot
Campanulaceae
Lobelia dunbariae Rock subsp. paniculata (Rock)
Lammers
Lobelia dunbarii Rock subsp. paniculata (Rock)
Lammers (spelling change)
Wagner et al. 2005– website
Dicot
Campanulaceae
Lobelia gaudichaudii A.DC.
Lobelia gaudichaudii A.DC. subsp. gaudichaudii
Lammers 2007:804
Dicot
Campanulaceae
Lobelia gloria-montis Rock subsp. gloria-montis
Lobelia gloria-montis Rock
Lammers 2007:805
Dicot
Campanulaceae
Lobelia gloria-montis Rock subsp. longibracteata
(Rock) Lammers
Lobelia gloria-montis Rock var. longibracteata (Rock)
Rock
Lammers 2007:806
Dicot
Campanulaceae
Lobelia xkauaiensis (A.Gray) A.Heller
Lobelia kauaensis (A.Gray) A.Heller (spelling change)
Lammers 2007:801
Lammers 2007:803
Dicot
Lammers 2005:17
Dicot
Campanulaceae
Lobelia koolauensis (Hosaka & Fosberg) Lammers
Lobelia gaudichaudii A.DC. subsp. koolauensis
(Hosaka & Fosberg) Lammers
Dicot
Campanulaceae
Lobelia niihauensis H.St.John
Neowimmeria niihauensis (H.St.John) O.Deg. &
I.Deg. var. forbesii (H.St.John) O.Deg. & I.Deg.
Wagner & Herbst 2003:16
Dicot
Campanulaceae
Trematolobelia kaalae (O.Deg. ) Lammers
Trematolobelia macrostachys (Hook. & Arn.) Zahlbr.
var. kaalae (O.Deg.) E.Wimm.
Lammers 2009:134
Dicot
Campanulaceae
Trematolobelia wimmeri O.Deg. & I.Deg.
Trematolobelia lustrialis H.St.John
Lammers 2009:140
Dicot
Caryophyllaceae
Schiedea diffusa A.Gray subsp. diffusa
Schiedea diffusa A.Gray
Wagner et al. 2005:103
Dicot
Caryophyllaceae
Schiedea diffusa A.Gray subsp. macraei (Sherff)
W.L.Wagner & Weller
Schiedea diffusa A.Gray var. macraei Sherff
Wagner et al. 2005:106
Dicot
Caryophyllaceae
Schiedea lychnoides Hillebr.
Alsinidendron lychnoides (Hillebr.) Sherff
Wagner et al. 2005:51
Dicot
Caryophyllaceae
Schiedea obovata (Sherff) W.L.Wagner & Weller
Alsinidendron obovatum Sherff
Wagner et al. 2005:57
Dicot
Caryophyllaceae
Schiedea pentandra W.L.Wagner & E.M.Harris
Schiedea pubescens Hillebr. var. purpurascens Sherff,
invalid name
Wagner & Harris 2000:159
Dicot
Caryophyllaceae
Schiedea pubescens Hillebr.
Schiedea pubescens Hillebr. var. pubescens
Wagner et al. 2005:99
Dicot
Caryophyllaceae
Schiedea spergulina A.Gray
Schiedea spergulina A.Gray var. leiopoda Sherff
Wagner et al. 2005:127
Dicot
Caryophyllaceae
Schiedea spergulina A.Gray
Schiedea spergulina A.Gray var. spergulina
Wagner et al. 2005:127
C-7
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Caryophyllaceae
Schiedea trinervis (H.Mann) Pax & K.Hoffm.
Alsinidendron trinerve H.Mann
Wagner et al. 2005:60
Dicot
Caryophyllaceae
Schiedea viscosa H.Mann
Alsinidendron viscosum (H.Mann) Sherff
Wagner et al. 2005:54
Dicot
Chenopodiaceae
Dysphania ambrosioides (L.) Mosyakin & Clemants
Chenopodium ambrosioides L.
Clemants & Mosyakin 2003:
270/Wagner et al. 2012:28
Dicot
Chenopodiaceae
Dysphania carinata (R.Br.) Mosyakin & Clemants
Chenopodium carinatum R.Br.
Clemants & Mosyakin 2003:
274/Wagner et al. 2012:28
Dicot
Chenopodiaceae
Salsola tragus L.
Salsola kali sensu Wagner et al. (1990), non L.
Wagner et al. 1997:55
Dicot
Clusiaceae
Hypericum mutilum L. subsp. mutilum
Hypericum mutilum L.
Robson 1990:116/Wagner &
Herbst 1995:18
Dicot
Convolvulaceae
Jacquemontia sandwicensis A.Gray
Jacquemontia ovalifolia (Vahl ex West) Hallier f.
subsp. sandwicensis (A.Gray) K.R.Robertson
Namoff et al. 2010
Bryophyllum beauverdii (Raym.-Hamet) A.Berger
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst
2003:21
Bryophyllum daigremontianum (Raym.-Hamet &
H.Perrier) A.Berger
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst
2003:21
Bryophyllum fedtschenkoi (Raym.-Hamet &
H.Perrier) Lauz.-March.
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst
2003:21
Bryophyllum pinnatum (Lam.) Oken
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst
2003:21
Kalanchoe tubiflora (Harv.) Raym.-Hamet
Bryophyllum tubiflorum Harv.
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst
2003:21
Dicot
Dicot
Dicot
Dicot
Crassulaceae
Crassulaceae
Crassulaceae
Crassulaceae
Kalanchoe beauverdii Raym.-Hamet
Kalanchoe daigremontiana Raym.-Hamet & H.Perrier
Kalanchoe fedtschenkoi Raym.-Hamet & H.Perrier
Kalanchoe pinnata (Lam.) Pers.
Dicot
Crassulaceae
Dicot
Cucurbitaceae
Sicyos albus (H.St.John) I.Telford
Sicyos alba (H.St.John) I.Telford (spelling change)
Wagner et al. 2012:30
Dicot
Cucurbitaceae
Sicyos lanceoloideus (H.St.John) W.L.Wagner &
D.R.Herbst
Sicyocarya lanceoloidea H.St.John
Wagner & Shannon 1999:444
Dicot
Cucurbitaceae
Sicyos lanceoloideus (H.St.John) W.L.Wagner &
D.R.Herbst
Sicyos sp. A
Wagner & Shannon 1999:444
Dicot
Cucurbitaceae
Sicyos macrophyllus A.Gray
Sicyocarya kipahuluensis H.St.John
Wagner & Shannon 1999:444
C-8
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Epacridaceae
Leptecophylla tameiameiae (Cham. & Schltdl.)
C.M.Weiller
Styphelia tameiameiae (Cham. & Schltdl.) F.Muell.
Weiller 1999/Staples et al.
2003:11
Dicot
Euphorbiaceae
Euphorbia albomarginata Torr. & A.Gray
Chamaesyce albomarginata (Torr. & A.Gray) Small
Govaerts et al. 2000:667
Dicot
Euphorbiaceae
Euphorbia arnottiana Endl.
Chamaesyce arnottiana (Endl.) O.Deg. & I.Deg.
Govaerts et al. 2000:674
Dicot
Euphorbiaceae
Euphorbia atrococca A.Heller
Chamaesyce atrococca (A.Heller) Croizat & O.Deg.
Govaerts et al. 2000:676
Dicot
Euphorbiaceae
Euphorbia celastroides Boiss. var. amplectens Sherff
Chamaesyce celastroides (Boiss.) Croizat & O.Deg.
var. amplectens (Sherff) O.Deg. & I.Deg.
Govaerts et al. 2000:692
Govaerts et al. 2000:692
Dicot
Euphorbiaceae
Euphorbia celastroides Boiss. var. celastroides
Chamaesyce celastroides (Boiss.) Croizat & O.Deg.
var. celastroides
Dicot
Euphorbiaceae
Euphorbia celastroides Boiss. var. hanapepensis Sherff
Chamaesyce celastroides (Boiss.) Croizat & O.Deg.
var. hanapepensis (Sherff) O.Deg. & I.Deg.
Govaerts et al. 2000:692
Dicot
Euphorbiaceae
Euphorbia celastroides Boiss. var. kaenana Sherff
Chamaesyce celastroides (Boiss.) Croizat & O.Deg.
var. kaenana (Sherff) O.Deg. & I.Deg.
Govaerts et al. 2000:692
Dicot
Euphorbiaceae
Euphorbia celastroides Boiss. var. laehiensis O.Deg.,
I.Deg. & Sherff
Chamaesyce celastroides (Boiss.) Croizat & O.Deg.
var. laehiensis (O.Deg., I.Deg. & Sherff) Koutnik
Govaerts et al. 2000:692
Dicot
Euphorbiaceae
Euphorbia celastroides Boiss. var. lorifolia A.Gray
Chamaesyce celastroides (Boiss.) Croizat & O.Deg.
var. lorifolia (A.Gray) O.Deg. & I.Deg.
Govaerts et al. 2000:692
Dicot
Euphorbiaceae
Euphorbia celastroides Boiss. var. stokesii
(C.N.Forbes) Sherff
Chamaesyce celastroides (Boiss.) Croizat & O.Deg.
var. stokesii (C.N.Forbes) O.Deg. & I.Deg.
Govaerts et al. 2000:692
Dicot
Euphorbiaceae
Euphorbia celastroides Boiss. var. tomentella (Boiss.)
Oudejans
Chamaesyce celastroides (Boiss.) Croizat & O.Deg.
var. tomentella (Boiss.) Koutnik
Govaerts et al. 2000:692
Dicot
Euphorbiaceae
Euphorbia clusiifolia Hook. & Arn.
Chamaesyce clusiifolia (Hook. & Arn.) Arthur
Govaerts et al. 2000:698
Dicot
Euphorbiaceae
Euphorbia degeneri Sherff
Chamaesyce degeneri (Sherff) Croizat & O.Deg.
Govaerts et al. 2000:709
Dicot
Euphorbiaceae
Euphorbia deppeana Boiss.
Chamaesyce deppeana (Boiss.) Millsp.
Govaerts et al. 2000:712
Dicot
Euphorbiaceae
Euphorbia eleanoriae (Lorence & W.L.Wagner)
Govaerts
Chamaesyce eleanoriae Lorence & W.L.Wagner
Govaerts et al. 2000:717
Dicot
Euphorbiaceae
Euphorbia halemanui Sherff
Chamaesyce halemanui (Sherff) Croizat & O.Deg.
Govaerts et al. 2000:742
Dicot
Euphorbiaceae
Euphorbia herbstii (W.L.Wagner) Oudejans
Chamaesyce herbstii W.L.Wagner
Govaerts et al. 2000:745
Dicot
Euphorbiaceae
Euphorbia hirta L.
Chamaesyce hirta (L.) Millsp.
Govaerts et al. 2000:320
Dicot
Euphorbiaceae
Euphorbia hypericifolia L.
Chamaesyce hypericifolia (L.) Millsp.
Govaerts et al. 2000:320
Dicot
Euphorbiaceae
Euphorbia hyssopifolia L.
Chamaesyce hyssopifolia (L.) Small
Govaerts et al. 2000:320
C-9
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Euphorbiaceae
Euphorbia kuwaleana O.Deg. & Sherff
Chamaesyce kuwaleana (O.Deg. & Sherff) O.Deg. &
I.Deg.
Dicot
Euphorbiaceae
Euphorbia maculata L.
Chamaesyce maculata (L.) Small
Govaerts et al. 2000:322
Dicot
Euphorbiaceae
Euphorbia multiformis Hook. & Arn. var. microphylla
Boiss.
Chamaesyce multiformis (Hook. & Arn.) Croizat &
Deg. var. microphylla (Boiss.) O.Deg. & I.Deg.
Govaerts et al. 2000:783
Dicot
Euphorbiaceae
Euphorbia multiformis Hook. & Arn. var. multiformis
Chamaesyce multiformis (Hook. & Arn.) Croizat &
Deg. var. multiformis
Govaerts et al. 2000:783
Dicot
Euphorbiaceae
Euphorbia olowaluana Sherff
Chamaesyce olowaluana (Sherff) Croizat & O.Deg.
Govaerts et al. 2000:793
Dicot
Euphorbiaceae
Euphorbia prostrata Aiton
Chamaesyce prostrata (Aiton) Small
Govaerts et al. 2000:324
Dicot
Euphorbiaceae
Euphorbia remyi A.Gray ex Boiss. var. hanaleiensis
Sherff
Chamaesyce remyi (A.Gray ex Boiss.) Croizat &
O.Deg. var. hanaleiensis (Sherff) O.Deg. & I.Deg.
Govaerts et al. 2000:818
Dicot
Euphorbiaceae
Euphorbia remyi A.Gray ex Boiss. var. kauaiensis O.
Deg. & Sherff
Chamaesyce remyi (A.Gray ex Boiss.) Croizat &
O.Deg. var. kauaiensis (O.Deg. & Sherff) O.Deg. &
I.Deg.
Govaerts et al. 2000:818
Dicot
Euphorbiaceae
Euphorbia remyi A.Gray ex Boiss. var. remyi
Chamaesyce remyi (A.Gray ex Boiss.) Croizat &
O.Deg. var. remyi
Govaerts et al. 2000:818
Dicot
Euphorbiaceae
Euphorbia rockii C.N.Forbes
Chamaesyce rockii (C.N.Forbes) Croizat & O.Deg.
Govaerts et al. 2000:821
Chamaesyce albomarginata sensu Wagner et al.
(1990), in part, non (Torr. & A.Gray) Small
Wagner et al. 1997:56; Govaerts
et al. 2000:832
Govaerts et al. 2000:764
Dicot
Euphorbiaceae
Euphorbia serpens Kunth
Dicot
Euphorbiaceae
Euphorbia serpens Kunth
Chamaesyce serpens (Kunth) Small
Govaerts et al. 2000:832
Govaerts et al. 2000:835
Dicot
Euphorbiaceae
Euphorbia skottsbergii Sherff var. skottsbergii
Chamaesyce skottsbergii (Sherff) Croizat & O.Deg.
var. skottsbergii
Dicot
Euphorbiaceae
Euphorbia skottsbergii Sherff var. vaccinioides Sherff
Chamaesyce skottsbergii (Sherff) Croizat & O.Deg.
var. vaccinioides (Sherff) Koutnik
Govaerts et al. 2000:835
Dicot
Euphorbiaceae
Euphorbia sparsiflora A.Heller
Chamaesyce sparsiflora (A.Heller) Koutnik
Govaerts et al. 2000:837
Dicot
Euphorbiaceae
Euphorbia thymifolia L.
Chamaesyce thymifolia (L.) Millsp.
Govaerts et al. 2000:848
Dicot
Euphorbiaceae
Homalanthus populifolius Graham
Omalanthus populifolius Graham (spelling change)
The Plant List website 2012
Dicot
Euphorbiaceae
Phyllanthus distichus Hook. & Arn.
Phyllanthus distichus Hook. & Arn. var. degeneri
(Sherff) Govaerts & Radcl.-Sm.
Herbst & Wagner 1999:20
Dicot
Euphorbiaceae
Phyllanthus distichus Hook. & Arn.
Phyllanthus distichus Hook. & Arn. var. ellipticus
(Müll.Arg.) Govaerts & Radcl.-Sm.
Herbst & Wagner 1999:20
Dicot
Fabaceae
Canavalia molokaiensis O.Deg., I.Deg. & J.D.Sauer
Canavalia galeata sensu Wysong et al. (2007), non
(Gaudich.) Vogel
Wagner et al. 2012:37
C-10
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Fabaceae
Centrosema molle Mart. ex Benth.
Centrosema pubescens sensu Flynn & Lorence (1998),
non Benth.
Fantz 1996:330/Wagner & Herbst
2003:24
Dicot
Fabaceae
Clitoria ternatea L.
Clitoria ternata L. (spelling change)
Staples & Herbst 2005:313
Dicot
Fabaceae
Desmanthus pernambucanus (L.) Thell.
Desmanthus virgatus (L.) Willd.
Wagner & Herbst 1995:20
Dicot
Fabaceae
Falcataria moluccana (Miq.) Barneby & J.W.Grimes
Paraserianthes falcataria (L.) I. Nielsen
Herbarium Pacificum Staff
1998:10/Staples & Herbst
2005:331
Dicot
Fabaceae
Indigofera spicata Forssk.
Indigofera hendecaphylla sensu Du Puy et al. (1993),
non Jacq.
Du Puy et al. 1993: 731/ Herbst
& Wagner 1999:21
Dicot
Fabaceae
Kummerowia striata (Thunb.) Schindl.
Lespedeza striata (Thunb.) Hook. & Arn.
Staples et al. 2002:10
Dicot
Fabaceae
Mezoneuron kavaiense (H.Mann) Hillebr.
Caesalpinia kavaiensis H.Mann
Bruneau et al. 2008:711/ Wagner
et al. 2012:37
Dicot
Fabaceae
Mucuna gigantea (Willd.) DC. subsp. gigantea
Mucuna gigantea (Willd.) DC.
Wilmot-Dear 1990:5/Wagner &
Herbst 1995:21
Dicot
Fabaceae
Mucuna sloanei Fawc. & Rendle var. sloanei
Mucuna urens sensu Wagner et al. (1990), non (L.)
Medik.
Wilmot-Dear 1990:25/Wagner &
Herbst 1995:20
Dicot
Fabaceae
Neonotonia wightii (Wight & Arn.) Lackey
Glycine wightii (Wight & Arn.) Verdc.
Herbarium Pacificum Staff
1996:4
Dicot
Fabaceae
Pueraria montana (Lour.) Merr. var. chinensis (Ohwi)
Sanjappa & Pradeep
Pueraria lobata (Willd.) Ohwi var. thomsoni (Benth.)
Maesen
Staples et al. 2002:11
Dicot
Fabaceae
Pueraria montana (Lour.) Merr. var. lobata (Willd.)
Sanjappa & Pradeep
Pueraria lobata (Willd.) Ohwi var. lobata
Herbst & Wagner 1999:21
Dicot
Fabaceae
Senna gaudichaudii (Hook. & Arn.) H.S.Irwin &
Barneby
Senna surattensis (Burm.f.) H.S.Irwin & Barneby
subsp. retusa (Vogel) Randell
Staples & Herbst 2005:326 call
this Senna surattensis (Burm.f.)
H.S.Irwin & Barneby subsp.
retusa (Vogel) Randell
Dicot
Fabaceae
Stylosanthes scabra Vogel
Stylosanthes fruticosa sensu Wagner et al. (1990), non
(Retz.) Alston
Herbst et al. 2004:7
Dicot
Fabaceae
Vachellia farnesiana (L.) Wight & Arn.
Acacia farnesiana (L.) Willd.
Mabberley 2008:890/Wagner et
al. 2012:36
Dicot
Flacourtiaceae
Xylosma hawaiiense Seem.
Drypetes sherffii Govaerts & Radcl.-Sm.
Herbst & Wagner 1999:21
Dicot
Gentianaceae
Schenkia sebaeoides Griseb.
Centaurium sebaeoides (Griseb.) Druce
Mansion 2004:725
Dicot
Geraniaceae
Geranium hillebrandii Aedo & Muñoz Garm.
Geranium humile Hillebr.
Aedo & Garmendia 1997:725/
Herbst & Wagner 1999:21
Dicot
Gesneriaceae
Cyrtandra confertiflora (Wawra) C.B.Clarke
Cyrtandra confertiflora (Wawra) C.B.Clarke var.
confertiflora
Wagner & Herbst 2003:29
C-11
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Gesneriaceae
Cyrtandra heinrichii H.St.John
Cyrtandra confertiflora (Wawra) C.B.Clarke var.
obovata (Wawra) C.B.Clarke
Dicot
Gesneriaceae
Cyrtandra kealiae Wawra subsp. kealiae
Cyrtandra limahuliensis H.St.John
Wagner & Lorence 2000:17
Dicot
Goodeniaceae
Scaevola taccada (Gaertn.) Roxb.
Scaevola sericea Vahl
Green 1991:121/Shannon et al.
1997:801
Dicot
Lamiaceae
Leonurus japonicus Houtt.
Leonurus sibiricus sensu Wagner et al. (1990), non L.
Harley & Paton 2001:244/
Staples et al. 2003:14
Dicot
Lamiaceae
Mentha xspicata L.
Mentha spicata L.
Staples & Herbst 2005:358
Staples et al. 2003:14
Wagner & Herbst 2003:29
Dicot
Lamiaceae
Mentha suaveolens Ehrh.
Mentha xrotundifolia sensu Wagner et al. (1990), non
(L.) Huds.
Dicot
Lamiaceae
Mentha suaveolens Ehrh.
Mentha xvillosa sensu Wagner et al. (1990), non Huds.
Staples et al. 2003:14
Dicot
Lamiaceae
Phyllostegia haliakalae Wawra
Phyllostegia imminuta (Sherff) H.St.John
Wagner 1999b:269
Dicot
Lamiaceae
Phyllostegia parviflora (Gaudich.) Benth. var. lydgatei
(Sherff) W.L.Wagner
Phyllostegia mollis Benth. var. lydgatei Sherff
Wagner 1999a:282
Dicot
Lamiaceae
Plectranthus scutellarioides (L.) R. Br.
Solenostemon scutellarioides (L.) Codd
Grayer et al. 2010/Wagner et al.
2012:48
Dicot
Lamiaceae
Stenogyne kaalae Wawra subsp. kaalae
Stenogyne kaalae Wawra
Wagner & Weller 1999:449
Dicot
Lamiaceae
Stenogyne kaalae Wawra subsp. sherffii (O.Deg.)
W.L.Wagner & Weller
Stenogyne sherffii O.Deg.
Wagner & Weller 1999:449
Dicot
Lauraceae
Cinnamomum burmanni (Nees) Blume
Cinnamomum burmanii (Nees) Blume (spelling
change)
Wagner & Herbst 2003:32
Herbst & Wagner 1999:23
Dicot
Lythraceae
Ammannia coccinea Rottb.
Ammannia auriculata sensu Wagner et al. (1990), non
Willd.
Dicot
Malvaceae
Hibiscus brackenridgei A.Gray subsp. molokaiana
(Rock ex Caum) F.D.Wilson
Hibiscus brackenridgei A.Gray var. molokaiana Rock
ex Caum
Wilson 1993:278/Herbst &
Wagner 1999:23
Dicot
Malvaceae
Hibiscus ovalifolius (Forssk.) Vahl
Hibiscus calyphyllus Cav.
Staples et al. 2003:14
Dicot
Malvaceae
Hibiscus tiliaceus L.
Talipariti tiliaceum (L.) Fryxell
Pfeil et al. 2002/Wagner & Herbst
2003:33
Dicot
Melastomataceae
Arthrostemma ciliatum Pav. ex D.Don
Arthrostema ciliatum Pav. ex D.Don (spelling change)
Imada et al. 2000:13
Dicot
Melastomataceae
Melastoma septemnervium Lour.
Melastoma candidum D.Don
Staples et al. 2003:15
Herbst & Wagner 1999:24
Dicot
Meliaceae
Toona ciliata M.Roem.
Toona ciliata M.Roem. var. australis (F.Muell.)
Bahadur
Dicot
Menispermaceae
Cocculus orbiculatus (L.) DC.
Cocculus trilobus (Thunb.) DC.
Wagner & Herbst 1995:22
Dicot
Moraceae
Ficus cf. platypoda (A.Cunn. ex Miq.) A.Cunn. ex
Miq.
Ficus rubiginosa sensu Nagata (1995), non Desf. ex
Vent.
Oppenheimer & Bartlett 2000:7
C-12
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Myricaceae
Morella cerifera (L.) Small
Myrica cerifera L.
Wilbur 1994:100/ Oppenheimer
2003:15
Dicot
Myricaceae
Morella faya (Aiton) Wilbur
Myrica faya Aiton
Wilbur 1994:103/Staples et al.
2002:13
Dicot
Myrsinaceae
Myrsine lessertiana A.DC.
Myrsine emarginata (Rock) Hosaka
Wagner & Herbst 2003:35
Dicot
Myrtaceae
Corymbia calophylla (R.Br. ex Lindl.) K.D.Hill &
L.A.S.Johnson
Eucalyptus calophylla R.Br. ex Lindl.
Hill & Johnson 1995:240/ Herbst
& Wagner 1999:24
Dicot
Myrtaceae
Corymbia citriodora (Hook.) K.D.Hill &
L.A.S.Johnson
Eucalyptus citriodora Hook.
Hill & Johnson 1995:388/ Herbst
& Wagner 1999:24
Dicot
Myrtaceae
Corymbia ficifolia (F.Muell.) K.D.Hill &
L.A.S.Johnson
Eucalyptus ficifolia F.Muell.
Hill & Johnson 1995:245/ Herbst
& Wagner 1999:24
Dicot
Myrtaceae
Corymbia gummifera (Sol. ex Gaertn.) K.D.Hill &
L.A.S.Johnson
Eucalyptus gummifera (Sol. ex Gaertn.) Hochr.
Hill & Johnson 1995:233/ Herbst
& Wagner 1999:24
Dicot
Myrtaceae
Leptospermum morrisonii Joy Thomps.
Leptospermum laevigatum sensu Wagner et al. (1990),
in part, non (Gaertn.) F.Muell.
Herbarium Pacificum Staff
1999:5
Dicot
Myrtaceae
Leptospermum petersonii F.M.Bailey
Leptospermum flavescens sensu Wagner et al. (1990),
in part, non Sm.
Herbarium Pacificum Staff
1999:5
Dicot
Myrtaceae
Leptospermum polygalifolium Salisb. subsp.
polygalifolium
Leptospermum flavescens Sm.
Herbarium Pacificum Staff
1999:6
Dicot
Myrtaceae
Syzygium sandwicense (A.Gray) Müll.Berol.
Syzygium sandwicensis (A.Gray) Nied.
Wagner et al. 2012:55
Herbst & Wagner 1999:24
Dicot
Nyctaginaceae
Boerhavia acutifolia (Choisy) J.W.Moore
Boerhavia glabrata sensu Wagner et al. (1990), non
Blume
Dicot
Ochnaceae
Ochna thomasiana Engl. & Gilg
Ochna kirkii sensu Wagner et al. (1990), non Oliv.
Herbarium Pacificum Staff
1998:12
Dicot
Oleaceae
Olea europaea L. subsp. cuspidata (Wall. ex G.Don)
Cif.
Olea europaea L. subsp. africana (P.Mill.) P.Green
Green & Wickens 1989:295/
Herbst & Wagner 1999:24
Dicot
Onagraceae
Fuchsia xhybrida Vilm.
Fuchsia hybrida Hort. ex Siebold & Voss
Staples & Herbst 2005:445
Oxalis debilis Kunth var. corymbosa (DC.) Lourteig
Oxalis corymbosa DC.
Herbst & Wagner 1999:25;
Staples & Herbst 2005:447
Coppens d'Eeckenbrugge et. al.
2001/Staples et al. 2003:17
Dicot
Oxalidaceae
Dicot
Passifloraceae
Passiflora tarminiana Coppens & Barney
Passiflora mollissima sensu Wagner et al. (1990), non
(Kunth) Bailey
Dicot
Passifloraceae
Passiflora xviolacea Loisel.
Passiflora xcaerulea sensu Wagner et al. (1990), non L.
Starr et al. 1999:13
Dicot
Piperaceae
Peperomia blanda (Jacq.) Kunth var. floribunda (Miq.)
H.Huber
Peperomia leptostachya Hook. & Arn., non (Nutt.)
Chapman
Huber 1987:293/Florence
1997:173/Herbst & Wagner
1999:25
Dicot
Piperaceae
Peperomia eekana C.DC.
Peperomia nahikuensis H.St.John
Wagner et al. 2012:58
C-13
Synonymy List Arranged By Family
Group
Dicot
Family
Polygonaceae
Current disposition
Emex spinosa (L.) Campd.
Old name (synonym, misapplied name)
Literature source
Emex australis sensu Wagner et al. (1990), in part, non
Steinh.
Parker & Parsons 2012b:71
Dicot
Polygonaceae
Fallopia convolvulus (L.) Á.Löve
Polygonum convolvulus L.
Ronse Decraene & Akeroyd
1988:369/Herbst & Wagner
1999:29
Dicot
Polygonaceae
Muehlenbeckia complexa (A.Cunn.) Meisn.
Muehlenbeckia axillaris sensu Wagner et al. (1990),
non (Hook.f.) Walp.
Staples & Herbst 2005:464
Polygonum capitatum Buch.-Ham. ex D. Don
Ronse Decraene & Akeroyd
1988:367/Herbst & Wagner
1999:29
Dicot
Polygonaceae
Persicaria capitata (Buch.-Ham. ex D.Don) Masam.
Dicot
Polygonaceae
Persicaria chinense (L.) Nakai
Polygonum chinense L.
Ronse Decraene & Akeroyd
1988:367/Herbst & Wagner
1999:29
Dicot
Polygonaceae
Persicaria glabra (Willd.) M.Gómez
Polygonum glabrum Willd.
Wilson 1990:632/Herbst &
Wagner 1999:30
Dicot
Polygonaceae
Persicaria punctata (Elliott) Small
Polygonum punctatum Elliott
Herbst & Wagner 1999:30/ Hinds
& Freeman 2005:586
Dicot
Polygonaceae
Rumex crispus L. subsp. crispus
Rumex crispus L.
Mosyakin & Wagner 1998:42
Dicot
Portulacaceae
Talinum fruticosum (L.) Juss.
Talinum triangulare (Jacq.) Willd.
Wagner & Herbst 1995:25
Dicot
Primulaceae
Lysimachia daphnoides (A.Gray) Hillebr.
Lysimachia kahiliensis H.St.John
Herbst & Wagner 1999:30
Dicot
Primulaceae
Lysimachia hillebrandii Hook.f. ex A.Gray
Lysimachia haupuensis H.St.John
Herbst & Wagner 1999:30
Dicot
Primulaceae
Lysimachia hillebrandii Hook.f. ex A.Gray
Lysimachia ovoidea H.St.John
Herbst & Wagner 1999:30
Dicot
Primulaceae
Lysimachia hillebrandii Hook.f. ex A.Gray
Lysimachia waianaeensis H.St.John
Herbst & Wagner 1999:30
Dicot
Primulaceae
Lysimachia remyi Hillebr.
Lysimachia caliginis H.St.John
Herbst & Wagner 1999:31
Dicot
Primulaceae
Lysimachia remyi Hillebr.
Lysimachia remyi Hillebr. subsp. caliginis (H.St.John)
K.L.Marr
Herbst & Wagner 1999:31
Herbst & Wagner 1999:31
Dicot
Primulaceae
Lysimachia remyi Hillebr.
Lysimachia remyi Hillebr. subsp. subherbacea
(H.St.John) K.L.Marr
Dicot
Primulaceae
Lysimachia remyi Hillebr.
Lysimachia subherbacea (Hillebr.) H.St.John
Herbst & Wagner 1999:31
Dicot
Rhizophoraceae
Bruguiera sexangula (Lour.) Poir.
Bruguiera gymnorhiza sensu Wagner et al. (1990), non
(L.) Savigny
Herbst & Wagner 1999:31
Dicot
Rosaceae
Pyracantha crenatoserrata (Hance) Rehder
Pyracantha angustifolia sensu Wagner et al. (1990), in
part, non (Franch.) C.K.Schneid.
Herbarium Pacificum Staff
1999:9
Dicot
Rosaceae
Pyracantha koidzumii Rehder
Pyracantha angustifolia sensu Wagner et al. (1990), in
part, non (Franch.) C.K.Schneid.
Herbarium Pacificum Staff
1999:9
C-14
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Rubiaceae
Atractocarpus fitzalanii (F.Muell.) Puttock
Randia fitzalanii (F.Muell.) F.Muell. ex Benth.
Puttock 1999:285
Dicot
Rubiaceae
Galium aparine L.
Sherardia arvensis sensu Oppenheimer (2008), non L.
Frohlich & Lau 2012:45
Dicot
Rubiaceae
Gynochthodes trimera (Hillebr.) Razafim. & B.Bremer
Morinda trimera Hillebr.
Razafimandimbison & Bremer
2011:295/Wagner et al. 2012:65
Dicot
Rubiaceae
Kadua acuminata Cham. & Schltdl.
Hedyotis acuminata (Cham. & Schltdl.) Steud.
Terrell et al. 2005:832/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua affinis DC.
Hedyotis terminalis (Hook. & Arn.) W.L.Wagner &
D.R.Herbst
Terrell et al. 2005:832/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua axillaris (Wawra) W.L.Wagner & Lorence
Hedyotis hillebrandii (Fosberg) W.L.Wagner &
D.R.Herbst
Terrell et al. 2005:832/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua centranthoides Hook. & Arn.
Hedyotis centranthoides (Hook. & Arn.) Steud.
Terrell et al. 2005:832/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua cookiana Cham. & Schltdl.
Hedyotis cookiana (Cham. & Schltdl.) Steud.
Terrell et al. 2005:833/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua cordata Cham. & Schltdl. subsp. cordata
Hedyotis schlechtendahliana Steud. subsp.
schlechtendahliana
Terrell et al. 2005:833/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua cordata Cham. & Schltdl. subsp. remyi
(Hillebr.) W.L.Wagner & Lorence
Hedyotis schlechtendahliana Steud. subsp. remyi
(Hillebr.) Fosberg
Terrell et al. 2005:833/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua cordata Cham. & Schltdl. subsp. remyi
(Hillebr.) W.L.Wagner & Lorence
Hedyotis schlechtendahliana Steud. var. remyi
(Hillebr.) Fosberg
Terrell et al. 2005:833/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua cordata Cham. & Schltdl. subsp. waimeae
(Wawra) W.L.Wagner & Lorence
Hedyotis glaucifolia (A.Gray) Fosberg var. waimeae
(Wawra) Fosberg
Terrell et al. 2005:833/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua cordata Cham. & Schltdl. subsp. waimeae
(Wawra) W.L.Wagner & Lorence
Hedyotis schlechtendahliana Steud. subsp. waimeae
(Wawra) W.L.Wagner & D.H.Lorence
Terrell et al. 2005:833/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua coriacea (Sm.) W.L.Wagner & Lorence
Hedyotis coriacea Sm.
Terrell et al. 2005:832/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua degeneri (Fosberg) W.L.Wagner & Lorence
subsp. coprosmifolia (Fosberg) W.L.Wagner &
Lorence
Hedyotis degeneri Fosberg var. coprosmifolia Fosberg
Terrell et al. 2005:833/ Kennedy
et al. 2010:23
Dicot
Rubiaceae
Kadua degeneri (Fosberg) W.L.Wagner & Lorence
subsp. degeneri
Hedyotis degeneri Fosberg var. degeneri
Terrell et al. 2005:833/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua elatior (H.Mann) W.L.Wagner & Lorence
Hedyotis elatior (H.Mann) Fosberg
Terrell et al. 2005:833/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua fluviatilis C.N.Forbes
Hedyotis fluviatilis (C.N.Forbes) Fosberg
Terrell et al. 2005:832/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua flynnii (W.L.Wagner & Lorence) W.L.Wagner
& Lorence
Hedyotis flynnii W.L.Wagner & Lorence
Terrell et al. 2005:833/ Kennedy
et al. 2010:24
C-15
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Rubiaceae
Kadua foggiana (Fosberg) W.L.Wagner & Lorence
Hedyotis foggiana Fosberg
Terrell et al. 2005:832/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua foliosa Hillebr.
Hedyotis foliosa (Hillebr.) Fosberg
Terrell et al. 2005:833/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua formosa Hillebr.
Hedyotis formosa (Hillebr.) Fosberg
Terrell et al. 2005:832/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua fosbergii (W.L.Wagner & D.R.Herbst)
W.L.Wagner & Lorence
Hedyotis fosbergii W.L.Wagner & Herbst
Terrell et al. 2005:832/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua knudsenii Hillebr.
Hedyotis knudsenii (Hillebr.) Fosberg
Terrell et al. 2005:832/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua laxiflora H.Mann
Hedyotis mannii Fosberg
Terrell et al. 2005:833/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua littoralis Hillebr.
Hedyotis littoralis (Hillebr.) Fosberg
Terrell et al. 2005:833/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua parvula A.Gray
Hedyotis parvula (A.Gray) Fosberg
Terrell et al. 2005:833/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua st.-johnii (B.C.Stone & Lane) W.L.Wagner &
Lorence
Hedyotis st.-johnii B.C.Stone & Lane
Terrell et al. 2005:833/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Kadua tryblium (D.R.Herbst & W.L.Wagner)
W.L.Wagner & Lorence
Hedyotis tryblium D.R.Herbst & W.L.Wagner
Terrell et al. 2005:832/ Kennedy
et al. 2010:24
Dicot
Rubiaceae
Nertera granadensis (L.f.) Druce
Coprosma granadensis (L.f.) Heads
Wagner et al. 2012:65
Dicot
Rubiaceae
Oldenlandia corymbosa L.
Hedyotis corymbosa (L.) Lam.
Groeninchkx et al. 2009:125/
Wagner et al. 2012:64
Dicot
Rubiaceae
Oldenlandiopsis callitrichoides (Griseb.) Terrell &
W.H.Lewis
Hedyotis callitrichoides (Griseb.) W.H.Lewis
Terrell & Lewis 1990:185/
Wagner et al. 2012:63
Dicot
Rubiaceae
Paederia foetida L.
Paederia scandens (Lour.) Merr.
Wagner & Herbst 1995:25
Dicot
Rubiaceae
Psydrax odorata (G.Forst.) A.C.Sm. & S.P.Darwin
Canthium odoratum (G. Forst.) Seem.
Smith 1988:230
Richardia scabra L.
Richardia brasiliensis sensu Wagner et al. (1990), in
part, non Gomes
Imada et al. 2008:14
Lorence & Flynn 2006:4
Dicot
Rubiaceae
Dicot
Rubiaceae
Spermacoce exilis (L.O.Williams) C.D.Adams
Spermacoce capitata sensu Wagner et al. (1997), non
Ruiz & Pav.
Dicot
Rubiaceae
Spermacoce exilis (L.O.Williams) C.D.Adams
Spermacoce mauritiana Gideon
Lorence et al. 1995:51
Dicot
Rubiaceae
Spermacoce prostrata Aubl.
Spermacoce ovalifolia sensu Lorence et al. (1995), non
(Mart. & Gal.) Hemsl.
Oppenheimer 2003:23
Rutaceae
Melicope adscendens (H.St.John & E.P.Hume)
T.G.Hartley & B.C.Stone
Pelea adscendens H.St.John & E.P.Hume
Hartley & Stone 1989:120
Dicot
C-16
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Rutaceae
Melicope anisata (H.Mann) T.G.Hartley & B.C.Stone
Pelea anisata H.Mann
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope balloui (Rock) T.G.Hartley & B.C.Stone
Pelea balloui Rock
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope barbigera A.Gray
Pelea barbigera (A.Gray) Hillebr.
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope christophersenii (H.St.John) T.G.Hartley &
B.C.Stone
Pelea christophersenii H.St.John
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope cinerea A.Gray
Pelea cinerea (A.Gray) Hillebr.
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope clusiifolia (A.Gray) T.G.Hartley &
B.C.Stone
Pelea clusiifolia A.Gray
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope cruciata (A.Heller) T.G.Hartley & B.C.Stone
Pelea cruciata A.Heller
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope degeneri (B.C.Stone) T.G.Hartley &
B.C.Stone
Pelea degeneri B.C.Stone
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope elliptica A.Gray
Pelea elliptica (A.Gray) Hillebr.
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope feddei (H.Lév.) T.G.Hartley & B.C.Stone
Pelea feddei H.Lév.
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope haleakalae (B.C.Stone) T.G.Hartley &
B.C.Stone
Pelea haleakalae B.C.Stone
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope haupuensis (H.St.John) T.G.Hartley &
B.C.Stone
Pelea haupuensis H.St.John
Hartley & Stone 1989:120
Dicot
Rutaceae
Melicope hawaiensis (Wawra) T.G.Hartley &
B.C.Stone
Pelea hawaiensis Wawra
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope hiiakae (B.C.Stone) T.G.Hartley &
B.C.Stone
Pelea hiiakae B.C.Stone
Wagner & Herbst 1999:1894
Dicot
Rutaceae
Melicope hosakae (H.St.John) W.L.Wagner &
R.K.Shannon
Melicope honoluluensis (H.St.John) T.G.Hartley &
B.C.Stone
Wagner & Shannon 1998:16
Dicot
Rutaceae
Melicope hosakae (H.St.John) W.L.Wagner &
R.K.Shannon
Pelea honoluluensis H.St.John
Wagner & Shannon 1998:16
Dicot
Rutaceae
Melicope hosakae (H.St.John) W.L.Wagner &
R.K.Shannon
Pelea hosakae H.St.John
Wagner & Shannon 1998:16
Dicot
Rutaceae
Melicope kaalaensis (H.St.John) T.G.Hartley &
B.C.Stone
Pelea kaalaensis H.St.John
Hartley & Stone 1989:121
Rutaceae
Melicope kavaiensis (H.Mann) T.G.Hartley &
B.C.Stone
Pelea kavaiensis H.Mann
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope knudsenii (Hillebr.) T.G.Hartley &
B.C.Stone
Pelea knudsenii Hillebr.
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope lydgatei (Hillebr.) T.G.Hartley & B.C.Stone
Pelea lydgatei Hillebr.
Hartley & Stone 1989:121
Dicot
C-17
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Rutaceae
Melicope macropus (Hillebr.) T.G.Hartley &
B.C.Stone
Dicot
Rutaceae
Melicope makahae (B.C.Stone) T.G.Hartley &
B.C.Stone
Pelea makahae B.C.Stone
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope molokaiensis (Hillebr.) T.G.Hartley &
B.C.Stone
Pelea molokaiensis Hillebr.
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope mucronulata (H.St.John) T.G.Hartley &
B.C.Stone
Pelea mucronulata H.St.John
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope munroi (H.St.John) T.G.Hartley &
B.C.Stone
Pelea munroi H.St.John
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope nealae (B.C.Stone) T.G.Hartley & B.C.Stone
Pelea nealae B.C.Stone
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope oahuensis (H.Lév.) T.G.Hartley & B.C.Stone
Pelea oahuensis H.Lév.
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope obovata (H.St.John) T.G.Hartley &
B.C.Stone
Pelea obovata H.St.John
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope orbicularis (Hillebr.) T.G.Hartley &
B.C.Stone
Pelea orbicularis Hillebr.
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope ovalis (H.St.John) T.G.Hartley & B.C.Stone
Pelea ovalis H.St.John
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope ovata (H.St.John & E.P.Hume) T.G.Hartley
& B.C.Stone
Pelea ovata H.St.John & E.P.Hume
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope pallida (Hillebr.) T.G.Hartley & B.C.Stone
Pelea pallida Hillebr.
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope paniculata (H.St.John) T.G.Hartley &
B.C.Stone
Pelea paniculata H.St.John
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope peduncularis (H.Lév.) T.G.Hartley &
B.C.Stone
Pelea peduncularis H.Lév.
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope pseudoanisata (Rock) T.G.Hartley &
B.C.Stone
Pelea pseudoanisata Rock
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope puberula (H.St.John) T.G.Hartley &
B.C.Stone
Pelea puberula H.St.John
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope quadrangularis (H.St.John & E.P.Hume)
T.G.Hartley & B.C.Stone
Pelea quadrangularis H.St.John & E.P.Hume
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone
Pelea radiata H.St.John
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope reflexa (H.St.John) T.G.Hartley & B.C.Stone
Pelea reflexa H.St.John
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope rotundifolia (A.Gray) T.G.Hartley &
B.C.Stone
Pelea rotundifolia A.Gray
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope saint-johnii (E.P.Hume) T.G.Hartley &
B.C.Stone
Pelea saint-johnii E.P.Hume
Hartley & Stone 1989:121
Pelea macropus Hillebr.
Hartley & Stone 1989:121
C-18
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Rutaceae
Melicope sandwicensis (Hook. & Arn.) T.G.Hartley &
B.C.Stone
Dicot
Rutaceae
Melicope sessilis (H.Lév.) T.G.Hartley & B.C.Stone
Melicope mauii T.G.Hartley & B.C.Stone
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope sessilis (H.Lév.) T.G.Hartley & B.C.Stone
Pelea parvifolia Hillebr.
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope sessilis (H.Lév.) T.G.Hartley & B.C.Stone
Pelea sessilis H.Lév.
Hartley & Stone 1989:121
Dicot
Rutaceae
Melicope volcanica (A.Gray) T.G.Hartley &
B.C.Stone
Pelea volcanica A.Gray
Hartley & Stone 1989:122
Rutaceae
Melicope waialealae (Wawra) T.G.Hartley &
B.C.Stone
Pelea waialealae Wawra
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope wailauensis (H.St.John) T.G.Hartley &
B.C.Stone
Pelea wailauensis H.St.John
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope wawraeana (Rock) T.G.Hartley & B.C.Stone
Pelea wawraeana Rock
Hartley & Stone 1989:122
Dicot
Rutaceae
Melicope zahlbruckneri (Rock) T.G.Hartley &
B.C.Stone
Pelea zahlbruckneri Rock
Hartley & Stone 1989:122
Dicot
Santalaceae
Santalum freycinetianum Gaudich.
Santalum freycinetianum Gaudich. var. freycinetianum
Harbaugh et al. 2010:833
Dicot
Santalaceae
Santalum haleakalae Hillebr. var. haleakalae
Santalum haleakalae Hillebr.
Harbaugh et al. 2010:833
Dicot
Santalaceae
Santalum haleakalae Hillebr. var. lanaiense (Rock)
Harbaugh
Santalum freycinetianum Gaudich. var. lanaiense Rock
Harbaugh et al. 2010:834
Dicot
Santalaceae
Santalum pyrularium A.Gray
Santalum freycinetianum Gaudich. var. pyrularium
(A.Gray) Stemmerm.
Harbaugh et al. 2010:836
Dicot
Sapotaceae
Planchonella sandwicensis (A.Gray) Pierre
Pouteria sandwicensis (A.Gray) Baehni & O.Deg.
Swenson et al. 2007/Wagner et al.
2012:69
Dicot
Sapotaceae
Sideroxylon polynesicum (Hillebr.) Smedmark &
Anderb.
Nesoluma polynesicum (Hillebr.) Baill.
Smedmark & Anderberg
2007:1502/Wagner et al. 2012:69
Dicot
Scrophulariaceae
Mazus pumilus (Burm.f.) Steenis
Mazus rugosus Lour.
Oppenheimer 2004:17
Dicot
Scrophulariaceae
Torenia glabra Osbeck
Torenia asiatica sensu Wagner et al. (1990), in part,
non L.
Staples et al. 2002:15,16
Dicot
Solanaceae
Brugmansia xcandida Pers.
Brugmansia candida Pers.
Staples & Herbst 2005:530
Dicot
Solanaceae
Capsicum annuum L.
Capsicum frutescens L.
Staples & Herbst 2005:532
Dicot
Solanaceae
Solanum lycopersicum L. var. cerasiforme (Dunal)
D.M.Spooner, G.J.Anderson & R.K.Jansen
Lycopersicon esculentum sensu Wagner et al. (1990),
non Mill.
Wagner et al. 1997:62
Dicot
Solanaceae
Solanum lycopersicum L. var. cerasiforme (Dunal)
D.M.Spooner, G.J.Anderson & R.K.Jansen
Lycopersicon pimpinellifolium sensu Wagner et al.
(1990), non (Jusl.) Mill.
Wagner et al. 1997:62
Dicot
Sterculiaceae
Waltheria indica L.
Waltheria pyrolifolia A.Gray
Wagner et al. 2012:71
Dicot
Pelea sandwicensis (Hook. & Arn.) A.Gray
Hartley & Stone 1989:121
C-19
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Dicot
Verbenaceae
Clerodendrum buchananii (Roxb.) Walp. var. fallax
(Lindl.) Bakh.
Dicot
Verbenaceae
Clerodendrum chinense (Osbeck) Mabb.
Clerodendrum philippinum Schauer
Wagner & Herbst 1995:26
Dicot
Verbenaceae
Stachytarpheta australis Moldenke
Stachytarpheta dichotoma sensu Wagner et al. (1990),
non (Ruiz & Pav.) Vahl
Munir 1992:156/Herbst &
Wagner 1999:32
Dicot
Verbenaceae
Stachytarpheta cayennensis (Rich.) Vahl
Stachytarpheta dichotoma (Ruiz & Pav.) Vahl
Munir 1992:145/Herbst &
Wagner 1999:32
Dicot
Verbenaceae
Stachytarpheta cayennensis (Rich.) Vahl
Stachytarpheta urticifolia (Salisb.) Sims
Munir 1992:145/Herbst &
Wagner 1999:32
Dicot
Viscaceae
Korthalsella complanata (Tiegh.) Engl.
Korthalsella taenioides (Comm. ex DC.) Engl. fma.
taenioides
Herbst & Wagner 1999:32
Dicot
Viscaceae
Korthalsella degeneri Danser
Korthalsella taenioides (Comm. ex DC.) Engl. fma.
horneana (Tiegh.) Molvray
Herbst & Wagner 1999:32
Dicot
Viscaceae
Korthalsella latissima (Tiegh.) Danser
Korthalsella taenioides (Comm. ex DC.) Engl. fma.
pendula (Wawra) Molvray
Herbst & Wagner 1999:32
Herbst & Wagner 1999:32
Herbst & Wagner 1999:32
Clerodendrum speciosissimum van Geert
Staples & Herbst 2005:559
Dicot
Viscaceae
Korthalsella platycaula (Tiegh.) Engl.
Korthalsella taenioides (Comm. ex DC.) Engl. fma.
taenioides
Dicot
Viscaceae
Korthalsella remyana Tiegh.
Korthalsella taenioides (Comm. ex DC.) Engl. fma.
remyana (Tiegh.) Molvray
Dicot
Vitaceae
Cissus quadrangularis L.
Cissus quadrangula L.
Frohlich & Lau 2008:10
Dracaena aurea H.Mann
proposed transfer to Dracaena
aurea H.Mann (Jankalski
2008:20)
Dracaena halemanuensis Jankal.
proposed resurrection of
Pleomele stenophylla H.St.John
(as Dracaena halemanuensis
Jankal.) (Jankalski 2008:21)
Pleomele stenophylla H.St.John
proposed resurrection of
Pleomele stenophylla H.St.John
(as Dracaena halemanuensis
Jankal.) (Jankalski 2008:21)
Dracaena rockii (H.St.John) Jankal.
proposed syn. under Dracaena
rockii (H.St.John) Jankal., along
with Pleomele rockii H.St.John
(Jankalski 2008:21)
Monocot
Monocot
Monocot
Monocot
Agavaceae
Agavaceae
Agavaceae
Agavaceae
Pleomele aurea (H.Mann) N.E.Br.
Pleomele aurea (H.Mann) N.E.Br.
Pleomele aurea (H.Mann) N.E.Br.
Pleomele auwahiensis H.St.John
C-20
Synonymy List Arranged By Family
Group
Monocot
Monocot
Monocot
Monocot
Monocot
Family
Agavaceae
Agavaceae
Agavaceae
Agavaceae
Agavaceae
Current disposition
Pleomele auwahiensis H.St.John
Pleomele fernaldii H.St.John
Pleomele forbesii O.Deg.
Pleomele halapepe H.St.John
Pleomele hawaiiensis O.Deg. & I.Deg.
Old name (synonym, misapplied name)
Literature source
Pleomele rockii H.St.John
proposed syn. under Dracaena
rockii (H.St.John) Jankal., along
with Pleomele rockii H.St.John
(Jankalski 2008:21)
Dracaena fernaldii (H.St.John) Jankal.
proposed transfer to Dracaena
fernaldii (H.St.John) Jankal.
(Jankalski 2008:20)
Dracaena forbesii (O.Deg.) Jankal.
proposed transfer to Dracaena
forbesii (O.Deg.) Jankal.
(Jankalski 2008:21)
Dracaena halaapepe (H.St.John) Jankal.
proposed transfer to Dracaena
halaapepe (H.St.John) Jankal.
(Jankalski 2008:21; note spelling
change)
Dracaena konaensis (H.St.John) Jankal.
proposed syn. under Dracaena
konaensis (H.St.John) Jankal.,
along with Pleomele konaensis
H.St.John (Jankalski 2008:21)
Monocot
Agavaceae
Pleomele hawaiiensis O.Deg. & I.Deg.
Pleomele konaensis H.St.John
proposed syn. under Dracaena
konaensis (H.St.John) Jankal.,
along with Pleomele konaensis
H.St.John (Jankalski 2008:21)
Monocot
Alismataceae
Echinodorus berteroi (Speng.) Fassett
Echinodorus rostratus (Nutt.) Engelm.
The Plant List website 2012
Monocot
Araceae
Xanthosoma robustum Schott
Xanthosoma roseum Schott
Herbst et al. 2004:3
Monocot
Arecaceae
Pritchardia glabrata Becc. & Rock
Pritchardia elliptica Rock & Caum
Hodel 2007:S-14
Monocot
Arecaceae
Pritchardia glabrata Becc. & Rock
Pritchardia lanaiensis Becc. & Rock
Hodel 2007:S-14
Monocot
Arecaceae
Pritchardia maideniana Becc.
Pritchardia affinis Becc.
Hodel 2007:S-26
Monocot
Arecaceae
Pritchardia martii (Gaudich.) H.Wendl.
Pritchardia kahukuensis Caum
Hodel 2007:S-20; Wagner et al.
2012:77
Monocot
Arecaceae
Pritchardia napaliensis H.St.John
Pritchardia limahuliensis H.St.John
Hodel 2007:S-37
Monocot
Arecaceae
Pritchardia remota Becc.
Pritchardia aylmer-robinsonii H.St.John
Hodel 2007:S-39
Monocot
Commelinaceae
Gibasis geniculata (Jacq.) Rohw.
Tradescantia geniculata Jacq.
Staples & Herbst 2005:648
Monocot
Costaceae
Cheilocostus speciosus (J.König) C.Specht
Costus speciosus (J.König) Sm.
Specht & Stevenson 2006:159/
Wagner et al. 2012:79
Monocot
Cyperaceae
Carex longii Mack.
Carex ovalis sensu Wagner et al. (1990), non Gooden
Wagner & Herbst 2003:51
C-21
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Monocot
Cyperaceae
Carex nealae R.W.Krauss
Carex thunbergii sensu Wagner et al. (1990), non
Steud.
Dragon & Barrington 2009:1898,
1906/ Wagner et al. 2012:79
Monocot
Cyperaceae
Cyperus congestus Vahl
Mariscus congestus (Vahl) C.B.Clarke
Tucker 1994:87/Wagner &
Herbst 1999:1899
Monocot
Cyperaceae
Cyperus cyperinus (Retz.) Suringar
Mariscus cyperinus (Retz.) Vahl
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus fauriei Kük.
Mariscus fauriei (Kük.) T.Koyama
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus haspan L.
Cyperus halpan L. (spelling change)
Tucker 1994:50/Hughes 1995:4
Monocot
Cyperaceae
Cyperus hillebrandii Boeck. var. decipiens (Hillebr.)
Kük.
Mariscus hillebrandii (Boeck.) T.Koyama subsp.
decipiens (Hillebr.) T.Koyama
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus hillebrandii Boeck. var. hillebrandii
Mariscus hillebrandii (Boeck.) T.Koyama subsp.
hillebrandii
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus hypochlorus Hillebr. var. brevior Kük.
Mariscus hypochlorus (Hillebr.) C.B.Clarke subsp.
brevior (Kük.) T.Koyama
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus hypochlorus Hillebr. var. hypochlorus
Mariscus hypochlorus (Hillebr.) C.B.Clarke
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus involucratus Rottb.
Cyperus alternifolius L. subsp. flabelliformis (Rottb.)
Kük.
Tucker 1994:20/Strong &
Wagner 1997:41
Monocot
Cyperaceae
Cyperus javanicus Houtt.
Mariscus javanicus (Houtt.) Merr. & Metcalfe
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus meyenianus Kunth
Mariscus meyenianus (Kunth) Nees
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus neokunthianus Kük.
Mariscus kunthianus Gaudich.
Tucker 1994:87/Wagner &
Herbst 1999:1900
Tucker 1994:180/Herbst &
Wagner 1999:20/Wagner &
Herbst 1999:1901
Monocot
Cyperaceae
Cyperus odoratus L.
Torulinium odoratum (L.) S.Hooper subsp.
auriculatum (Nees & Meyen ex Kunth) T.Koyama
Monocot
Cyperaceae
Cyperus pennatiformis Kük. var. bryanii Kük.
Mariscus pennatiformis (Kük.) T.Koyama subsp.
bryanii (Kük.) T.Koyama
Tucker 1994:87/Wagner, Herbst
et al. 1999:1900
Monocot
Cyperaceae
Cyperus pennatiformis Kük. var. pennatiformis
Mariscus pennatiformis (Kük.) T.Koyama subsp.
pennatiformis
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus phleoides (Nees ex Kunth) Hillebr. var.
hawaiiensis (H.Mann) Kük.
Mariscus phleoides Nees ex Kunth subsp. hawaiensis
(H.Mann) T.Koyama
Tucker 1994:87/Wagner &
Herbst 1999:1900
Monocot
Cyperaceae
Cyperus phleoides (Nees ex Kunth) Hillebr. var.
phleoides
Mariscus phleoides Nees ex Kunth subsp. phleoides
Tucker 1994:87/Wagner &
Herbst 1999:1900
C-22
Synonymy List Arranged By Family
Group
Monocot
Family
Cyperaceae
Current disposition
Old name (synonym, misapplied name)
Literature source
Cyperus polystachyos Rottb.
Pycreus polystachyos (Rottb.) P.Beauv. subsp.
holocericeus (Link) T.Koyama
Tucker 1994:72/Herbst &
Wagner 1999:20/Wagner &
Herbst 1999:1901
Tucker 1994:72/Herbst &
Wagner 1999:20/Wagner &
Herbst 1999:1901
Monocot
Cyperaceae
Cyperus polystachyos Rottb.
Pycreus polystachyos (Rottb.) P.Beauv. subsp.
polystachyos
Monocot
Cyperaceae
Cyperus rockii Kük.
Mariscus rockii (Kük.) T.Koyama
Tucker 1994:87/Wagner &
Herbst 1999:1901
Monocot
Cyperaceae
Cyperus sandwicensis Kük.
Mariscus sandwicensis (Kük.) T.Koyama
Tucker 1994:87/Wagner &
Herbst 1999:1901
Monocot
Cyperaceae
Cyperus sanguinolentus Vahl
Pycreus sanguinolentus (Vahl) Nees
Tucker 1994:66/Herbst &
Wagner 1999:20/Wagner &
Herbst 1999:1901
Monocot
Cyperaceae
Fimbristylis littoralis Gaudich.
Fimbristylis miliacea (L.) Vahl
Strong 2004:1069/Imada 2007:35
Monocot
Cyperaceae
Gahnia lacera (A.Rich.) Steud.
Gahnia lanaiensis O.Deg., I.Deg. & J.Kern
Koyama et al. 2010:30
Monocot
Cyperaceae
Morelotia gahniiformis Gaudich.
Gahnia gahniiformis (Gaudich.) A.Heller
Lorence et al. 1995:34/Wagner &
Herbst 2003:52
Monocot
Cyperaceae
Schoenoplectiella juncoides (Roxb.) Lye
Schoenoplectus juncoides (Roxb.) Palla
Lye 2003:25/Wagner et al.
2012:82
Monocot
Cyperaceae
Schoenoplectus tabernaemontani (C.C.Gmel.) Palla
Schoenoplectus lacustris (L.) Palla subsp. validus
(Vahl) T.Koyama
Smith 1995:98/Kennedy et al.
2010:21
Monocot
Eriocaulaceae
Eriocaulon scariosum Sm.
Eriocaulon sp. A
Herbst & Wagner 1999:20
Monocot
Iridaceae
Aristea gerrardii Weim.
Aristea cf. compressa sensu Herbarium Pacificum Staff
(1999), non Buchinger ex Baker
Staples et al. 2003:13
Monocot
Lemnaceae
Landoltia punctata (G.Mey.) Les & D.J.Crawford
Spirodela punctata (G.Mey.) C.H.Thomps.
Staples et al. 2002:11
Wagner et al. 1997:58
Monocot
Lemnaceae
Lemna aequinoctialis Welw.
Lemna perpusilla sensu Wagner et al. (1990), non
Torr.
Monocot
Liliaceae
Asparagus plumosus Baker
Asparagus setaceus sensu Lorence et al. (1995), non
(Kunth) Jessop
Staples & Herbst 2005:682
Herbst & Wagner
1999:23/Staples & Herbst
2005:689
Monocot
Liliaceae
Hippeastrum striatum (Lam.) H.E.Moore
Hippeastrum puniceum sensu Wagner et al. (1990),
non (Lam.) Voss
Monocot
Orchidaceae
Habenaria rodeiensis Barb.Rodr.
Habenaria sp.
Herbst & Wagner 1999:24
Agrostis exarata Trin.
Agrostis exarata Trin. var. monolepis (Torr.)
A.S.Hitchc.
Snow 2008:38
Monocot
Poaceae
C-23
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Monocot
Poaceae
Andropogon bicornis L.
Andropogon glomeratus sensu Herbst & Clayton
(1998), in part, non (Walter) Britton, Sterns &
Poggenb.
Monocot
Poaceae
Andropogon bicornis L.
Andropogon virginicus sensu Wagner et al. (1990), in
part, non L.
Snow & Lau 2010:46
Monocot
Poaceae
Andropogon bicornis L.
Schizachyrium scoparium sensu Lorence et al. (1995),
in part, non (Michx.) Nash
Snow & Lau 2010:46
Monocot
Poaceae
Andropogon glomeratus (Walter) Britton, Sterns &
Poggenb. var. pumilus (Vasey) L.H.Dewey
Andropogon virginicus sensu Starr et al. (2002), non
L.
Snow & Lau 2010:47
Herbst & Clayton 1998:18/ Snow
& Lau 2010:47
Snow & Lau 2010:46
Monocot
Poaceae
Andropogon glomeratus (Walter) Britton, Sterns &
Poggenb. var. pumilus (Vasey) L.H.Dewey
Schizachyrium condensatum sensu Lorence et al.
(1995) et Herbst et al. (2004), in part, non (Kunth)
Nees
Monocot
Poaceae
Andropogon virginicus L. var. virginicus
Andropogon virginicus L.
Snow & Lau 2010:48
Monocot
Poaceae
Bromus catharticus Vahl
Bromus willdenowii Kunth
Herbst & Clayton 1998:20
Monocot
Poaceae
Bromus diandrus Roth
Bromus rigidus Roth
Snow 2008:38
Monocot
Poaceae
Bromus hordeaceus L.
Bromus hordeaceus L. subsp. hordeaceus
Snow 2008:39
Monocot
Poaceae
Bromus hordeaceus L.
Bromus hordeaceus L. subsp. molliformis (J.Lloyd)
Maire & Weiller
Snow 2008:39
Monocot
Poaceae
Bromus hordeaceus L.
Bromus molliformis J.Lloyd
Snow 2008:39
Monocot
Poaceae
Bromus hordeaceus L.
Bromus mollis L.
Snow 2008:39
Monocot
Poaceae
Cenchrus americanus (L.) Morrone
Pennisetum glaucum (L.) R.Br.
Chemisquy et al. 2010:127
Monocot
Poaceae
Cenchrus caninus (Reinw. ex Blume) Morrone
Pennisetum macrostachyum (Brongn.) Trin.
Chemisquy et al. 2010:127
Monocot
Poaceae
Cenchrus clandestinus (Hochst. ex Chiov.) Morrone
Pennisetum clandestinum Hochst. ex Chiov.
Chemisquy et al. 2010:127/
Wagner et al. 2012:98
Monocot
Poaceae
Cenchrus complanatus (Nees) Morrone
Pennisetum complanatum (Nees) Hemsl.
Chemisquy et al. 2010:127/
Wagner et al. 2012:98
Monocot
Poaceae
Cenchrus longisetus M.C.Johnst.
Pennisetum villosum R.Br. ex Fresen.
Wagner et al. 2005–
Monocot
Poaceae
Cenchrus polystachios (L.) Morrone
Pennisetum polystachion (L.) Schult.
Chemisquy et al. 2010:129/
Wagner et al. 2012:99
Monocot
Poaceae
Cenchrus purpureus (Schumach.) Morrone
Pennisetum purpureum Schumach.
Chemisquy et al. 2010:129/
Wagner et al. 2012:99
Monocot
Poaceae
Cenchrus setaceus (Forssk.) Morrone
Pennisetum setaceum (Forssk.) Chiov.
Chemisquy et al. 2010:129/
Wagner et al. 2012:99
C-24
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Monocot
Poaceae
Dichanthelium cynodon (Reichardt) C.A.Clark &
Gould
Panicum cynodon Reichardt
Clayton & Snow 2010:55;
Wagner et al. 2012:92
Monocot
Poaceae
Dichanthelium hillebrandianum (Hitchc.) C.A.Clark &
Gould
Panicum hillebrandianum A.S. Hitchc.
Clayton & Snow 2010:55;
Wagner et al. 2012:92
Monocot
Poaceae
Dichanthelium isachnoides (Munro ex Hillebr.)
C.A.Clark & Gould
Panicum isachnoides Munro ex Hillebr.
Clayton & Snow 2010:55;
Wagner et al. 2012:92
Monocot
Poaceae
Dichanthelium koolauense (H.St.John & Hosaka)
C.A.Clark & Gould
Panicum koolauense H.St.John & Hosaka
Clayton & Snow 2010:55;
Wagner et al. 2012:92
Monocot
Poaceae
Dichelachne micrantha (Cav.) Domin
Dichelachne sciurea (R.Br.) Hook.f.
Herbst & Clayton 1998:22
Monocot
Poaceae
Dichelachne micrantha (Cav.) Domin
Garnotia acutigluma sensu Lorence & Flynn (1999),
non (Steud.) Ohwi
Snow & Davidse 2011:18
Monocot
Poaceae
Digitaria eriantha Steud.
Digitaria pentzii Stent
Herbst & Clayton 1998:23
Monocot
Poaceae
Diplachne fusca subsp. uninervia (J.Presl)
P.M.Peterson & N.Snow
Leptochloa fusca (L.) Kunth subsp. uninervia (J.Presl)
N.Snow
Peterson et al. 2012:1327/
Wagner et al. 2012:96
Monocot
Poaceae
Diplachne fusca subsp. uninervia (J.Presl)
P.M.Peterson & N.Snow
Leptochloa uninervia (J.Presl) A.S.Hitchc. & Chase
Imada et al. 2000:14; Peterson et
al. 2012:1327/Wagner et al.
2012:96
Monocot
Poaceae
Disakisperma dubia (Kunth) P.M.Peterson & N.Snow
Leptochloa dubia (Kunth) Nees
Peterson et al. 2012:1327/
Wagner et al. 2012:96
Monocot
Poaceae
Echinochloa crusgalli (L.) P.Beauv.
Echinochloa walteri sensu Wagner et al. (1990), non
(Pursh) A.Heller
Herbst & Clayton 1998:24
Monocot
Poaceae
Echinochloa esculenta (A.Braun) H.Scholz
Echinochloa utilis Ohwi & Yabuno
Snow 2008:39
Monocot
Poaceae
Echinochloa oryzoides (Ard.) Fritsch
Echinochloa stagnina sensu Wagner et al. (1990), non
(Retz.) P.Beauv.
Herbst & Clayton 1998:24
Herbst & Clayton 1998:24
Monocot
Poaceae
Echinochloa picta (J.König) P.W.Michael
Echinochloa stagnina sensu Wagner et al. (1990), non
(Retz.) P.Beauv.
Monocot
Poaceae
Eragrostis amabilis (L.) Wight & Arn. ex Nees
Eragrostis tenella (L.) P.Beauv. ex Roem. &
J.A.Schultes
Lorence et al. 1995:45
Monocot
Poaceae
Eragrostis brownii (Kunth) Nees ex Steud.
Eragrostis molokaiensis H.St.John
Herbst & Clayton 1998:26,27
Monocot
Poaceae
Eragrostis curvula (Schrad.) Nees
Leptochloa uninervia sensu Warren & Herbst (1994),
in part, non (J.Presl) A.S.Hitchc. & Chase
Herbst & Clayton 1998:29
Monocot
Poaceae
Eragrostis leptostachya (R.Br.) Steud.
Eragrostis hosakai O.Deg.
Herbst & Clayton 1998:26,27
Monocot
Poaceae
Eragrostis pectinacea (Michx.) Nees var. pectinacea
Eragrostis pectinacea (Michx.) Nees
Snow & Lau 2010:51
Eragrostis pilosa (L.) P.Beauv.
Eragrostis pectinacea sensu Wagner et al. (1990), in
part, non (Michx.) Nees
Herbst & Clayton 1998: 27
Monocot
Poaceae
C-25
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Monocot
Poaceae
Eragrostis pilosa (L.) P.Beauv.
Eragrostis monticola sensu Wagner et al. (1990), in
part, non (Gaudich.) Hillebr.
Monocot
Poaceae
Eragrostis pilosa (L.) P.Beauv.
Eragrostis leptophylla sensu Wagner et al. (1990), in
part, non Hitchc.
Herbst & Clayton 1998: 27
Monocot
Poaceae
Eragrostis variabilis (Gaudich.) Steud.
Eragrostis hobdyi H.St.John
Snow & Lau 2010:50
Monocot
Poaceae
Eustachys petraea (Sw.) Desv.
Chloris petraea Sw.
Wagner & Herbst 1995:24
Monocot
Poaceae
Festuca bromoides L.
Vulpia bromoides (L.) Gray
Wagner et al. 2012:101
Monocot
Poaceae
Festuca myuros L.
Vulpia myuros (L.) C.C.Gmel.
Wagner et al. 2012:101
Snow 2008:39
Herbst & Clayton 1998:28
Herbst & Clayton 1998: 27
Monocot
Poaceae
Glyceria notata Chevall.
Glyceria fluitans sensu Wagner et al. (1990), non (L.)
R.Br.
Monocot
Poaceae
Hordeum murinum L. subsp. leporinum (Link)
Arcang.
Hordeum leporinum Link
Monocot
Poaceae
Hyparrhenia hirta (L.) Stapf
Andropogon virginicus sensu Herbarium Pacificum
Staff (1999), non L.
Snow & Lau 2010:48
Sun & Phillips 2008:611/ Snow
& Lau 2010:52
Monocot
Poaceae
Ischaemum ciliare Retz.
Ischaemum indicum sensu Wagner et al. (1990), non
(Houtt.) Merr.
Monocot
Poaceae
Koeleria macrantha (Ledeb.) Schult.
Koeleria nitida Nutt.
Herbst & Clayton 1998:29
Monocot
Poaceae
Lachnagrostis filiformis (G.Forst.) Trin.
Agrostis avenacea J.G.Gmel.
Snow 2008:39
Monocot
Poaceae
Leymus triticoides (Buckley) Pilg.
Elymus triticoides Buckley
Soreng et al. 2003:425
Monocot
Poaceae
Lolium temulentum L.
Lolium tementulum L. (spelling change)
Clayton & Snow 2010:26
Monocot
Poaceae
Melinis repens (Willd.) Zizka
Rhynchelytrum repens (Willd.) C.E.Hubb.
Herbst & Clayton 1998:29
Monocot
Poaceae
Nassella cernua (Stebbins & Love) Barkworth
Stipa cernua Stebbins & Love
Barkworth 1990:609/Snow
2008:39
Monocot
Poaceae
Oplismenus hirtellus (L.) P.Beauv. subsp. hirtellus
Oplismenus hirtellus (L.) P.Beauv.
Snow & Lau 2010:52
Snow & Davidse 2011:18
Monocot
Poaceae
Paspalum arundinaceum Poir.
Paspalum aff. laxum sensu Snow & Lau (2010), non
Lam.
Monocot
Poaceae
Paspalum arundinaceum Poir.
Paspalum paniculatum sensu Oppenheimer (2007),
non L.
Snow & Lau 2010:53/ Snow &
Davidse 2011:18
Monocot
Poaceae
Paspalum mandiocanum Trin. var. mandiocanum
Paspalum aff. thunbergii sensu Snow & Lau (2010),
non Kunth ex Steud.
Snow & Davidse 2011:18
Herbst & Clayton 1998:33
Herbst & Clayton 1998:33
Monocot
Poaceae
Phalaris aquatica L.
Phalaris tuberosa L. var. stenoptera sensu Wagner et
al. (1990), non (Hack.) Hitchc.
Monocot
Poaceae
Phragmites australis (Cav.) Trin. ex Steud.
Phragmites karka sensu Wagner et al. (1990), non
(Retz.) Trin.
C-26
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Monocot
Poaceae
Piptatherum miliaceum (L.) Coss.
Oryzopsis miliacea (L.) Benth. & Hook.f. ex Aschers.
& Schweinf.
Monocot
Poaceae
Polypogon viridis (Gouan) Breistr.
Agrostis semiverticillata (Forssk.) C.Christens.
Herbst & Clayton 1998:18, 33, 34
Monocot
Poaceae
Rytidosperma penicillatum (Labill.) Connor & Edgar
Danthonia pilosa sensu Wagner et al. (1990), non
R.Br.
Herbst & Clayton 1998:34;
Darbyshire et al. 2010:670
Monocot
Poaceae
Rytidosperma penicillatum (Labill.) Connor & Edgar
Rytidosperma pilosum sensu Herbst & Clayton (1998),
non (R.Br.) Connor & Edgar
Darbyshire et al. 2010:670
Monocot
Poaceae
Rytidosperma semiannulare (Labill.) Connor & Edgar
Danthonia semiannularis (Labill.) R.Br.
Herbst & Clayton 1998:34
Monocot
Poaceae
Schedonorus arundinaceus (Schreb.) Dumort.
Festuca arundinacea Schreb.
Snow 2008:40
Monocot
Poaceae
Setaria parviflora (Poir.) Kerguélen
Setaria gracilis Kunth
Herbst & Clayton 1998:35
Monocot
Poaceae
Sorghum halepense (L.) Pers.
Sorghum halpense (L.) Pers. (spelling change)
Clayton & Snow 2010:84
Monocot
Poaceae
Sporobolus africanus (Poir.) Robyns & Tournay
Sporobolus indicus sensu Wagner et al. (1990) et
Warren (1993), in part, non (L.) R.Br.
Herbst & Clayton 1998:36
Monocot
Poaceae
Sporobolus diandrus (Retz.) P.Beauv.
Sporobolus diander (Retz.) P.Beauv. (spelling change)
Clayton & Snow 2010:47
Monocot
Poaceae
Sporobolus elongatus R.Br.
Sporobolus pyramidalis sensu Herbst & Clayton
(1998), non P.Beauv.
Snow 2008:41
Snow 2008:41
Snow 2008:40
Monocot
Poaceae
Sporobolus elongatus R.Br.
Sporobolus pyramidatus sensu Wagner & Herbst
(1995), in part, non (Lam.) Hitchc.
Monocot
Poaceae
Sporobolus pyramidatus (Lam.) Hitchc.
Sporobolus piliferus sensu Snow (2008), non (Trin.)
Kunth
Snow & Lau 2010:56
Monocot
Poaceae
Thysanolaena latifolia (Roxb ex Hornem.) Honda
Thysanolaena maxima (Roxb.) Kuntze
Snow & Lau 2010:56
Monocot
Poaceae
Trisetum inaequale Whitney
Trisetum glomeratum sensu Wagner et al. (1990), in
part, non (Kunth) Trin.
Herbst & Clayton 1998:37
Urochloa brizantha (Hochst. ex A.Rich.) R.D.Webster
Brachiaria brizantha (Hochst. ex A.Rich.) Stapf
Zuloaga et al. 2003:630/ Clayton
& Snow 2010:60/ Wagner et al.
2012:88
Urochloa decumbens (Stapf) R.D.Webster
Brachiaria brizantha sensu Starr et al. (2006), in part,
non (Hochst. ex A.Rich.) Stapf
Snow & Lau 2010:49
Brachiaria decumbens Stapf
Zuloaga et al. 2003:630/ Clayton
& Snow 2010:60/ Wagner et al.
2012:89
Brachiaria distachya (L.) Stapf
Zuloaga et al. 2003:631/ Wagner
& Herbst 2003:58/ Clayton &
Snow 2010:60
Monocot
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Poaceae
Urochloa decumbens (Stapf) R.D.Webster
Urochloa distachya (L.) T.Q.Nguyen
C-27
Synonymy List Arranged By Family
Group
Monocot
Family
Poaceae
Current disposition
Urochloa distachya (L.) T.Q.Nguyen
Old name (synonym, misapplied name)
Literature source
Brachiaria subquadripara (Trin.) Hitchc.
Zuloaga et al. 2003:631/ Wagner
& Herbst 2003:58/ Clayton &
Snow 2010:60
Monocot
Poaceae
Urochloa distachya (L.) T.Q.Nguyen
Urochloa subquadripara (Trin.) R.D.Webster
Zuloaga et al. 2003:631/ Wagner
& Herbst 2003:58/ Clayton &
Snow 2010:60
Monocot
Poaceae
Urochloa maxima (Jacq.) R.D.Webster
Panicum maximum Jacq.
Zuloaga et al. 2003:632/ Wagner
et al. 2012:97
Brachiaria mollis (Sw.) Parodi
Zuloaga et al. 2003:632/ Wagner
& Herbst 2003:58/ Clayton &
Snow 2010:61
Brachiaria mutica (Forssk.) Stapf
Zuloaga et al. 2003:632/ Wagner
& Herbst 2003:58/ Clayton &
Snow 2010:61
Brachiaria plantaginea (Link) Hitchc.
Zuloaga et al. 2003:633/ Wagner
& Herbst 2003:58/ Clayton &
Snow 2010:61
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Urochloa mollis (Sw.) O.Morrone & F.Zuloaga
Urochloa mutica (Forssk.) T.Q.Nguyen
Urochloa plantaginea (Link) R.D.Webster
Monocot
Poaceae
Urochloa reptans (L.) Stapf
Brachiaria reptans (L.) C.A.Gardner & C.E.Hubb.
Zuloaga et al. 2003:633/ Wagner
& Herbst 2003:58/ Clayton &
Snow 2010:61
Monocot
Potamogetonaceae
Stuckenia pectinata (L.) Börner
Coleogeton pectinatus (L.) Les & R.R.Haynes
Haynes et al. 1998:241/Herbst &
Wagner 1999:30
Monocot
Potamogetonaceae
Stuckenia pectinata (L.) Börner
Potamogeton pectinatus L.
Haynes et al. 1998:241/Herbst &
Wagner 1999:30
Pteridophyte
Aspleniaceae
Asplenium dielerectum Viane
Diellia erecta Brack. fma. alexandri (Hillebr.)
W.H.Wagner
Snow et al. 2011:12 (status of f.
alexandri under study)
Pteridophyte
Aspleniaceae
Asplenium dielerectum Viane
Diellia erecta Brack. fma. erecta
Snow et al. 2011:12
Pteridophyte
Aspleniaceae
Asplenium dielerectum Viane
Diellia erecta Brack. fma. pumila (Brack.)
W.H.Wagner
Snow et al. 2011:12 (status of f.
pumila under study)
Pteridophyte
Aspleniaceae
Asplenium dielfalcatum Viane
Diellia falcata Brack.
Snow et al. 2011:12
Pteridophyte
Aspleniaceae
Asplenium diellaciniatum Viane
Diellia laciniata (Hillebr.) Diels
Wagner et al. 2012:103
Pteridophyte
Aspleniaceae
Asplenium dielmannii Viane
Diellia mannii (D.C.Eaton) W.J.Rob.
Snow et al. 2011:12
Pteridophyte
Aspleniaceae
Asplenium dielpallidum N.Snow
Diellia pallida W.H.Wagner
Snow et al. 2011:12
Pteridophyte
Aspleniaceae
Asplenium xflagrum W.H.Wagner & D.D.Palmer
Asplenium hobdyi W.H.Wagner x A. normale D.Don
Palmer 2003:76
Pteridophyte
Aspleniaceae
Asplenium xjoellaui N.Snow
Asplenium xlauii (W.H.Wagner) N.Snow
Snow 2011:161
Pteridophyte
Aspleniaceae
Asplenium xjoellaui N.Snow
Diellia falcata Brack. x D. unisora W.H.Wagner
Snow 2011:161
C-28
Synonymy List Arranged By Family
Group
Pteridophyte
Family
Aspleniaceae
Current disposition
Old name (synonym, misapplied name)
Literature source
Asplenium xjoellaui N.Snow
Diellia xlauii W.H.Wagner
Snow 2011:161
Palmer 2003:76
Pteridophyte
Aspleniaceae
Asplenium xkokeense W.H.Wagner
Asplenium aethiopicum (Burm.f.) Bech. x A.
polyodon G.Forst.
Pteridophyte
Aspleniaceae
Asplenium leucostegioides Baker
Diellia leucostegioides (Baker) W.H.Wagner
Snow et al. 2011:12
Pteridophyte
Aspleniaceae
Asplenium xsphenocookii W.H.Wagner
Asplenium polyodon G.Forst. x A. sphenotomum
Hillebr.
Palmer 2003:76
Pteridophyte
Aspleniaceae
Asplenium unisorum (W.H.Wagner) Viane
Diellia unisora W.H.Wagner
Snow et al. 2011:12
Asplenium xwaikamoi W.H.Wagner
Asplenium acuminatum Hook. & Arn. x A.
aethiopicum (Burm.f.) Bech.
Palmer 2003:76
Blechnum hawaiianum Christenh.
proposal to conserve name
Blechnum hawaiianum Christenh.
over B. norfolkense (Christenhusz
2011:1772)
Blechnum norfolkense Christenh.
proposed transfer to Blechnum
norfolkense Christenh.
(Christenhusz et al. 2011:20)
Doodia lyonii O.Deg.
Blechnum lyonii (O.Deg.) Christenh.
proposed transfer to Blechnum
lyonii (O.Deg.) Christenh.
(Christenhusz et al. 2011:20)
Botrychium subbifoliatum Brack.
Sceptridium subbifoliatum (Brack.) W.H.Wagner &
F.S.Wagner
Snow et al. 2011:14
Palmer 2003:186
Thomson 2004:800
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Aspleniaceae
Blechnaceae
Blechnaceae
Blechnaceae
Botrychiaceae
Doodia kunthiana Gaudich.
Doodia kunthiana Gaudich.
Pteridophyte
Dennstaedtiaceae
Microlepia xadulterina W.H.Wagner
Microlepia speluncae (L.) T.Moore x M. strigosa
(Thunb.) C.Presl
Pteridophyte
Dennstaedtiaceae
Pteridium aquilinum (L.) Kuhn subsp. decompositum
(Gaudich.) Lamoureux ex J.A.Thomson
Pteridium aquilinum (L.) Kuhn var. decompositum
(Gaudich.) R.M.Tryon
Pteridophyte
Dicksoniaceae
Cibotium xheleniae D.D.Palmer
Cibotium chamissoi Kaulf. x C. menziesii Hook.
Palmer 2003:98
Smith et al. 2006:717/Snow et al.
2011:13
Pteridophyte
Dryopteridaceae
Dryopteris rubiginosa (Brack.) Kuntze
Nothoperanema rubiginosa (Brack.) A.R.Sm. &
D.D.Palmer
Pteridophyte
Grammitidaceae
Adenophorus xabbottiae W.H.Wagner
Adenophorus hymenophylloides (Kaulf.) Hook. &
Grev. x A. tamariscinus (Kaulf.) Hook. & Grev.
Palmer 2003:36
Pteridophyte
Grammitidaceae
Adenophorus xbishopii (W.H.Wagner) xxxxx
Adenophorus oahuensis (Copel.) L.E.Bishop x A.
pinnatifidus Gaudich.
Palmer 2003:41
Pteridophyte
Grammitidaceae
Adenophorus xcarsonii Ranker
Adenophorus hymenophylloides (Kaulf.) Hook. &
Grev. x A. tripinnatifidus Gaudich.
Palmer 2003:36
Grammitis tenella Kaulf.
Ranker 2008:171/Snow et al.
2011:14
Pteridophyte
Grammitidaceae
Adenophorus tenellus (Kaulf.) Ranker
C-29
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Pteridophyte
Grammitidaceae
Oreogrammitis baldwinii (Baker) Parris
Grammitis baldwinii (Baker) Copel.
Parris 2007:256
Pteridophyte
Grammitidaceae
Oreogrammitis forbesiana (W.H.Wagner) Parris
Grammitis forbesiana W.H.Wagner
Parris 2007:260
Pteridophyte
Grammitidaceae
Oreogrammitis hookeri (Brack.) Parris
Grammitis hookeri (Brack.) Copel.
Parris 2007:260
Pteridophyte
Hymenophyllaceae
Crepidomanes draytonianum (Brack.) Ebihara & K.
Iwats.
Vandenboschia draytoniana (Brack.) Copel.
Ebihara et al. 2006:238/Snow et
al. 2011:13
Pteridophyte
Hymenophyllaceae
Crepidomanes minutum (Blume) K.Iwats.
Gonocormus minutus (Blume) Bosch
Ebihara et al. 2006:238/Snow et
al. 2011:13
Pteridophyte
Hymenophyllaceae
Crepidomanes proliferum (Blume) Bostock
Gonocormus prolifer (Blume) Prantl
Ebihara et al. 2006:238/Snow et
al. 2011:13
Pteridophyte
Hymenophyllaceae
Hymenophyllum lanceolatum Hook. & Arn.
Sphaerocionium lanceolatum (Hook. & Arn.) Copel.
Ebihara et al. 2006:230/Snow et
al. 2011:13
Pteridophyte
Hymenophyllaceae
Hymenophyllum obtusum Hook. & Arn.
Sphaerocionium obtusum (Hook. & Arn.) Copel.
Ebihara et al. 2006:230/Snow et
al. 2011:13
Pteridophyte
Hymenophyllaceae
Hymenophyllum recurvum Gaudich.
Mecodium recurvum (Gaudich.) Copel.
Ebihara et al. 2006:231/Snow et
al. 2011:13
Pteridophyte
Lindsaeaceae
xLindsaeosoria flynnii W.H.Wagner
Lindsaea ensifolia Sw. x Sphenomeris chinensis (L.)
Maxon
Palmer 2003:174
Pteridophyte
Lycopodiaceae
Huperzia xcarlquistii Beitel & W.H.Wagner
Huperzia erubescens (Brack.) Holub x H. subintegra
(Hillebr.) Beitel & W.H.Wagner
Palmer 2003:265
Pteridophyte
Lycopodiaceae
Huperzia xgillettii Beitel & W.H.Wagner
Huperzia serrata (Thunb. ex Murray) Trevis. x H.
subintegra (Hillebr.) Beitel & W.H.Wagner
Palmer 2003:265
Palmer 2003:259
Palmer 2003:265
Pteridophyte
Lycopodiaceae
Huperzia xkoolauensis (W.H.Wagner) Kartesz
Huperzia nutans (Brack.) Rothm. x H. phyllantha
(Hook. & Arn.) Holub
Pteridophyte
Lycopodiaceae
Huperzia xmedeirosii Beitel & W.H.Wagner
Huperzia haleakalae (Brack.) Holub x H. subintegra
(Hillebr.) Beitel & W.H.Wagner
Pteridophyte
Lycopodiaceae
Huperzia xsulcinervia (Spring) Trevis.
Huperzia erubescens (Brack.) Holub x H. serrata
(Thunb. ex Murray) Trevis.
Palmer 2003:263
Hovenkamp & Miyamoto
2005:293
Pteridophyte
Nephrolepidaceae
Nephrolepis brownii (Desv.) Hovenkamp & Miyam.
Nephrolepis multiflora (Roxb.) F.M.Jarrett ex
C.V.Morton
Pteridophyte
Nephrolepidaceae
Nephrolepis exaltata (L.) Schott
Nephrolepis exaltata (L.) Schott subsp. hawaiiensis
W.H.Wagner
Hovenkamp & Miyamoto
2005:300
Pteridophyte
Ophioglossaceae
Ophioglossum pendulum L.
Ophioderma pendulum (L.) C.Presl subsp. falcatum
(C.Presl) R.T.Clausen
Smith et al. 2006:709/Wagner et
al. 2012:105
Pteridophyte
Ophioglossaceae
Ophioglossum pendulum L.
Ophioderma pendulum (L.) C.Presl subsp. pendulum
Smith et al. 2006:709/Wagner et
al. 2012:105
Pteridophyte
Polypodiaceae
Stenogrammitis saffordii (Maxon) Labiak
Lellingeria saffordii (Maxon) A.R.Sm. & R.C.Moran
Labiak 2011:147
C-30
Synonymy List Arranged By Family
Group
Family
Current disposition
Old name (synonym, misapplied name)
Literature source
Pteridophyte
Pteridaceae
Doryopteris subdecipiens W.H.Wagner
Doryopteris decipiens (Hook.) J.Sm. x D. decora
Brack.
Pteridophyte
Thelypteridaceae
Cyclosorus boydiae (D.C.Eaton) W.H.Wagner
Christella boydiae (D.C.Eaton) Holttum
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Cyclosorus cyatheoides (Kaulf.) Farw.
Christella cyatheoides (Kaulf.) Holttum
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Cyclosorus dentatus (Forssk.) Ching
Christella dentata (Forssk.) Brownsey & Jermy
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Cyclosorus hudsonianus (Brack.) Ching
Pneumatopteris hudsoniana (Brack.) Holttum
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Cyclosorus xintermedia W.H.Shieh & J.L.Tsai
Christella dentata (Forssk.) Brownsey & Jermy x C.
parasitica (L.) H.Lév.
Snow et al. 2011:15
Snow et al. 2011:15
Palmer 2003:133
Pteridophyte
Thelypteridaceae
Cyclosorus xintermedia W.H.Shieh & J.L.Tsai
Christella xintermedia (W.H.Shieh & J.L.Tsai)
D.D.Palmer
Pteridophyte
Thelypteridaceae
Cyclosorus xpalmeri (W.H.Wagner) W.H.Wagner
Christella cyatheoides (Kaulf.) Holttum x C. dentata
(Forssk.) Brownsey & Jermy
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Cyclosorus xpalmeri (W.H.Wagner) W.H.Wagner
Christella xpalmeri W.H.Wagner
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Cyclosorus parasiticus (L.) Farw.
Christella parasitica (L.) H.Lév.
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Cyclosorus pendens (D.D.Palmer) N.Snow
Pneumatopteris pendens D.D.Palmer
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Cyclosorus sandwicensis (Brack.) Copel.
Pneumatopteris sandwicensis (Brack.) Holttum
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Cyclosorus wailele (Flynn) W.H.Wagner
Christella wailele (Flynn) D.D.Palmer
Snow et al. 2011:15
Pteridophyte
Thelypteridaceae
Thelypteris globulifera (Brack.) C.F.Reed
Amauropelta globulifera (Brack.) Holttum
Snow et al. 2011:15
C-31
Synonymy List Arranged By Old Name
Appendix D. Synonymy list for Hawaiian native and naturalized vascular plant checklist, arranged alphabetically by old scientific name
This appendix is provided as a user-friendly alternative to the synonymy list
presented in Appendix A. While identical in content, this list reassembles the fields,
presenting old scientific names first in strictly alphabetical sequence, without regard
to their major group or family classification, followed by their current disposition.
The literature citation provides a source for the name change; while primary
sources were preferred, secondary sources are often cited. In some cases, both
primary and secondary sources are cited; these are denoted by a slash symbol
Old name (synonym, misapplied name)
between the two citations. Occasionally, a name has been changed, and later
changed again [e.g., Blechum brownei (Acanthaceae) became a synonym of B.
pyramidatum, which later was synonymized into Ruellia blechum]. In such cases, the
connection between the oldest and currently accepted names may not be apparent,
and citations, separated by a semicolon, are provided for the name change to an
intermediate name, followed by one for the current disposition.
Acacia farnesiana (L.) Willd.
Vachellia farnesiana (L.) Wight & Arn.
Dicot
Fabaceae
Literature source
Mabberley 2008:890/Wagner et al.
2012:36
Adenophorus hymenophylloides (Kaulf.) Hook. &
Grev. x A. tamariscinus (Kaulf.) Hook. & Grev.
Adenophorus xabbottiae W.H.Wagner
Pteridophyte
Grammitidaceae
Palmer 2003:36
Adenophorus xcarsonii Ranker
Pteridophyte
Grammitidaceae
Palmer 2003:36
Adenophorus xbishopii (W.H.Wagner) xxxxx
Pteridophyte
Grammitidaceae
Adenostemma viscosum J.R.Forst. & G.Forst.
Lachnagrostis filiformis (G.Forst.) Trin.
Dicot
Monocot
Asteraceae
Poaceae
Palmer 2003:41
King & Robinson 1987:60/ Herbst
& Wagner 1999:14
Snow 2008:39
Agrostis exarata Trin.
Polypogon viridis (Gouan) Breistr.
Schiedea lychnoides Hillebr.
Schiedea obovata (Sherff) W.L.Wagner & Weller
Schiedea trinervis (H.Mann) Pax & K.Hoffm.
Schiedea viscosa H.Mann
Alyxia stellata (J.R.Forst. & G.Forst.) Roem. & Schult.
Thelypteris globulifera (Brack.) C.F.Reed
Monocot
Monocot
Dicot
Dicot
Dicot
Dicot
Dicot
Pteridophyte
Poaceae
Poaceae
Caryophyllaceae
Caryophyllaceae
Caryophyllaceae
Caryophyllaceae
Apocynaceae
Thelypteridaceae
Snow 2008:38
Herbst & Clayton 1998:18, 33, 34
Wagner et al. 2005:51
Wagner et al. 2005:57
Wagner et al. 2005:60
Wagner et al. 2005:54
Middleton 2002:56
Snow et al. 2011:15
Ammannia coccinea Rottb.
Dicot
Lythraceae
Herbst & Wagner 1999:23
Monocot
Monocot
Poaceae
Poaceae
Snow & Lau 2010:46
Snow & Lau 2010:48
Monocot
Poaceae
Snow & Lau 2010:48
Monocot
Poaceae
Snow & Lau 2010:47
Adenophorus hymenophylloides (Kaulf.) Hook. &
Grev. x A. tripinnatifidus Gaudich.
Adenophorus oahuensis (Copel.) L.E.Bishop x A.
pinnatifidus Gaudich.
Adenostemma lavenia sensu Wagner et al. (1990), non
(L.) Kuntze
Agrostis avenacea J.G.Gmel.
Agrostis exarata Trin. var. monolepis (Torr.)
A.S.Hitchc.
Agrostis semiverticillata (Forssk.) C.Christens.
Alsinidendron lychnoides (Hillebr.) Sherff
Alsinidendron obovatum Sherff
Alsinidendron trinerve H.Mann
Alsinidendron viscosum (H.Mann) Sherff
Alyxia oliviformis Gaudich.
Amauropelta globulifera (Brack.) Holttum
Ammannia auriculata sensu Wagner et al. (1990), non
Willd.
Current disposition
Andropogon glomeratus sensu Herbst & Clayton
(1998), in part, non (Walter) Britton, Sterns & Poggenb. Andropogon bicornis L.
Andropogon virginicus L.
Andropogon virginicus L. var. virginicus
Andropogon virginicus sensu Herbarium Pacificum
Staff (1999), non L.
Hyparrhenia hirta (L.) Stapf
Andropogon virginicus sensu Starr et al. (2002), non L.
Andropogon glomeratus (Walter) Britton, Sterns &
Poggenb. var. pumilus (Vasey) L.H.Dewey
D-1
Group
Family
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Andropogon virginicus sensu Wagner et al. (1990), in
part, non L.
Aristea cf. compressa sensu Herbarium Pacificum Staff
(1999), non Buchinger ex Baker
Arthrostema ciliatum Pav. ex D.Don (spelling change)
Asparagus setaceus sensu Lorence et al. (1995), non
(Kunth) Jessop
Asplenium acuminatum Hook. & Arn. x A. aethiopicum
(Burm.f.) Bech.
Asplenium aethiopicum (Burm.f.) Bech. x A. polyodon
G.Forst.
Asplenium hobdyi W.H.Wagner x A. normale D.Don
Asplenium xlauii (W.H.Wagner) N.Snow
Asplenium polyodon G.Forst. x A. sphenotomum
Hillebr.
Aster subulatus Michx. var. sandwicensis (A.Gray)
A.G.Jones
Current disposition
Group
Family
Literature source
Andropogon bicornis L.
Monocot
Poaceae
Snow & Lau 2010:46
Aristea gerrardii Weim.
Arthrostemma ciliatum Pav. ex D.Don
Monocot
Dicot
Iridaceae
Melastomataceae
Staples et al. 2003:13
Imada et al. 2000:13
Asparagus plumosus Baker
Monocot
Liliaceae
Staples & Herbst 2005:682
Asplenium xwaikamoi W.H.Wagner
Pteridophyte
Aspleniaceae
Palmer 2003:76
Asplenium xkokeense W.H.Wagner
Asplenium xflagrum W.H.Wagner & D.D.Palmer
Asplenium xjoellaui N.Snow
Pteridophyte
Pteridophyte
Pteridophyte
Aspleniaceae
Aspleniaceae
Aspleniaceae
Palmer 2003:76
Palmer 2003:76
Snow 2011:161
Asplenium xsphenocookii W.H.Wagner
Symphyotrichum expansum (Poepp. ex Spreng.)
G.L.Nesom
Pteridophyte
Aspleniaceae
Dicot
Asteraceae
Palmer 2003:76
Nesom 1994:292; Sundberg
2004:907; Nesom 2005:2133
Begonia foliosa Kunth var. miniata sensu Wagner et al.
(1990), non (Planch.) L.B.Sm. & B.G.Shubert
Begonia reniformis sensu Wagner et al. (1990), non
Dryand.
Begonia foliosa Kunth
Dicot
Begoniaceae
Imada 2007:35
Begonia vitifolia Schott
Dicot
Begoniaceae
Blechnum hawaiianum Christenh.
Doodia kunthiana Gaudich.
Pteridophyte
Blechnaceae
Blechnum lyonii (O.Deg.) Christenh.
Doodia lyonii O.Deg.
Pteridophyte
Blechnaceae
Blechnum norfolkense Christenh.
Doodia kunthiana Gaudich.
Pteridophyte
Blechnaceae
Blechum brownei Juss.
Ruellia blechum L.
Dicot
Acanthaceae
Blechum pyramidatum (Lam.) Urb.
Boerhavia glabrata sensu Wagner et al. (1990), non
Blume
Ruellia blechum L.
Dicot
Acanthaceae
Staples et al. 2002:6
proposal to conserve name
Blechnum hawaiianum Christenh.
over B. norfolkense (Christenhusz
2011:1772)
proposed transfer to Blechnum
lyonii (O.Deg.) Christenh.
(Christenhusz et al. 2011:20)
proposed transfer to Blechnum
norfolkense Christenh.
(Christenhusz et al. 2011:20)
Wagner & Herbst 1995:14; Tripp
et al. 2009:900/Wagner et al.
2012:3
Tripp et al. 2009:900/Wagner et al.
2012:3
Boerhavia acutifolia (Choisy) J.W.Moore
Dicot
Nyctaginaceae
Boltonia indica (L.) Benth.
Kalimeris indica (L.) Sch.Bip. subsp. indica
Dicot
Asteraceae
D-2
Herbst & Wagner 1999:24
Gu & Hoch 1997:785/Herbst &
Wagner 1999:14
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Current disposition
Group
Family
Literature source
Zuloaga et al. 2003:630/ Clayton &
Snow 2010:60/ Wagner et al.
2012:88
Brachiaria brizantha (Hochst. ex A.Rich.) Stapf
Brachiaria brizantha sensu Starr et al. (2006), in part,
non (Hochst. ex A.Rich.) Stapf
Urochloa brizantha (Hochst. ex A.Rich.) R.D.Webster
Monocot
Poaceae
Urochloa decumbens (Stapf) R.D.Webster
Monocot
Poaceae
Brachiaria decumbens Stapf
Urochloa decumbens (Stapf) R.D.Webster
Monocot
Poaceae
Brachiaria distachya (L.) Stapf
Urochloa distachya (L.) T.Q.Nguyen
Monocot
Poaceae
Brachiaria mollis (Sw.) Parodi
Urochloa mollis (Sw.) O.Morrone & F.Zuloaga
Monocot
Poaceae
Brachiaria mutica (Forssk.) Stapf
Urochloa mutica (Forssk.) T.Q.Nguyen
Monocot
Poaceae
Brachiaria plantaginea (Link) Hitchc.
Urochloa plantaginea (Link) R.D.Webster
Monocot
Poaceae
Brachiaria reptans (L.) C.A.Gardner & C.E.Hubb.
Urochloa reptans (L.) Stapf
Monocot
Poaceae
Brachiaria subquadripara (Trin.) Hitchc.
Urochloa distachya (L.) T.Q.Nguyen
Monocot
Poaceae
Brassica campestris L.
Brassica campestris sensu Wagner et al. (1990), in part,
non L.
Brassica nigra sensu Wagner et al. (1990), in part, non
(L.) W.D.J.Koch
Bromus hordeaceus L. subsp. hordeaceus
Bromus hordeaceus L. subsp. molliformis (J.Lloyd)
Maire & Weiller
Bromus molliformis J.Lloyd
Bromus mollis L.
Bromus rigidus Roth
Bromus willdenowii Kunth
Brugmansia candida Pers.
Bruguiera gymnorhiza sensu Wagner et al. (1990), non
(L.) Savigny
Brassica rapa L.
Dicot
Brassicaceae
Snow & Lau 2010:49
Zuloaga et al. 2003:630/ Clayton &
Snow 2010:60/ Wagner et al.
2012:89
Zuloaga et al. 2003:631/ Wagner &
Herbst 2003:58/ Clayton & Snow
2010:60
Zuloaga et al. 2003:632/ Wagner &
Herbst 2003:58/ Clayton & Snow
2010:61
Zuloaga et al. 2003:632/ Wagner &
Herbst 2003:58/ Clayton & Snow
2010:61
Zuloaga et al. 2003:633/ Wagner &
Herbst 2003:58/ Clayton & Snow
2010:61
Zuloaga et al. 2003:633/ Wagner &
Herbst 2003:58/ Clayton & Snow
2010:61
Zuloaga et al. 2003:631/ Wagner &
Herbst 2003:58/ Clayton & Snow
2010:60
Al-Shehbaz 1985:290/Wagner &
Herbst 1995:17
Sinapis arvensis L.
Dicot
Brassicaceae
Herbarium Pacificum Staff 1998:9
Hirschfeldia incana (L.) Lagr.-Foss.
Bromus hordeaceus L.
Dicot
Monocot
Brassicaceae
Poaceae
Lorence et al. 1995:26
Snow 2008:39
Bromus hordeaceus L.
Bromus hordeaceus L.
Bromus hordeaceus L.
Bromus diandrus Roth
Bromus catharticus Vahl
Brugmansia xcandida Pers.
Monocot
Monocot
Monocot
Monocot
Monocot
Dicot
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Solanaceae
Snow 2008:39
Snow 2008:39
Snow 2008:39
Snow 2008:38
Herbst & Clayton 1998:20
Staples & Herbst 2005:530
Bruguiera sexangula (Lour.) Poir.
Dicot
Rhizophoraceae
Herbst & Wagner 1999:31
D-3
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Bryophyllum beauverdii (Raym.-Hamet) A.Berger
Kalanchoe beauverdii Raym.-Hamet
Dicot
Crassulaceae
Bryophyllum daigremontianum (Raym.-Hamet &
H.Perrier) A.Berger
Kalanchoe daigremontiana Raym.-Hamet & H.Perrier
Dicot
Crassulaceae
Bryophyllum fedtschenkoi (Raym.-Hamet & H.Perrier)
Lauz.-March.
Kalanchoe fedtschenkoi Raym.-Hamet & H.Perrier
Dicot
Crassulaceae
Bryophyllum pinnatum (Lam.) Oken
Kalanchoe pinnata (Lam.) Pers.
Dicot
Crassulaceae
Bryophyllum tubiflorum Harv.
Kalanchoe tubiflora (Harv.) Raym.-Hamet
Dicot
Crassulaceae
Caesalpinia kavaiensis H.Mann
Calotropis gigantea sensu Starr et al. (2003, 2004), in
part, non (L.) W.T.Aiton
Canavalia galeata sensu Wysong et al. (2007), non
(Gaudich.) Vogel
Canthium odoratum (G. Forst.) Seem.
Capsella rubella Reut.
Capsicum frutescens L.
Carex ovalis sensu Wagner et al. (1990), non Gooden
Mezoneuron kavaiense (H.Mann) Hillebr.
Dicot
Fabaceae
Literature source
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst 2003:21
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst 2003:21
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst 2003:21
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst 2003:21
Staples et al. 2002:9/Staples &
Herbst 2005:255; Mort et al.
2001:83/Wagner & Herbst 2003:21
Bruneau et al. 2008:711/ Wagner et
al. 2012:37
Calotropis procera (Aiton) W.T.Aiton
Dicot
Asclepiadaceae
Starr & Starr 2012:87
Canavalia molokaiensis O.Deg., I.Deg. & J.D.Sauer
Psydrax odorata (G.Forst.) A.C.Sm. & S.P.Darwin
Capsella bursa-pastoris (L.) Medik.
Capsicum annuum L.
Carex longii Mack.
Dicot
Dicot
Dicot
Dicot
Monocot
Fabaceae
Rubiaceae
Brassicaceae
Solanaceae
Cyperaceae
Carex nealae R.W.Krauss
Thevetia peruviana (Pers.) K.Schum.
Schenkia sebaeoides Griseb.
Monocot
Dicot
Dicot
Cyperaceae
Apocynaceae
Gentianaceae
Centrosema molle Mart. ex Benth.
Euphorbia albomarginata Torr. & A.Gray
Dicot
Dicot
Fabaceae
Euphorbiaceae
Euphorbia serpens Kunth
Euphorbia arnottiana Endl.
Euphorbia atrococca A.Heller
Dicot
Dicot
Dicot
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Wagner et al. 2012:37
Smith 1988:230
Wagner & Herbst 1995:17
Staples & Herbst 2005:532
Wagner & Herbst 2003:51
Dragon & Barrington 2009:1898,
1906/ Wagner et al. 2012:79
Wagner & Herbst 1999:1858
Mansion 2004:725
Fantz 1996:330/Wagner & Herbst
2003:24
Govaerts et al. 2000:667
Wagner et al. 1997:56; Govaerts et
al. 2000:832
Govaerts et al. 2000:674
Govaerts et al. 2000:676
Euphorbia celastroides Boiss. var. amplectens Sherff
Dicot
Euphorbiaceae
Govaerts et al. 2000:692
Euphorbia celastroides Boiss. var. celastroides
Dicot
Euphorbiaceae
Govaerts et al. 2000:692
Carex thunbergii sensu Wagner et al. (1990), non Steud.
Cascabela peruviana (Pers.) Raf.
Centaurium sebaeoides (Griseb.) Druce
Centrosema pubescens sensu Flynn & Lorence (1998),
non Benth.
Chamaesyce albomarginata (Torr. & A.Gray) Small
Chamaesyce albomarginata sensu Wagner et al. (1990),
in part, non (Torr. & A.Gray) Small
Chamaesyce arnottiana (Endl.) O.Deg. & I.Deg.
Chamaesyce atrococca (A.Heller) Croizat & O.Deg.
Chamaesyce celastroides (Boiss.) Croizat & O.Deg. var.
amplectens (Sherff) O.Deg. & I.Deg.
Chamaesyce celastroides (Boiss.) Croizat & O.Deg. var.
celastroides
Current disposition
D-4
Group
Family
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Current disposition
Group
Family
Literature source
Chamaesyce celastroides (Boiss.) Croizat & O.Deg. var.
hanapepensis (Sherff) O.Deg. & I.Deg.
Euphorbia celastroides Boiss. var. hanapepensis Sherff
Dicot
Euphorbiaceae
Govaerts et al. 2000:692
Chamaesyce celastroides (Boiss.) Croizat & O.Deg. var.
kaenana (Sherff) O.Deg. & I.Deg.
Euphorbia celastroides Boiss. var. kaenana Sherff
Dicot
Euphorbiaceae
Govaerts et al. 2000:692
Chamaesyce celastroides (Boiss.) Croizat & O.Deg. var.
laehiensis (O.Deg., I.Deg. & Sherff) Koutnik
Euphorbia celastroides Boiss. var. laehiensis O.Deg.,
I.Deg. & Sherff
Dicot
Euphorbiaceae
Govaerts et al. 2000:692
Chamaesyce celastroides (Boiss.) Croizat & O.Deg. var.
lorifolia (A.Gray) O.Deg. & I.Deg.
Euphorbia celastroides Boiss. var. lorifolia A.Gray
Dicot
Euphorbiaceae
Govaerts et al. 2000:692
Chamaesyce celastroides (Boiss.) Croizat & O.Deg. var.
stokesii (C.N.Forbes) O.Deg. & I.Deg.
Euphorbia celastroides Boiss. var. stokesii (C.N.Forbes)
Sherff
Dicot
Euphorbiaceae
Govaerts et al. 2000:692
Chamaesyce celastroides (Boiss.) Croizat & O.Deg. var.
tomentella (Boiss.) Koutnik
Chamaesyce clusiifolia (Hook. & Arn.) Arthur
Chamaesyce degeneri (Sherff) Croizat & O.Deg.
Chamaesyce deppeana (Boiss.) Millsp.
Euphorbia celastroides Boiss. var. tomentella (Boiss.)
Oudejans
Euphorbia clusiifolia Hook. & Arn.
Euphorbia degeneri Sherff
Euphorbia deppeana Boiss.
Euphorbia eleanoriae (Lorence & W.L.Wagner)
Govaerts
Euphorbia halemanui Sherff
Euphorbia herbstii (W.L.Wagner) Oudejans
Euphorbia hirta L.
Euphorbia hypericifolia L.
Euphorbia hyssopifolia L.
Dicot
Dicot
Dicot
Dicot
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Govaerts et al. 2000:692
Govaerts et al. 2000:698
Govaerts et al. 2000:709
Govaerts et al. 2000:712
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Govaerts et al. 2000:717
Govaerts et al. 2000:742
Govaerts et al. 2000:745
Govaerts et al. 2000:320
Govaerts et al. 2000:320
Govaerts et al. 2000:320
Euphorbia kuwaleana O.Deg. & Sherff
Euphorbia maculata L.
Dicot
Dicot
Euphorbiaceae
Euphorbiaceae
Govaerts et al. 2000:764
Govaerts et al. 2000:322
Chamaesyce multiformis (Hook. & Arn.) Croizat &
Deg. var. microphylla (Boiss.) O.Deg. & I.Deg.
Chamaesyce multiformis (Hook. & Arn.) Croizat &
Deg. var. multiformis
Chamaesyce olowaluana (Sherff) Croizat & O.Deg.
Chamaesyce prostrata (Aiton) Small
Euphorbia multiformis Hook. & Arn. var. microphylla
Boiss.
Dicot
Euphorbiaceae
Govaerts et al. 2000:783
Euphorbia multiformis Hook. & Arn. var. multiformis
Euphorbia olowaluana Sherff
Euphorbia prostrata Aiton
Dicot
Dicot
Dicot
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Govaerts et al. 2000:783
Govaerts et al. 2000:793
Govaerts et al. 2000:324
Chamaesyce remyi (A.Gray ex Boiss.) Croizat &
O.Deg. var. hanaleiensis (Sherff) O.Deg. & I.Deg.
Chamaesyce remyi (A.Gray ex Boiss.) Croizat &
O.Deg. var. kauaiensis (O.Deg. & Sherff) O.Deg. &
I.Deg.
Chamaesyce remyi (A.Gray ex Boiss.) Croizat &
O.Deg. var. remyi
Euphorbia remyi A.Gray ex Boiss. var. hanaleiensis
Sherff
Dicot
Euphorbiaceae
Govaerts et al. 2000:818
Euphorbia remyi A.Gray ex Boiss. var. kauaiensis O.
Deg. & Sherff
Dicot
Euphorbiaceae
Govaerts et al. 2000:818
Euphorbia remyi A.Gray ex Boiss. var. remyi
Dicot
Euphorbiaceae
Govaerts et al. 2000:818
Chamaesyce eleanoriae Lorence & W.L.Wagner
Chamaesyce halemanui (Sherff) Croizat & O.Deg.
Chamaesyce herbstii W.L.Wagner
Chamaesyce hirta (L.) Millsp.
Chamaesyce hypericifolia (L.) Millsp.
Chamaesyce hyssopifolia (L.) Small
Chamaesyce kuwaleana (O.Deg. & Sherff) O.Deg. &
I.Deg.
Chamaesyce maculata (L.) Small
D-5
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Chamaesyce rockii (C.N.Forbes) Croizat & O.Deg.
Chamaesyce serpens (Kunth) Small
Chamaesyce skottsbergii (Sherff) Croizat & O.Deg. var.
skottsbergii
Current disposition
Euphorbia rockii C.N.Forbes
Euphorbia serpens Kunth
Group
Dicot
Dicot
Family
Euphorbiaceae
Euphorbiaceae
Literature source
Govaerts et al. 2000:821
Govaerts et al. 2000:832
Euphorbia skottsbergii Sherff var. skottsbergii
Dicot
Euphorbiaceae
Govaerts et al. 2000:835
Chamaesyce skottsbergii (Sherff) Croizat & O.Deg. var.
vaccinioides (Sherff) Koutnik
Chamaesyce sparsiflora (A.Heller) Koutnik
Chamaesyce thymifolia (L.) Millsp.
Euphorbia skottsbergii Sherff var. vaccinioides Sherff
Euphorbia sparsiflora A.Heller
Euphorbia thymifolia L.
Dicot
Dicot
Dicot
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Chenopodium ambrosioides L.
Dysphania ambrosioides (L.) Mosyakin & Clemants
Dicot
Chenopodiaceae
Chenopodium carinatum R.Br.
Chloris petraea Sw.
Christella boydiae (D.C.Eaton) Holttum
Christella cyatheoides (Kaulf.) Holttum
Dysphania carinata (R.Br.) Mosyakin & Clemants
Eustachys petraea (Sw.) Desv.
Cyclosorus boydiae (D.C.Eaton) W.H.Wagner
Cyclosorus cyatheoides (Kaulf.) Farw.
Dicot
Monocot
Pteridophyte
Pteridophyte
Chenopodiaceae
Poaceae
Thelypteridaceae
Thelypteridaceae
Govaerts et al. 2000:835
Govaerts et al. 2000:837
Govaerts et al. 2000:848
Clemants & Mosyakin 2003:
270/Wagner et al. 2012:28
Clemants & Mosyakin 2003:
274/Wagner et al. 2012:28
Wagner & Herbst 1995:24
Snow et al. 2011:15
Snow et al. 2011:15
Cyclosorus xpalmeri (W.H.Wagner) W.H.Wagner
Cyclosorus dentatus (Forssk.) Ching
Pteridophyte
Pteridophyte
Thelypteridaceae
Thelypteridaceae
Snow et al. 2011:15
Snow et al. 2011:15
Cyclosorus xintermedia W.H.Shieh & J.L.Tsai
Pteridophyte
Thelypteridaceae
Snow et al. 2011:15
Cyclosorus xintermedia W.H.Shieh & J.L.Tsai
Cyclosorus xpalmeri (W.H.Wagner) W.H.Wagner
Cyclosorus parasiticus (L.) Farw.
Cyclosorus wailele (Flynn) W.H.Wagner
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Thelypteridaceae
Thelypteridaceae
Thelypteridaceae
Thelypteridaceae
Leucanthemum vulgare Lam.
Tanacetum parthenium (L.) Sch.Bip.
Cibotium xheleniae D.D.Palmer
Cinnamomum burmanni (Nees) Blume
Cissus quadrangularis L.
Clermontia peleana Rock subsp. peleana
Clermontia peleana Rock subsp. singuliflora (Rock)
Lammers
Dicot
Dicot
Pteridophyte
Dicot
Dicot
Dicot
Asteraceae
Asteraceae
Dicksoniaceae
Lauraceae
Vitaceae
Campanulaceae
Snow et al. 2011:15
Snow et al. 2011:15
Snow et al. 2011:15
Snow et al. 2011:15
Soreng & Cope 1991:162/Herbst &
Wagner 1999:15
Herbst & Wagner 1999:15
Palmer 2003:98
Wagner & Herbst 2003:32
Frohlich & Lau 2008:10
Lammers 1991:34
Dicot
Campanulaceae
Lammers 1991:32
Dicot
Dicot
Campanulaceae
Verbenaceae
Lammers 1991:25
Wagner & Herbst 1995:26
Dicot
Verbenaceae
Staples & Herbst 2005:559
Christella cyatheoides (Kaulf.) Holttum x C. dentata
(Forssk.) Brownsey & Jermy
Christella dentata (Forssk.) Brownsey & Jermy
Christella dentata (Forssk.) Brownsey & Jermy x C.
parasitica (L.) H.Lév.
Christella xintermedia (W.H.Shieh & J.L.Tsai)
D.D.Palmer
Christella xpalmeri W.H.Wagner
Christella parasitica (L.) H.Lév.
Christella wailele (Flynn) D.D.Palmer
Chrysanthemum leucanthemum L.
Chrysanthemum parthenium (L.) Bernh.
Cibotium chamissoi Kaulf. x C. menziesii Hook.
Cinnamomum burmanii (Nees) Blume (spelling change)
Cissus quadrangula L.
Clermontia peleana Rock
Clermontia singuliflora (Rock) Rock
Clermontia waikoluensis H.St.John
Clerodendrum philippinum Schauer
Clerodendrum speciosissimum van Geert
Clermontia arborescens (H.Mann) Hillebr. subsp.
waikoluensis (H.St.John) Lammers
Clerodendrum chinense (Osbeck) Mabb.
Clerodendrum buchananii (Roxb.) Walp. var. fallax
(Lindl.) Bakh.
D-6
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Clitoria ternata L. (spelling change)
Cocculus trilobus (Thunb.) DC.
Current disposition
Clitoria ternatea L.
Cocculus orbiculatus (L.) DC.
Group
Dicot
Dicot
Family
Fabaceae
Menispermaceae
Coleogeton pectinatus (L.) Les & R.R.Haynes
Coprosma granadensis (L.f.) Heads
Stuckenia pectinata (L.) Börner
Nertera granadensis (L.f.) Druce
Monocot
Dicot
Potamogetonaceae
Rubiaceae
Costus speciosus (J.König) Sm.
Cheilocostus speciosus (J.König) C.Specht
Monocot
Costaceae
Literature source
Staples & Herbst 2005:313
Wagner & Herbst 1995:22
Haynes et al. 1998:241/Herbst &
Wagner 1999:30
Wagner et al. 2012:65
Specht & Stevenson 2006:159/
Wagner et al. 2012:79
Cyanea lobata H.Mann subsp. baldwinii (C.N.Forbes &
G.C.Munro) H.St.John
Cyanea hirtella (H.Mann) Hillebr.
Dicot
Dicot
Campanulaceae
Campanulaceae
Lammers 1999:387
Wagner & Herbst 2003:15
Cyanea floribunda E.Wimm.
Cyanea dunbariae Rock
Cyanea coriacea (A.Gray) Hillebr.
Dicot
Dicot
Dicot
Campanulaceae
Campanulaceae
Campanulaceae
Cyanea cylindrocalyx (Rock) Lammers
Dicot
Campanulaceae
Cyanea grimesiana Gaudich. var. mauiensis Rock
Cyanea mauiensis (Rock) Lammers
Dicot
Campanulaceae
Cyanea grimesiana Gaudich. var. munroi Hosaka
Cyanea lanceolata (Gaudich.) Lammers, Givnish &
Sytsma subsp. calycina (Cham.) Lammers, Givnish &
Sytsma
Cyanea lobata H.Mann
Cyanea spathulata (Hillebr.) A.Heller subsp.
longipetiolata Lammers
Cyanea spathulata (Hillebr.) A.Heller subsp. spathulata
Cyanea sylvestris A.Heller
Cyperus alternifolius L. subsp. flabelliformis (Rottb.)
Kük.
Cyperus halpan L. (spelling change)
Cyrtandra confertiflora (Wawra) C.B.Clarke var.
confertiflora
Cyrtandra confertiflora (Wawra) C.B.Clarke var.
obovata (Wawra) C.B.Clarke
Cyrtandra limahuliensis H.St.John
Cyanea munroi (Hosaka) Lammers
Dicot
Campanulaceae
Wagner & Herbst 2003:15
Herbst & Wagner 1999:16
Lammers 2004:96
Lammers 1998:31/Herbst &
Wagner 1999:16
Lammers 1998:31/Herbst &
Wagner 1999:17
Lammers 1998:31/Herbst &
Wagner 1999:17
Cyanea calycina (Cham.) Lammers
Cyanea lobata H.Mann subsp. lobata
Dicot
Dicot
Campanulaceae
Campanulaceae
Cyanea kahiliensis (H.St.John) Lammers
Cyanea spathulata (Hillebr.) A.Heller
Cyanea hirtella (H.Mann) Hillebr.
Dicot
Dicot
Dicot
Campanulaceae
Campanulaceae
Campanulaceae
Cyperus involucratus Rottb.
Cyperus haspan L.
Monocot
Monocot
Cyperaceae
Cyperaceae
Lammers 1998:32/Herbst &
Wagner 1999:16
Lammers 1999:387
Lammers 1998:33/Herbst &
Wagner 1999:16
Lammers 2004:100
Wagner et al. 2005– website
Tucker 1994:20/Strong & Wagner
1997:41
Tucker 1994:50/Hughes 1995:4
Cyrtandra confertiflora (Wawra) C.B.Clarke
Dicot
Gesneriaceae
Wagner & Herbst 2003:29
Cyrtandra heinrichii H.St.John
Cyrtandra kealiae Wawra subsp. kealiae
Dicot
Dicot
Gesneriaceae
Gesneriaceae
Danthonia pilosa sensu Wagner et al. (1990), non R.Br.
Danthonia semiannularis (Labill.) R.Br.
Rytidosperma penicillatum (Labill.) Connor & Edgar
Rytidosperma semiannulare (Labill.) Connor & Edgar
Monocot
Monocot
Poaceae
Poaceae
Delissea eleeleensis H.St.John
Cyanea eleeleensis (H.St.John) Lammers
Dicot
Campanulaceae
Wagner & Herbst 2003:29
Wagner & Lorence 2000:17
Herbst & Clayton 1998:34;
Darbyshire et al. 2010:670
Herbst & Clayton 1998:34
Lammers 1992:129/Herbst &
Wagner 1999:16
Cyanea baldwinii C.N.Forbes & G.C.Munro
Cyanea degeneriana E.Wimm.
Cyanea degeneriana sensu Wagner et al. (1990), non
E.Wimm.
Cyanea dunbarii Rock (spelling change)
Cyanea fauriei H.Lév.
Cyanea grimesiana Gaudich. subsp. cylindrocalyx
(Rock) Lammers
D-7
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Current disposition
Group
Family
Literature source
Lammers 1998:33/Herbst &
Wagner 1999:16
Lammers 1998:33/Herbst &
Wagner 1999:16
Lammers 1992:130/Herbst &
Wagner 1999:16
Delissea habenata H.St.John
Cyanea habenata (H.St.John) Lammers
Dicot
Campanulaceae
Delissea kahiliensis H.St.John
Cyanea kahiliensis (H.St.John) Lammers
Dicot
Campanulaceae
Delissea kolekoleensis H.St.John
Delissea niihauensis H.St.John subsp. kauaiensis
(Lammers) Lammers
Delissea niihauensis H.St.John subsp. niihauensis
Delissea rivularis (Rock) E.Wimm.
Delissea sinuata Hillebr. subsp. lanaiensis (Rock)
Lammers
Delissea sinuata Hillebr. subsp. sinuata
Delissea subcordata Gaudich.
Cyanea kolekoleensis (H.St.John) Lammers
Dicot
Campanulaceae
Delissea kauaiensis (Lammers) Lammers
Delissea niihauensis H.St.John
Cyanea rivularis Rock
Dicot
Dicot
Dicot
Campanulaceae
Campanulaceae
Campanulaceae
Delissea lanaiensis (Rock) Lammers
Delissea sinuata Hillebr.
Delissea subcordata Gaudich. subsp. subcordata
Delissea subcordata Gaudich. subsp. obtusifolia
(Wawra) Lammers
Delissea kauaiensis (Lammers) Lammers
Dicot
Dicot
Dicot
Campanulaceae
Campanulaceae
Campanulaceae
Lammers 2005:19
Lammers 2005:17
Lammers 2005:62
Lammers 1998:34/Herbst &
Wagner 1999:18
Lammers 1998:34
Lammers 2005:33
Dicot
Dicot
Campanulaceae
Campanulaceae
Lammers 2005:34
Lammers 2005:19
Delissea niihauensis H.St.John
Delissea undulata Gaudich.
Desmanthus pernambucanus (L.) Thell.
Dichelachne micrantha (Cav.) Domin
Dicot
Dicot
Dicot
Monocot
Campanulaceae
Campanulaceae
Fabaceae
Poaceae
Asplenium dielerectum Viane
Asplenium dielerectum Viane
Pteridophyte
Pteridophyte
Aspleniaceae
Aspleniaceae
Diellia erecta Brack. fma. pumila (Brack.) W.H.Wagner
Diellia falcata Brack.
Diellia falcata Brack. x D. unisora W.H.Wagner
Diellia laciniata (Hillebr.) Diels
Diellia xlauii W.H.Wagner
Diellia leucostegioides (Baker) W.H.Wagner
Diellia mannii (D.C.Eaton) W.J.Rob.
Diellia pallida W.H.Wagner
Diellia unisora W.H.Wagner
Digitaria pentzii Stent
Doryopteris decipiens (Hook.) J.Sm. x D. decora Brack.
Asplenium dielerectum Viane
Asplenium dielfalcatum Viane
Asplenium xjoellaui N.Snow
Asplenium diellaciniatum Viane
Asplenium xjoellaui N.Snow
Asplenium leucostegioides Baker
Asplenium dielmannii Viane
Asplenium dielpallidum N.Snow
Asplenium unisorum (W.H.Wagner) Viane
Digitaria eriantha Steud.
Doryopteris subdecipiens W.H.Wagner
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Monocot
Pteridophyte
Aspleniaceae
Aspleniaceae
Aspleniaceae
Aspleniaceae
Aspleniaceae
Aspleniaceae
Aspleniaceae
Aspleniaceae
Aspleniaceae
Poaceae
Pteridaceae
Dracaena aurea H.Mann
Pleomele aurea (H.Mann) N.E.Br.
Monocot
Agavaceae
Lammers 2005:17
Lammers 2005:21
Wagner & Herbst 1995:20
Herbst & Clayton 1998:22
Snow et al. 2011:12 (status of f.
alexandri under study)
Snow et al. 2011:12
Snow et al. 2011:12 (status of f.
pumila under study)
Snow et al. 2011:12
Snow 2011:161
Wagner et al. 2012:103
Snow 2011:161
Snow et al. 2011:12
Snow et al. 2011:12
Snow et al. 2011:12
Snow et al. 2011:12
Herbst & Clayton 1998:23
Palmer 2003:133
proposed transfer to Dracaena
aurea H.Mann (Jankalski 2008:20)
Delissea subcordata Gaudich. var. obtusifolia Wawra
Delissea undulata Gaudich. subsp. kauaiensis Lammers
Delissea undulata Gaudich. subsp. niihauensis
(H.St.John) Lammers
Delissea undulata Gaudich. subsp. undulata
Desmanthus virgatus (L.) Willd.
Dichelachne sciurea (R.Br.) Hook.f.
Diellia erecta Brack. fma. alexandri (Hillebr.)
W.H.Wagner
Diellia erecta Brack. fma. erecta
D-8
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Dracaena fernaldii (H.St.John) Jankal.
Pleomele fernaldii H.St.John
Monocot
Agavaceae
Dracaena forbesii (O.Deg.) Jankal.
Pleomele forbesii O.Deg.
Monocot
Agavaceae
Dracaena halaapepe (H.St.John) Jankal.
Pleomele halapepe H.St.John
Monocot
Agavaceae
Dracaena halemanuensis Jankal.
Pleomele aurea (H.Mann) N.E.Br.
Monocot
Agavaceae
Dracaena konaensis (H.St.John) Jankal.
Pleomele hawaiiensis O.Deg. & I.Deg.
Monocot
Agavaceae
Dracaena rockii (H.St.John) Jankal.
Drypetes sherffii Govaerts & Radcl.-Sm.
Pleomele auwahiensis H.St.John
Xylosma hawaiiense Seem.
Monocot
Dicot
Agavaceae
Flacourtiaceae
Dubautia dolosa (O.Deg. & Sherff) G.D.Carr
Echinochloa stagnina sensu Wagner et al. (1990), non
(Retz.) P.Beauv.
Echinochloa stagnina sensu Wagner et al. (1990), non
(Retz.) P.Beauv.
Echinochloa utilis Ohwi & Yabuno
Echinochloa walteri sensu Wagner et al. (1990), non
(Pursh) A.Heller
Echinodorus rostratus (Nutt.) Engelm.
Eclipta alba (L.) Hassk.
Dubautia waianapanapaensis G.D.Carr
Dicot
Asteraceae
Literature source
proposed transfer to Dracaena
fernaldii (H.St.John) Jankal.
(Jankalski 2008:20)
proposed transfer to Dracaena
forbesii (O.Deg.) Jankal. (Jankalski
2008:21)
proposed transfer to Dracaena
halaapepe (H.St.John) Jankal.
(Jankalski 2008:21; note spelling
change)
proposed resurrection of Pleomele
stenophylla H.St.John (as Dracaena
halemanuensis Jankal. (Jankalski
2008:21)
proposed syn. under Dracaena
konaensis (H.St.John) Jankal.,
along with Pleomele konaensis
H.St.John (Jankalski 2008:21)
proposed syn. under Dracaena
rockii (H.St.John) Jankal., along
with Pleomele rockii H.St.John
(Jankalski 2008:21)
Herbst & Wagner 1999:21
Carr 1995:1580/Wagner & Herbst
2003:7
Echinochloa oryzoides (Ard.) Fritsch
Monocot
Poaceae
Herbst & Clayton 1998:24
Echinochloa picta (J.König) P.W.Michael
Echinochloa esculenta (A.Braun) H.Scholz
Monocot
Monocot
Poaceae
Poaceae
Herbst & Clayton 1998:24
Snow 2008:39
Echinochloa crusgalli (L.) P.Beauv.
Echinodorus berteroi (Speng.) Fassett
Eclipta prostrata (L.) L.
Pseudelephantopus spicatus (B.Juss. ex Aubl.)
C.F.Baker
Leymus triticoides (Buckley) Pilg.
Monocot
Monocot
Dicot
Poaceae
Alismataceae
Asteraceae
Herbst & Clayton 1998:24
The Plant List website 2012
Wagner & Herbst 1995:16
Dicot
Monocot
Asteraceae
Poaceae
Herbst & Wagner 1999:15
Soreng et al. 2003:425
Emex spinosa (L.) Campd.
Eragrostis variabilis (Gaudich.) Steud.
Eragrostis leptostachya (R.Br.) Steud.
Dicot
Monocot
Monocot
Polygonaceae
Poaceae
Poaceae
Parker & Parsons 2012b:71
Snow & Lau 2010:50
Herbst & Clayton 1998:26,27
Elephantopus spicatus Juss. ex Aubl.
Elymus triticoides Buckley
Emex australis sensu Wagner et al. (1990), in part, non
Steinh.
Eragrostis hobdyi H.St.John
Eragrostis hosakai O.Deg.
Current disposition
D-9
Group
Family
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Eragrostis leptophylla sensu Wagner et al. (1990), in
part, non Hitchc.
Eragrostis molokaiensis H.St.John
Eragrostis monticola sensu Wagner et al. (1990), in part,
non (Gaudich.) Hillebr.
Eragrostis pectinacea (Michx.) Nees
Eragrostis pectinacea sensu Wagner et al. (1990), in
part, non (Michx.) Nees
Eragrostis tenella (L.) P.Beauv. ex Roem. &
J.A.Schultes
Eriocaulon sp. A
Eucalyptus calophylla R.Br. ex Lindl.
Eucalyptus citriodora Hook.
Eucalyptus ficifolia F.Muell.
Eucalyptus gummifera (Sol. ex Gaertn.) Hochr.
Festuca arundinacea Schreb.
Ficus rubiginosa sensu Nagata (1995), non Desf. ex
Vent.
Filago gallica L.
Fimbristylis miliacea (L.) Vahl
Fuchsia hybrida Hort. ex Siebold & Voss
Gahnia gahniiformis (Gaudich.) A.Heller
Gahnia lanaiensis O.Deg., I.Deg. & J.Kern
Garnotia acutigluma sensu Lorence & Flynn (1999),
non (Steud.) Ohwi
Geranium humile Hillebr.
Glyceria fluitans sensu Wagner et al. (1990), non (L.)
R.Br.
Glycine wightii (Wight & Arn.) Verdc.
Gnaphalium japonicum sensu Wagner et al. (1990), non
Euchiton japonicus (Thunb.) A.Anderb.
Gnaphalium japonicum Thunb.
Gnaphalium purpureum L.
Current disposition
Group
Family
Literature source
Eragrostis pilosa (L.) P.Beauv.
Eragrostis brownii (Kunth) Nees ex Steud.
Monocot
Monocot
Poaceae
Poaceae
Herbst & Clayton 1998: 27
Herbst & Clayton 1998:26,27
Eragrostis pilosa (L.) P.Beauv.
Eragrostis pectinacea (Michx.) Nees var. pectinacea
Monocot
Monocot
Poaceae
Poaceae
Herbst & Clayton 1998: 27
Snow & Lau 2010:51
Eragrostis pilosa (L.) P.Beauv.
Monocot
Poaceae
Herbst & Clayton 1998: 27
Eragrostis amabilis (L.) Wight & Arn. ex Nees
Eriocaulon scariosum Sm.
Corymbia calophylla (R.Br. ex Lindl.) K.D.Hill &
L.A.S.Johnson
Monocot
Monocot
Poaceae
Eriocaulaceae
Dicot
Myrtaceae
Dicot
Myrtaceae
Dicot
Myrtaceae
Dicot
Monocot
Myrtaceae
Poaceae
Lorence et al. 1995:45
Herbst & Wagner 1999:20
Hill & Johnson 1995:240/ Herbst
& Wagner 1999:24
Hill & Johnson 1995:388/ Herbst
& Wagner 1999:24
Hill & Johnson 1995:245/ Herbst
& Wagner 1999:24
Hill & Johnson 1995:233/ Herbst
& Wagner 1999:24
Snow 2008:40
Ficus cf. platypoda (A.Cunn. ex Miq.) A.Cunn. ex Miq.
Logfia gallica (L.) Coss. & Germ.
Fimbristylis littoralis Gaudich.
Fuchsia xhybrida Vilm.
Dicot
Dicot
Monocot
Dicot
Moraceae
Asteraceae
Cyperaceae
Onagraceae
Morelotia gahniiformis Gaudich.
Gahnia lacera (A.Rich.) Steud.
Monocot
Monocot
Cyperaceae
Cyperaceae
Dichelachne micrantha (Cav.) Domin
Monocot
Poaceae
Geranium hillebrandii Aedo & Muñoz Garm.
Dicot
Geraniaceae
Snow & Davidse 2011:18
Aedo & Garmendia 1997:725/
Herbst & Wagner 1999:21
Glyceria notata Chevall.
Neonotonia wightii (Wight & Arn.) Lackey
Monocot
Dicot
Poaceae
Fabaceae
Snow 2008:39
Herbarium Pacificum Staff 1996:4
Euchiton sphaericus (Willd.) A.Anderb.
Euchiton japonicus (Thunb.) A.Anderb.
Gamochaeta purpurea (L.) Cabrera
Dicot
Dicot
Dicot
Asteraceae
Asteraceae
Asteraceae
Wagner et al. 1997:54
Wagner et al. 1997:54
Wagner et al. 1997:54
Corymbia citriodora (Hook.) K.D.Hill & L.A.S.Johnson
Corymbia ficifolia (F.Muell.) K.D.Hill &
L.A.S.Johnson
Corymbia gummifera (Sol. ex Gaertn.) K.D.Hill &
L.A.S.Johnson
Schedonorus arundinaceus (Schreb.) Dumort.
D-10
Oppenheimer & Bartlett 2000:7
Wagner et al. 1997:54
Strong 2004:1069/Imada 2007:35
Staples & Herbst 2005:445
Lorence et al. 1995:34/Wagner &
Herbst 2003:52
Koyama et al. 2010:30
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Gnaphalium sandwicensium Gaudich. var. molokaiense
O.Deg. & Sherff
Gnaphalium sandwicensium Gaudich. var.
sandwicensium
Current disposition
Pseudognaphalium sandwicensium (Gaudich.)
A.Anderb. var. hawaiiense (O.Deg. & Sherff)
W.L.Wagner
Pseudognaphalium sandwicensium (Gaudich.)
A.Anderb. var. kilaueanum (O.Deg. & Sherff)
W.L.Wagner
Pseudognaphalium sandwicensium (Gaudich.)
A.Anderb. var. molokaiense (O.Deg. & Sherff)
W.L.Wagner
Pseudognaphalium sandwicensium (Gaudich.)
A.Anderb. var. sandwicensium
Gonocormus minutus (Blume) Bosch
Crepidomanes minutum (Blume) K.Iwats.
Pteridophyte
Hymenophyllaceae
Gonocormus prolifer (Blume) Prantl
Grammitis baldwinii (Baker) Copel.
Grammitis forbesiana W.H.Wagner
Grammitis hookeri (Brack.) Copel.
Crepidomanes proliferum (Blume) Bostock
Oreogrammitis baldwinii (Baker) Parris
Oreogrammitis forbesiana (W.H.Wagner) Parris
Oreogrammitis hookeri (Brack.) Parris
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Hymenophyllaceae
Grammitidaceae
Grammitidaceae
Grammitidaceae
Grammitis tenella Kaulf.
Habenaria sp.
Adenophorus tenellus (Kaulf.) Ranker
Habenaria rodeiensis Barb.Rodr.
Pteridophyte
Monocot
Grammitidaceae
Orchidaceae
Hedyotis acuminata (Cham. & Schltdl.) Steud.
Dicot
Rubiaceae
Hedyotis callitrichoides (Griseb.) W.H.Lewis
Kadua acuminata Cham. & Schltdl.
Oldenlandiopsis callitrichoides (Griseb.) Terrell &
W.H.Lewis
Dicot
Rubiaceae
Hedyotis centranthoides (Hook. & Arn.) Steud.
Kadua centranthoides Hook. & Arn.
Dicot
Rubiaceae
Hedyotis cookiana (Cham. & Schltdl.) Steud.
Kadua cookiana Cham. & Schltdl.
Dicot
Rubiaceae
Hedyotis coriacea Sm.
Kadua coriacea (Sm.) W.L.Wagner & Lorence
Dicot
Rubiaceae
Hedyotis corymbosa (L.) Lam.
Dicot
Rubiaceae
Dicot
Rubiaceae
Hedyotis degeneri Fosberg var. degeneri
Oldenlandia corymbosa L.
Kadua degeneri (Fosberg) W.L.Wagner & Lorence
subsp. coprosmifolia (Fosberg) W.L.Wagner & Lorence
Kadua degeneri (Fosberg) W.L.Wagner & Lorence
subsp. degeneri
Dicot
Rubiaceae
Hedyotis elatior (H.Mann) Fosberg
Kadua elatior (H.Mann) W.L.Wagner & Lorence
Dicot
Rubiaceae
Hedyotis fluviatilis (C.N.Forbes) Fosberg
Kadua fluviatilis C.N.Forbes
Dicot
Kadua flynnii (W.L.Wagner & Lorence) W.L.Wagner &
Lorence
Dicot
Rubiaceae
Gnaphalium sandwicensium Gaudich. var. hawaiiense
O.Deg. & Sherff
Gnaphalium sandwicensium Gaudich. var. kilaueanum
O.Deg. & Sherff
Hedyotis degeneri Fosberg var. coprosmifolia Fosberg
Hedyotis flynnii W.L.Wagner & Lorence
D-11
Group
Family
Literature source
Dicot
Asteraceae
Wagner et al. 1997:54
Dicot
Asteraceae
Wagner et al. 1997:54
Dicot
Asteraceae
Wagner et al. 1997:54
Dicot
Asteraceae
Wagner et al. 1997:54
Ebihara et al. 2006:238/Snow et al.
2011:13
Ebihara et al. 2006:238/Snow et al.
2011:13
Parris 2007:256
Parris 2007:260
Parris 2007:260
Ranker 2008:171/Snow et al.
2011:14
Herbst & Wagner 1999:24
Terrell et al. 2005:832/ Kennedy et
al. 2010:23
Terrell & Lewis 1990:185/ Wagner
et al. 2012:63
Terrell et al. 2005:832/ Kennedy et
al. 2010:23
Terrell et al. 2005:833/ Kennedy et
al. 2010:23
Terrell et al. 2005:832/ Kennedy et
al. 2010:23
Groeninchkx et al. 2009:125/
Wagner et al. 2012:64
Terrell et al. 2005:833/ Kennedy et
al. 2010:23
Terrell et al. 2005:833/ Kennedy et
al. 2010:24
Terrell et al. 2005:833/ Kennedy et
al. 2010:24
Terrell et al. 2005:832/ Kennedy et
al. 2010:24
Terrell et al. 2005:833/ Kennedy et
al. 2010:24
Rubiaceae
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Current disposition
Group
Family
Hedyotis foggiana Fosberg
Kadua foggiana (Fosberg) W.L.Wagner & Lorence
Dicot
Rubiaceae
Hedyotis foliosa (Hillebr.) Fosberg
Kadua foliosa Hillebr.
Dicot
Rubiaceae
Hedyotis formosa (Hillebr.) Fosberg
Kadua formosa Hillebr.
Kadua fosbergii (W.L.Wagner & D.R.Herbst)
W.L.Wagner & Lorence
Kadua cordata Cham. & Schltdl. subsp. waimeae
(Wawra) W.L.Wagner & Lorence
Dicot
Rubiaceae
Dicot
Rubiaceae
Dicot
Rubiaceae
Kadua axillaris (Wawra) W.L.Wagner & Lorence
Dicot
Rubiaceae
Hedyotis knudsenii (Hillebr.) Fosberg
Kadua knudsenii Hillebr.
Dicot
Rubiaceae
Hedyotis littoralis (Hillebr.) Fosberg
Kadua littoralis Hillebr.
Dicot
Rubiaceae
Hedyotis mannii Fosberg
Kadua laxiflora H.Mann
Dicot
Rubiaceae
Hedyotis parvula (A.Gray) Fosberg
Hedyotis schlechtendahliana Steud. subsp. remyi
(Hillebr.) Fosberg
Hedyotis schlechtendahliana Steud. subsp.
schlechtendahliana
Kadua parvula A.Gray
Kadua cordata Cham. & Schltdl. subsp. remyi (Hillebr.)
W.L.Wagner & Lorence
Dicot
Rubiaceae
Dicot
Rubiaceae
Kadua cordata Cham. & Schltdl. subsp. cordata
Dicot
Rubiaceae
Hedyotis schlechtendahliana Steud. subsp. waimeae
(Wawra) W.L.Wagner & D.H.Lorence
Hedyotis schlechtendahliana Steud. var. remyi (Hillebr.)
Fosberg
Kadua cordata Cham. & Schltdl. subsp. waimeae
(Wawra) W.L.Wagner & Lorence
Kadua cordata Cham. & Schltdl. subsp. remyi (Hillebr.)
W.L.Wagner & Lorence
Kadua st.-johnii (B.C.Stone & Lane) W.L.Wagner &
Lorence
Dicot
Rubiaceae
Dicot
Rubiaceae
Dicot
Rubiaceae
Hedyotis fosbergii W.L.Wagner & Herbst
Hedyotis glaucifolia (A.Gray) Fosberg var. waimeae
(Wawra) Fosberg
Hedyotis hillebrandii (Fosberg) W.L.Wagner &
D.R.Herbst
Hedyotis st.-johnii B.C.Stone & Lane
Hedyotis terminalis (Hook. & Arn.) W.L.Wagner &
D.R.Herbst
Hedyotis tryblium D.R.Herbst & W.L.Wagner
Hibiscus brackenridgei A.Gray var. molokaiana Rock ex
Caum
Hibiscus calyphyllus Cav.
Hippeastrum puniceum sensu Wagner et al. (1990), non
(Lam.) Voss
Hordeum leporinum Link
Huperzia erubescens (Brack.) Holub x H. serrata
(Thunb. ex Murray) Trevis.
Literature source
Terrell et al. 2005:832/ Kennedy et
al. 2010:24
Terrell et al. 2005:833/ Kennedy et
al. 2010:24
Terrell et al. 2005:832/ Kennedy et
al. 2010:24
Terrell et al. 2005:832/ Kennedy et
al. 2010:24
Terrell et al. 2005:833/ Kennedy et
al. 2010:23
Terrell et al. 2005:832/ Kennedy et
al. 2010:23
Terrell et al. 2005:832/ Kennedy et
al. 2010:24
Terrell et al. 2005:833/ Kennedy et
al. 2010:24
Terrell et al. 2005:833/ Kennedy et
al. 2010:24
Terrell et al. 2005:833/ Kennedy et
al. 2010:24
Terrell et al. 2005:833/ Kennedy et
al. 2010:23
Terrell et al. 2005:833/ Kennedy et
al. 2010:23
Terrell et al. 2005:833/ Kennedy et
al. 2010:23
Terrell et al. 2005:833/ Kennedy et
al. 2010:23
Terrell et al. 2005:833/ Kennedy et
al. 2010:24
Terrell et al. 2005:832/ Kennedy et
al. 2010:23
Terrell et al. 2005:832/ Kennedy et
al. 2010:24
Kadua affinis DC.
Kadua tryblium (D.R.Herbst & W.L.Wagner)
W.L.Wagner & Lorence
Dicot
Rubiaceae
Dicot
Rubiaceae
Hibiscus brackenridgei A.Gray subsp. molokaiana
(Rock ex Caum) F.D.Wilson
Hibiscus ovalifolius (Forssk.) Vahl
Dicot
Dicot
Malvaceae
Malvaceae
Hippeastrum striatum (Lam.) H.E.Moore
Hordeum murinum L. subsp. leporinum (Link) Arcang.
Monocot
Monocot
Liliaceae
Poaceae
Wilson 1993:278/Herbst & Wagner
1999:23
Staples et al. 2003:14
Herbst & Wagner 1999:23/Staples
& Herbst 2005:689
Herbst & Clayton 1998:28
Huperzia xsulcinervia (Spring) Trevis.
Pteridophyte
Lycopodiaceae
Palmer 2003:263
D-12
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Huperzia erubescens (Brack.) Holub x H. subintegra
(Hillebr.) Beitel & W.H.Wagner
Huperzia haleakalae (Brack.) Holub x H. subintegra
(Hillebr.) Beitel & W.H.Wagner
Huperzia nutans (Brack.) Rothm. x H. phyllantha
(Hook. & Arn.) Holub
Current disposition
Group
Family
Literature source
Huperzia xcarlquistii Beitel & W.H.Wagner
Pteridophyte
Lycopodiaceae
Palmer 2003:265
Huperzia xmedeirosii Beitel & W.H.Wagner
Pteridophyte
Lycopodiaceae
Palmer 2003:265
Huperzia xkoolauensis (W.H.Wagner) Kartesz
Pteridophyte
Lycopodiaceae
Palmer 2003:259
Huperzia serrata (Thunb. ex Murray) Trevis. x H.
subintegra (Hillebr.) Beitel & W.H.Wagner
Huperzia xgillettii Beitel & W.H.Wagner
Pteridophyte
Lycopodiaceae
Hypericum mutilum L.
Hypericum mutilum L. subsp. mutilum
Dicot
Clusiaceae
Impatiens oliveri C.Wright ex W.Wats.
Impatiens wallerana Hook.f (spelling change)
Indigofera hendecaphylla sensu Du Puy et al. (1993),
non Jacq.
Ischaemum indicum sensu Wagner et al. (1990), non
(Houtt.) Merr.
Impatiens sodeni Engl. & Warb. ex Engl.
Impatiens walleriana Hook.f
Dicot
Dicot
Balsaminaceae
Balsaminaceae
Indigofera spicata Forssk.
Dicot
Fabaceae
Ischaemum ciliare Retz.
Monocot
Poaceae
Palmer 2003:265
Robson 1990:116/Wagner &
Herbst 1995:18
Grey-Wilson 1982:20/Herbst &
Wagner 1999:15
Staples & Herbst 2005:172
Du Puy et al. 1993: 731/ Herbst &
Wagner 1999:21
Sun & Phillips 2008:611/ Snow &
Lau 2010:52
Jacquemontia sandwicensis A.Gray
Koeleria macrantha (Ledeb.) Schult.
Dicot
Monocot
Convolvulaceae
Poaceae
Namoff et al. 2010
Herbst & Clayton 1998:29
Korthalsella complanata (Tiegh.) Engl.
Dicot
Viscaceae
Herbst & Wagner 1999:32
Korthalsella platycaula (Tiegh.) Engl.
Dicot
Viscaceae
Herbst & Wagner 1999:32
Korthalsella degeneri Danser
Dicot
Viscaceae
Herbst & Wagner 1999:32
Korthalsella latissima (Tiegh.) Danser
Dicot
Viscaceae
Herbst & Wagner 1999:32
Korthalsella remyana Tiegh.
Lactuca sativa L.
Dicot
Dicot
Viscaceae
Asteraceae
Herbst & Wagner 1999:32
Koopman et al. 2001
Nicolson 1996:341; Nesom
2000:58/Nesom 2001b:514
Nicolson 1996:341; Nesom
2000:58/Nesom 2001b:514
Nicolson 1996:341; Nesom
2000:58/Nesom 2001b:514
Labiak 2011:147
Wagner et al. 1997:58
Harley & Paton 2001:244/ Staples
et al. 2003:14
Jacquemontia ovalifolia (Vahl ex West) Hallier f. subsp.
sandwicensis (A.Gray) K.R.Robertson
Koeleria nitida Nutt.
Korthalsella taenioides (Comm. ex DC.) Engl. f.
taenioides
Korthalsella taenioides (Comm. ex DC.) Engl. f.
taenioides
Korthalsella taenioides (Comm. ex DC.) Engl. fma.
horneana (Tiegh.) Molvray
Korthalsella taenioides (Comm. ex DC.) Engl. fma.
pendula (Wawra) Molvray
Korthalsella taenioides (Comm. ex DC.) Engl. fma.
remyana (Tiegh.) Molvray
Lactuca serriola L.
Lagenifera [Lagenophora] erici C.N.Forbes
Lagenifera [Lagenophora] helenae C.N.Forbes &
Lydgate
Keysseria erici (C.N.Forbes) Cabrera
Dicot
Asteraceae
Keysseria helenae (C.N.Forbes & Lydgate) Cabrera
Dicot
Asteraceae
Lagenifera [Lagenophora] maviensis H.Mann
Lellingeria saffordii (Maxon) A.R.Sm. & R.C.Moran
Lemna perpusilla sensu Wagner et al. (1990), non Torr.
Keysseria maviensis (H.Mann) Cabrera
Stenogrammitis saffordii (Maxon) Labiak
Lemna aequinoctialis Welw.
Dicot
Pteridophyte
Monocot
Asteraceae
Polypodiaceae
Lemnaceae
Leonurus sibiricus sensu Wagner et al. (1990), non L.
Leonurus japonicus Houtt.
Dicot
Lamiaceae
D-13
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Lepidium hyssopifolium sensu Wagner et al. (1990),
non Desv.
Lepidium africanum (Burm.f.) DC.
Dicot
Brassicaceae
Leptochloa dubia (Kunth) Nees
Leptochloa fusca (L.) Kunth subsp. uninervia (J.Presl)
N.Snow
Disakisperma dubia (Kunth) P.M.Peterson & N.Snow
Diplachne fusca subsp. uninervia (J.Presl) P.M.Peterson
& N.Snow
Monocot
Poaceae
Monocot
Poaceae
Diplachne fusca subsp. uninervia (J.Presl) P.M.Peterson
& N.Snow
Monocot
Poaceae
Wagner & Herbst 1995:17
Peterson et al. 2012:1327/ Wagner
et al. 2012:96
Peterson et al. 2012:1327/ Wagner
et al. 2012:96
Imada et al. 2000:14; Peterson et
al. 2012:1327/Wagner et al.
2012:96
Eragrostis curvula (Schrad.) Nees
Monocot
Poaceae
Herbst & Clayton 1998:29
Leptospermum petersonii F.M.Bailey
Leptospermum polygalifolium Salisb. subsp.
polygalifolium
Dicot
Myrtaceae
Herbarium Pacificum Staff 1999:5
Dicot
Myrtaceae
Herbarium Pacificum Staff 1999:6
Leptospermum morrisonii Joy Thomps.
Kummerowia striata (Thunb.) Schindl.
Dicot
Dicot
Myrtaceae
Fabaceae
Herbarium Pacificum Staff 1999:5
Staples et al. 2002:10
xLindsaeosoria flynnii W.H.Wagner
Melanthera bryanii (Sherff) W.L.Wagner & H.Rob.
Lipochaeta connata (Gaudich.) DC. subsp. acris (Sherff)
W.L.Wagner & H.Rob.
Lipochaeta connata (Gaudich.) DC. subsp. connata
Melanthera fauriei (H.Lév.) W.L.Wagner & H.Rob.
Melanthera integrifolia (Nutt.) W.L.Wagner & H.Rob.
Melanthera kamolensis (O.Deg. & Sherff) W.L.Wagner
& H.Rob.
Melanthera lavarum (Gaudich.) W.L.Wagner & H.Rob.
Lipochaeta lobata (Gaudich.) DC. subsp. leptophylla
(O.Deg. & Sherff) W.L.Wagner & H.Rob.
Lipochaeta lobata (Gaudich.) DC. subsp. lobata
Pteridophyte
Dicot
Lindsaeaceae
Asteraceae
Palmer 2003:174
Wagner & Robinson 2002:552
Dicot
Dicot
Dicot
Dicot
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Wagner & Robinson 2002:545
Wagner & Robinson 2002:545
Wagner & Robinson 2002:552
Wagner & Robinson 2002:552
Dicot
Dicot
Asteraceae
Asteraceae
Wagner & Robinson 2002:552
Wagner & Robinson 2002:553
Dicot
Dicot
Asteraceae
Asteraceae
Wagner & Robinson 2002:547
Wagner & Robinson 2002:546
Dicot
Asteraceae
Wagner & Robinson 2002:554
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Wagner & Robinson 2002:554
Wagner & Robinson 2002:554
Wagner & Robinson 2002:554
Wagner & Robinson 2002:554
Wagner & Robinson 2002:554
Wagner & Robinson 2002:554
Leptochloa uninervia (J.Presl) A.S.Hitchc. & Chase
Leptochloa uninervia sensu Warren & Herbst (1994), in
part, non (J.Presl) A.S.Hitchc. & Chase
Leptospermum flavescens sensu Wagner et al. (1990),
in part, non Sm.
Leptospermum flavescens Sm.
Leptospermum laevigatum sensu Wagner et al. (1990),
in part, non (Gaertn.) F.Muell.
Lespedeza striata (Thunb.) Hook. & Arn.
Lindsaea ensifolia Sw. x Sphenomeris chinensis (L.)
Maxon
Lipochaeta bryanii Sherff
Lipochaeta connata (Gaudich.) DC. var. acris (Sherff)
R.C.Gardner
Lipochaeta connata (Gaudich.) DC. var. connata
Lipochaeta fauriei H.Lév.
Lipochaeta integrifolia (Nutt.) Gray
Lipochaeta kamolensis O.Deg. & Sherff
Lipochaeta lavarum (Gaudich.) DC.
Lipochaeta lobata (Gaudich.) DC. var. leptophylla
O.Deg. & Sherff
Lipochaeta lobata (Gaudich.) DC. var. lobata
Lipochaeta micrantha (Nutt.) A.Gray var. exigua
(O.Deg. & Sherff) R.C.Gardner
Lipochaeta micrantha (Nutt.) A.Gray var. micrantha
Lipochaeta perdita Sherff
Lipochaeta populifolia (Sherff) R.C.Gardner
Lipochaeta remyi A.Gray
Lipochaeta subcordata A.Gray
Lipochaeta tenuifolia A.Gray
Current disposition
Melanthera micrantha (Nutt.) W.L.Wagner & H.Rob.
subsp. exigua (O.Deg. & Sherff) W.L.Wagner & H.Rob.
Melanthera micrantha (Nutt.) W.L.Wagner & H.Rob.
subsp. micrantha
Melanthera perdita (Sherff) W.L.Wagner & H.Rob.
Melanthera populifolia (Sherff) W.L.Wagner & H.Rob.
Melanthera remyi (A.Gray) W.L.Wagner & H.Rob.
Melanthera subcordata (A.Gray) W.L.Wagner & H.Rob.
Melanthera tenuifolia (A.Gray) W.L.Wagner & H.Rob.
D-14
Group
Family
Literature source
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Current disposition
Melanthera tenuis (O.Deg. & Sherff) W.L.Wagner &
Lipochaeta tenuis O.Deg. & Sherff
H.Rob.
Lipochaeta venosa Sherff
Melanthera venosa (Sherff) W.L.Wagner & H.Rob.
Melanthera waimeaensis (H.St.John) W.L.Wagner &
Lipochaeta waimeaensis H.St.John
H.Rob.
Lobelia dunbarii Rock subsp. dunbarii (spelling change) Lobelia dunbariae Rock subsp. dunbariae
Lobelia dunbarii Rock subsp. paniculata (Rock)
Lobelia dunbariae Rock subsp. paniculata (Rock)
Lammers (spelling change)
Lammers
Lobelia gaudichaudii A.DC. subsp. gaudichaudii
Lobelia gaudichaudii A.DC.
Lobelia gaudichaudii A.DC. subsp. koolauensis (Hosaka
& Fosberg) Lammers
Lobelia koolauensis (Hosaka & Fosberg) Lammers
Lobelia gloria-montis Rock
Lobelia gloria-montis Rock subsp. gloria-montis
Lobelia gloria-montis Rock var. longibracteata (Rock)
Lobelia gloria-montis Rock subsp. longibracteata
Rock
(Rock) Lammers
Lobelia kauaensis (A.Gray) A.Heller (spelling change)
Lobelia xkauaiensis (A.Gray) A.Heller
Lolium tementulum L. (spelling change)
Lolium temulentum L.
Lycopersicon esculentum sensu Wagner et al. (1990),
Solanum lycopersicum L. var. cerasiforme (Dunal)
non Mill.
D.M.Spooner, G.J.Anderson & R.K.Jansen
Lycopersicon pimpinellifolium sensu Wagner et al.
Solanum lycopersicum L. var. cerasiforme (Dunal)
(1990), non (Jusl.) Mill.
D.M.Spooner, G.J.Anderson & R.K.Jansen
Lysimachia caliginis H.St.John
Lysimachia remyi Hillebr.
Lysimachia haupuensis H.St.John
Lysimachia hillebrandii Hook.f. ex A.Gray
Lysimachia kahiliensis H.St.John
Lysimachia daphnoides (A.Gray) Hillebr.
Lysimachia ovoidea H.St.John
Lysimachia hillebrandii Hook.f. ex A.Gray
Lysimachia remyi Hillebr. subsp. caliginis (H.St.John)
K.L.Marr
Lysimachia remyi Hillebr.
Lysimachia remyi Hillebr. subsp. subherbacea
(H.St.John) K.L.Marr
Lysimachia remyi Hillebr.
Lysimachia subherbacea (Hillebr.) H.St.John
Lysimachia remyi Hillebr.
Lysimachia waianaeensis H.St.John
Lysimachia hillebrandii Hook.f. ex A.Gray
Group
Family
Dicot
Dicot
Asteraceae
Asteraceae
Wagner & Robinson 2002:555
Wagner & Robinson 2002:555
Dicot
Dicot
Asteraceae
Campanulaceae
Wagner & Robinson 2002:555
Wagner et al. 2005– website
Dicot
Dicot
Campanulaceae
Campanulaceae
Wagner et al. 2005– website
Lammers 2007:804
Dicot
Dicot
Campanulaceae
Campanulaceae
Lammers 2007:803
Lammers 2007:805
Dicot
Dicot
Monocot
Campanulaceae
Campanulaceae
Poaceae
Lammers 2007:806
Lammers 2007:801
Clayton & Snow 2010:26
Dicot
Solanaceae
Wagner et al. 1997:62
Dicot
Dicot
Dicot
Dicot
Dicot
Solanaceae
Primulaceae
Primulaceae
Primulaceae
Primulaceae
Wagner et al. 1997:62
Herbst & Wagner 1999:31
Herbst & Wagner 1999:30
Herbst & Wagner 1999:30
Herbst & Wagner 1999:30
Dicot
Primulaceae
Herbst & Wagner 1999:31
Dicot
Dicot
Dicot
Primulaceae
Primulaceae
Primulaceae
Herbst & Wagner 1999:31
Herbst & Wagner 1999:31
Herbst & Wagner 1999:30
Tucker 1994:87/Wagner & Herbst
1999:1899
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Mariscus congestus (Vahl) C.B.Clarke
Cyperus congestus Vahl
Monocot
Cyperaceae
Mariscus cyperinus (Retz.) Vahl
Cyperus cyperinus (Retz.) Suringar
Monocot
Cyperaceae
Mariscus fauriei (Kük.) T.Koyama
Mariscus hillebrandii (Boeck.) T.Koyama subsp.
decipiens (Hillebr.) T.Koyama
Mariscus hillebrandii (Boeck.) T.Koyama subsp.
hillebrandii
Cyperus fauriei Kük.
Cyperus hillebrandii Boeck. var. decipiens (Hillebr.)
Kük.
Monocot
Cyperaceae
Monocot
Cyperaceae
Cyperus hillebrandii Boeck. var. hillebrandii
Monocot
Cyperaceae
D-15
Literature source
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Current disposition
Mariscus hypochlorus (Hillebr.) C.B.Clarke
Mariscus hypochlorus (Hillebr.) C.B.Clarke subsp.
brevior (Kük.) T.Koyama
Cyperus hypochlorus Hillebr. var. hypochlorus
Monocot
Cyperaceae
Cyperus hypochlorus Hillebr. var. brevior Kük.
Monocot
Cyperaceae
Mariscus javanicus (Houtt.) Merr. & Metcalfe
Cyperus javanicus Houtt.
Monocot
Cyperaceae
Mariscus kunthianus Gaudich.
Cyperus neokunthianus Kük.
Monocot
Cyperaceae
Mariscus meyenianus (Kunth) Nees
Mariscus pennatiformis (Kük.) T.Koyama subsp.
bryanii (Kük.) T.Koyama
Mariscus pennatiformis (Kük.) T.Koyama subsp.
pennatiformis
Mariscus phleoides Nees ex Kunth subsp. hawaiensis
(H.Mann) T.Koyama
Cyperus meyenianus Kunth
Monocot
Cyperaceae
Cyperus pennatiformis Kük. var. bryanii Kük.
Monocot
Cyperaceae
Monocot
Cyperaceae
Monocot
Cyperaceae
Mariscus phleoides Nees ex Kunth subsp. phleoides
Cyperus pennatiformis Kük. var. pennatiformis
Cyperus phleoides (Nees ex Kunth) Hillebr. var.
hawaiiensis (H.Mann) Kük.
Cyperus phleoides (Nees ex Kunth) Hillebr. var.
phleoides
Monocot
Cyperaceae
Mariscus rockii (Kük.) T.Koyama
Cyperus rockii Kük.
Monocot
Cyperaceae
Mariscus sandwicensis (Kük.) T.Koyama
Mazus rugosus Lour.
Cyperus sandwicensis Kük.
Mazus pumilus (Burm.f.) Steenis
Monocot
Dicot
Cyperaceae
Scrophulariaceae
Mecodium recurvum (Gaudich.) Copel.
Melastoma candidum D.Don
Melicope honoluluensis (H.St.John) T.G.Hartley &
B.C.Stone
Melicope mauii T.G.Hartley & B.C.Stone
Mentha xrotundifolia sensu Wagner et al. (1990), non
(L.) Huds.
Mentha spicata L.
Mentha xvillosa sensu Wagner et al. (1990), non Huds.
Microlepia speluncae (L.) T.Moore x M. strigosa
(Thunb.) C.Presl
Hymenophyllum recurvum Gaudich.
Melastoma septemnervium Lour.
Melicope hosakae (H.St.John) W.L.Wagner &
R.K.Shannon
Melicope sessilis (H.Lév.) T.G.Hartley & B.C.Stone
Pteridophyte
Dicot
Hymenophyllaceae
Melastomataceae
Literature source
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner, Herbst et
al. 1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1900
Tucker 1994:87/Wagner & Herbst
1999:1901
Tucker 1994:87/Wagner & Herbst
1999:1901
Oppenheimer 2004:17
Ebihara et al. 2006:231/Snow et al.
2011:13
Staples et al. 2003:15
Dicot
Dicot
Rutaceae
Rutaceae
Wagner & Shannon 1998:16
Hartley & Stone 1989:121
Mentha suaveolens Ehrh.
Mentha xspicata L.
Mentha suaveolens Ehrh.
Dicot
Dicot
Dicot
Lamiaceae
Lamiaceae
Lamiaceae
Staples et al. 2003:14
Staples & Herbst 2005:358
Staples et al. 2003:14
Microlepia xadulterina W.H.Wagner
Pteridophyte
Dennstaedtiaceae
Morinda trimera Hillebr.
Gynochthodes trimera (Hillebr.) Razafim. & B.Bremer
Dicot
Rubiaceae
Dicot
Fabaceae
Dicot
Fabaceae
Palmer 2003:186
Razafimandimbison & Bremer
2011:295/Wagner et al. 2012:65
Wilmot-Dear 1990:5/Wagner &
Herbst 1995:21
Wilmot-Dear 1990:25/Wagner &
Herbst 1995:20
Dicot
Polygonaceae
Staples & Herbst 2005:464
Mucuna gigantea (Willd.) DC.
Mucuna gigantea (Willd.) DC. subsp. gigantea
Mucuna urens sensu Wagner et al. (1990), non (L.)
Medik.
Mucuna sloanei Fawc. & Rendle var. sloanei
Muehlenbeckia axillaris sensu Wagner et al. (1990), non
(Hook.f.) Walp.
Muehlenbeckia complexa (A.Cunn.) Meisn.
D-16
Group
Family
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Munroidendron racemosum (C.N.Forbes) Sherff
Current disposition
Polyscias racemosa (C.N.Forbes) Lowry &
G.M.Plunkett
Dicot
Araliaceae
Myrica cerifera L.
Morella cerifera (L.) Small
Dicot
Myricaceae
Myrica faya Aiton
Myrsine emarginata (Rock) Hosaka
Nasturtium sarmentosum (G.Forst. ex DC.) Schinz &
Guillaumin
Morella faya (Aiton) Wilbur
Myrsine lessertiana A.DC.
Dicot
Dicot
Myricaceae
Myrsinaceae
Lowry & Plunkett 2010:74
Wilbur 1994:100/ Oppenheimer
2003:15
Wilbur 1994:103/Staples et al.
2002:13
Wagner & Herbst 2003:35
Rorippa sarmentosa (G.Forst. ex DC.) J.F.Macbr.
Dicot
Brassicaceae
Wagner & Herbst 1995:18
Lobelia niihauensis H.St.John
Dicot
Campanulaceae
Nephrolepis exaltata (L.) Schott
Pteridophyte
Nephrolepidaceae
Nephrolepis brownii (Desv.) Hovenkamp & Miyam.
Sideroxylon polynesicum (Hillebr.) Smedmark &
Anderb.
Pteridophyte
Nephrolepidaceae
Dicot
Sapotaceae
Dryopteris rubiginosa (Brack.) Kuntze
Pteridophyte
Dryopteridaceae
Ochna kirkii sensu Wagner et al. (1990), non Oliv.
Odontonema strictum sensu Wagner et al. (1990), non
(Nees) Kuntze
Ochna thomasiana Engl. & Gilg
Dicot
Ochnaceae
Wagner & Herbst 2003:16
Hovenkamp & Miyamoto
2005:300
Hovenkamp & Miyamoto
2005:293
Smedmark & Anderberg
2007:1502/Wagner et al. 2012:69
Smith et al. 2006:717/Snow et al.
2011:13
Herbarium Pacificum Staff
1998:12
Odontonema cuspidatum (Nees) Kuntze
Dicot
Acanthaceae
Olea europaea L. subsp. africana (P.Mill.) P.Green
Omalanthus populifolius Graham (spelling change)
Ophioderma pendulum (L.) C.Presl subsp. falcatum
(C.Presl) R.T.Clausen
Olea europaea L. subsp. cuspidata (Wall. ex G.Don) Cif.
Homalanthus populifolius Graham
Dicot
Dicot
Oleaceae
Euphorbiaceae
Ophioglossum pendulum L.
Pteridophyte
Ophioglossaceae
Ophioderma pendulum (L.) C.Presl subsp. pendulum
Oplismenus hirtellus (L.) P.Beauv.
Opuntia vulgaris sensu Wagner et al. (1990), non Mill.
Oryzopsis miliacea (L.) Benth. & Hook.f. ex Aschers. &
Schweinf.
Ophioglossum pendulum L.
Oplismenus hirtellus (L.) P.Beauv. subsp. hirtellus
Opuntia monacantha (Willd.) Haw.
Pteridophyte
Monocot
Dicot
Ophioglossaceae
Poaceae
Cactaceae
Piptatherum miliaceum (L.) Coss.
Monocot
Poaceae
Oxalis corymbosa DC.
Paederia scandens (Lour.) Merr.
Oxalis debilis Kunth var. corymbosa (DC.) Lourteig
Paederia foetida L.
Dicot
Dicot
Oxalidaceae
Rubiaceae
Panicum cynodon Reichardt
Dichanthelium cynodon (Reichardt) C.A.Clark & Gould
Dichanthelium hillebrandianum (Hitchc.) C.A.Clark &
Gould
Monocot
Poaceae
Monocot
Poaceae
Neowimmeria niihauensis (H.St.John) O.Deg. & I.Deg.
var. forbesii (H.St.John) O.Deg. & I.Deg.
Nephrolepis exaltata (L.) Schott subsp. hawaiiensis
W.H.Wagner
Nephrolepis multiflora (Roxb.) F.M.Jarrett ex
C.V.Morton
Nesoluma polynesicum (Hillebr.) Baill.
Nothoperanema rubiginosa (Brack.) A.R.Sm. &
D.D.Palmer
Panicum hillebrandianum A.S. Hitchc.
D-17
Group
Family
Literature source
Lorence et al. 1995:20
Green & Wickens 1989:295/
Herbst & Wagner 1999:24
The Plant List website 2012
Smith et al. 2006:709/Wagner et al.
2012:105
Smith et al. 2006:709/Wagner et al.
2012:105
Snow & Lau 2010:52
Staples et al. 2002:6
Snow 2008:40
Herbst & Wagner 1999:25; Staples
& Herbst 2005:447
Wagner & Herbst 1995:25
Clayton & Snow 2010:55; Wagner
et al. 2012:92
Clayton & Snow 2010:55; Wagner
et al. 2012:92
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Monocot
Poaceae
Panicum koolauense H.St.John & Hosaka
Current disposition
Dichanthelium isachnoides (Munro ex Hillebr.)
C.A.Clark & Gould
Dichanthelium koolauense (H.St.John & Hosaka)
C.A.Clark & Gould
Monocot
Poaceae
Panicum maximum Jacq.
Urochloa maxima (Jacq.) R.D.Webster
Monocot
Poaceae
Paraserianthes falcataria (L.) I. Nielsen
Paspalum aff. laxum sensu Snow & Lau (2010), non
Lam.
Paspalum aff. thunbergii sensu Snow & Lau (2010), non
Kunth ex Steud.
Paspalum paniculatum sensu Oppenheimer (2007), non
L.
Passiflora xcaerulea sensu Wagner et al. (1990), non L.
Passiflora mollissima sensu Wagner et al. (1990), non
(Kunth) Bailey
Falcataria moluccana (Miq.) Barneby & J.W.Grimes
Dicot
Fabaceae
Literature source
Clayton & Snow 2010:55; Wagner
et al. 2012:92
Clayton & Snow 2010:55; Wagner
et al. 2012:92
Zuloaga et al. 2003:632/ Wagner et
al. 2012:97
Herbarium Pacificum Staff
1998:10/Staples & Herbst
2005:331
Paspalum arundinaceum Poir.
Monocot
Poaceae
Snow & Davidse 2011:18
Paspalum mandiocanum Trin. var. mandiocanum
Monocot
Poaceae
Paspalum arundinaceum Poir.
Passiflora xviolacea Loisel.
Monocot
Dicot
Poaceae
Passifloraceae
Passiflora tarminiana Coppens & Barney
Melicope adscendens (H.St.John & E.P.Hume)
T.G.Hartley & B.C.Stone
Melicope anisata (H.Mann) T.G.Hartley & B.C.Stone
Melicope balloui (Rock) T.G.Hartley & B.C.Stone
Melicope barbigera A.Gray
Melicope christophersenii (H.St.John) T.G.Hartley &
B.C.Stone
Melicope cinerea A.Gray
Melicope clusiifolia (A.Gray) T.G.Hartley & B.C.Stone
Melicope cruciata (A.Heller) T.G.Hartley & B.C.Stone
Melicope degeneri (B.C.Stone) T.G.Hartley &
B.C.Stone
Melicope elliptica A.Gray
Melicope feddei (H.Lév.) T.G.Hartley & B.C.Stone
Melicope haleakalae (B.C.Stone) T.G.Hartley &
B.C.Stone
Melicope haupuensis (H.St.John) T.G.Hartley &
B.C.Stone
Melicope hawaiensis (Wawra) T.G.Hartley & B.C.Stone
Melicope hiiakae (B.C.Stone) T.G.Hartley & B.C.Stone
Melicope hosakae (H.St.John) W.L.Wagner &
R.K.Shannon
Dicot
Passifloraceae
Snow & Davidse 2011:18
Snow & Lau 2010:53/ Snow &
Davidse 2011:18
Starr et al. 1999:13
Coppens d'Eeckenbrugge et. al.
2001/Staples et al. 2003:17
Dicot
Dicot
Dicot
Dicot
Rutaceae
Rutaceae
Rutaceae
Rutaceae
Hartley & Stone 1989:120
Hartley & Stone 1989:122
Hartley & Stone 1989:121
Hartley & Stone 1989:122
Dicot
Dicot
Dicot
Dicot
Rutaceae
Rutaceae
Rutaceae
Rutaceae
Hartley & Stone 1989:121
Hartley & Stone 1989:122
Hartley & Stone 1989:122
Hartley & Stone 1989:121
Dicot
Dicot
Dicot
Rutaceae
Rutaceae
Rutaceae
Hartley & Stone 1989:122
Hartley & Stone 1989:122
Hartley & Stone 1989:121
Dicot
Rutaceae
Hartley & Stone 1989:122
Dicot
Dicot
Dicot
Rutaceae
Rutaceae
Rutaceae
Hartley & Stone 1989:120
Hartley & Stone 1989:121
Wagner & Herbst 1999:1894
Dicot
Rutaceae
Wagner & Shannon 1998:16
Panicum isachnoides Munro ex Hillebr.
Pelea adscendens H.St.John & E.P.Hume
Pelea anisata H.Mann
Pelea balloui Rock
Pelea barbigera (A.Gray) Hillebr.
Pelea christophersenii H.St.John
Pelea cinerea (A.Gray) Hillebr.
Pelea clusiifolia A.Gray
Pelea cruciata A.Heller
Pelea degeneri B.C.Stone
Pelea elliptica (A.Gray) Hillebr.
Pelea feddei H.Lév.
Pelea haleakalae B.C.Stone
Pelea haupuensis H.St.John
Pelea hawaiensis Wawra
Pelea hiiakae B.C.Stone
Pelea honoluluensis H.St.John
D-18
Group
Family
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Pelea hosakae H.St.John
Pelea kaalaensis H.St.John
Pelea kavaiensis H.Mann
Pelea knudsenii Hillebr.
Pelea lydgatei Hillebr.
Pelea macropus Hillebr.
Pelea makahae B.C.Stone
Pelea molokaiensis Hillebr.
Pelea mucronulata H.St.John
Pelea munroi H.St.John
Pelea nealae B.C.Stone
Pelea oahuensis H.Lév.
Pelea obovata H.St.John
Pelea orbicularis Hillebr.
Pelea ovalis H.St.John
Pelea ovata H.St.John & E.P.Hume
Pelea pallida Hillebr.
Pelea paniculata H.St.John
Pelea parvifolia Hillebr.
Pelea peduncularis H.Lév.
Pelea pseudoanisata Rock
Pelea puberula H.St.John
Pelea quadrangularis H.St.John & E.P.Hume
Pelea radiata H.St.John
Pelea reflexa H.St.John
Pelea rotundifolia A.Gray
Current disposition
Melicope hosakae (H.St.John) W.L.Wagner &
R.K.Shannon
Melicope kaalaensis (H.St.John) T.G.Hartley &
B.C.Stone
Melicope kavaiensis (H.Mann) T.G.Hartley &
B.C.Stone
Melicope knudsenii (Hillebr.) T.G.Hartley & B.C.Stone
Melicope lydgatei (Hillebr.) T.G.Hartley & B.C.Stone
Melicope macropus (Hillebr.) T.G.Hartley & B.C.Stone
Melicope makahae (B.C.Stone) T.G.Hartley &
B.C.Stone
Melicope molokaiensis (Hillebr.) T.G.Hartley &
B.C.Stone
Melicope mucronulata (H.St.John) T.G.Hartley &
B.C.Stone
Melicope munroi (H.St.John) T.G.Hartley & B.C.Stone
Melicope nealae (B.C.Stone) T.G.Hartley & B.C.Stone
Melicope oahuensis (H.Lév.) T.G.Hartley & B.C.Stone
Melicope obovata (H.St.John) T.G.Hartley & B.C.Stone
Melicope orbicularis (Hillebr.) T.G.Hartley &
B.C.Stone
Melicope ovalis (H.St.John) T.G.Hartley & B.C.Stone
Melicope ovata (H.St.John & E.P.Hume) T.G.Hartley &
B.C.Stone
Melicope pallida (Hillebr.) T.G.Hartley & B.C.Stone
Melicope paniculata (H.St.John) T.G.Hartley &
B.C.Stone
Melicope sessilis (H.Lév.) T.G.Hartley & B.C.Stone
Melicope peduncularis (H.Lév.) T.G.Hartley &
B.C.Stone
Melicope pseudoanisata (Rock) T.G.Hartley &
B.C.Stone
Melicope puberula (H.St.John) T.G.Hartley &
B.C.Stone
Melicope quadrangularis (H.St.John & E.P.Hume)
T.G.Hartley & B.C.Stone
Melicope radiata (H.St.John) T.G.Hartley & B.C.Stone
Melicope reflexa (H.St.John) T.G.Hartley & B.C.Stone
Melicope rotundifolia (A.Gray) T.G.Hartley &
B.C.Stone
D-19
Group
Family
Literature source
Dicot
Rutaceae
Wagner & Shannon 1998:16
Dicot
Rutaceae
Hartley & Stone 1989:121
Dicot
Dicot
Dicot
Dicot
Rutaceae
Rutaceae
Rutaceae
Rutaceae
Hartley & Stone 1989:121
Hartley & Stone 1989:121
Hartley & Stone 1989:121
Hartley & Stone 1989:121
Dicot
Rutaceae
Hartley & Stone 1989:121
Dicot
Rutaceae
Hartley & Stone 1989:121
Dicot
Dicot
Dicot
Dicot
Dicot
Rutaceae
Rutaceae
Rutaceae
Rutaceae
Rutaceae
Hartley & Stone 1989:121
Hartley & Stone 1989:121
Hartley & Stone 1989:121
Hartley & Stone 1989:122
Hartley & Stone 1989:121
Dicot
Dicot
Rutaceae
Rutaceae
Hartley & Stone 1989:121
Hartley & Stone 1989:122
Dicot
Dicot
Rutaceae
Rutaceae
Hartley & Stone 1989:121
Hartley & Stone 1989:121
Dicot
Dicot
Rutaceae
Rutaceae
Hartley & Stone 1989:122
Hartley & Stone 1989:121
Dicot
Rutaceae
Hartley & Stone 1989:122
Dicot
Rutaceae
Hartley & Stone 1989:121
Dicot
Rutaceae
Hartley & Stone 1989:121
Dicot
Dicot
Dicot
Rutaceae
Rutaceae
Rutaceae
Hartley & Stone 1989:122
Hartley & Stone 1989:121
Hartley & Stone 1989:122
Dicot
Rutaceae
Hartley & Stone 1989:122
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Pelea zahlbruckneri Rock
Current disposition
Melicope saint-johnii (E.P.Hume) T.G.Hartley &
B.C.Stone
Melicope sandwicensis (Hook. & Arn.) T.G.Hartley &
B.C.Stone
Melicope sessilis (H.Lév.) T.G.Hartley & B.C.Stone
Melicope volcanica (A.Gray) T.G.Hartley & B.C.Stone
Melicope waialealae (Wawra) T.G.Hartley & B.C.Stone
Melicope wailauensis (H.St.John) T.G.Hartley &
B.C.Stone
Melicope wawraeana (Rock) T.G.Hartley & B.C.Stone
Melicope zahlbruckneri (Rock) T.G.Hartley &
B.C.Stone
Pennisetum clandestinum Hochst. ex Chiov.
Cenchrus clandestinus (Hochst. ex Chiov.) Morrone
Monocot
Poaceae
Pennisetum complanatum (Nees) Hemsl.
Pennisetum glaucum (L.) R.Br.
Pennisetum macrostachyum (Brongn.) Trin.
Cenchrus complanatus (Nees) Morrone
Cenchrus americanus (L.) Morrone
Cenchrus caninus (Reinw. ex Blume) Morrone
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Pennisetum polystachion (L.) Schult.
Cenchrus polystachios (L.) Morrone
Monocot
Poaceae
Pennisetum purpureum Schumach.
Cenchrus purpureus (Schumach.) Morrone
Monocot
Poaceae
Pennisetum setaceum (Forssk.) Chiov.
Pennisetum villosum R.Br. ex Fresen.
Cenchrus setaceus (Forssk.) Morrone
Cenchrus longisetus M.C.Johnst.
Monocot
Monocot
Poaceae
Poaceae
Peperomia leptostachya Hook. & Arn., non (Nutt.)
Chapman
Peperomia nahikuensis H.St.John
Phalaris tuberosa L. var. stenoptera sensu Wagner et al.
(1990), non (Hack.) Hitchc.
Phragmites karka sensu Wagner et al. (1990), non
(Retz.) Trin.
Peperomia blanda (Jacq.) Kunth var. floribunda (Miq.)
H.Huber
Peperomia eekana C.DC.
Dicot
Dicot
Piperaceae
Piperaceae
Hartley & Stone 1989:122
Chemisquy et al. 2010:127/
Wagner et al. 2012:98
Chemisquy et al. 2010:127/
Wagner et al. 2012:98
Chemisquy et al. 2010:127
Chemisquy et al. 2010:127
Chemisquy et al. 2010:129/
Wagner et al. 2012:99
Chemisquy et al. 2010:129/
Wagner et al. 2012:99
Chemisquy et al. 2010:129/
Wagner et al. 2012:99
Wagner et al. 2005–
Huber 1987:293/Florence
1997:173/Herbst & Wagner
1999:25
Wagner et al. 2012:58
Phalaris aquatica L.
Monocot
Poaceae
Herbst & Clayton 1998:33
Phragmites australis (Cav.) Trin. ex Steud.
Monocot
Poaceae
Herbst & Clayton 1998:33
Phyllanthus distichus Hook. & Arn.
Dicot
Euphorbiaceae
Herbst & Wagner 1999:20
Phyllanthus distichus Hook. & Arn.
Phyllostegia haliakalae Wawra
Phyllostegia parviflora (Gaudich.) Benth. var. lydgatei
(Sherff) W.L.Wagner
Dicot
Dicot
Euphorbiaceae
Lamiaceae
Herbst & Wagner 1999:20
Wagner 1999b:269
Dicot
Lamiaceae
Wagner 1999a:282
Pelea saint-johnii E.P.Hume
Pelea sandwicensis (Hook. & Arn.) A.Gray
Pelea sessilis H.Lév.
Pelea volcanica A.Gray
Pelea waialealae Wawra
Pelea wailauensis H.St.John
Pelea wawraeana Rock
Phyllanthus distichus Hook. & Arn. var. degeneri
(Sherff) Govaerts & Radcl.-Sm.
Phyllanthus distichus Hook. & Arn. var. ellipticus
(Müll.Arg.) Govaerts & Radcl.-Sm.
Phyllostegia imminuta (Sherff) H.St.John
Phyllostegia mollis Benth. var. lydgatei Sherff
D-20
Group
Family
Literature source
Dicot
Rutaceae
Hartley & Stone 1989:121
Dicot
Dicot
Dicot
Dicot
Rutaceae
Rutaceae
Rutaceae
Rutaceae
Hartley & Stone 1989:121
Hartley & Stone 1989:121
Hartley & Stone 1989:122
Hartley & Stone 1989:122
Dicot
Dicot
Rutaceae
Rutaceae
Hartley & Stone 1989:122
Hartley & Stone 1989:122
Dicot
Rutaceae
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Current disposition
Group
Family
Pleomele konaensis H.St.John
Pleomele hawaiiensis O.Deg. & I.Deg.
Monocot
Agavaceae
Pleomele rockii H.St.John
Pleomele auwahiensis H.St.John
Monocot
Agavaceae
Pleomele stenophylla H.St.John
Pluchea symphytifolia sensu Wagner et al. (1990), non
(Mill.) Gillis
Pneumatopteris hudsoniana (Brack.) Holttum
Pneumatopteris pendens D.D.Palmer
Pneumatopteris sandwicensis (Brack.) Holttum
Pleomele aurea (H.Mann) N.E.Br.
Monocot
Agavaceae
Pluchea carolinensis (Jacq.) G.Don
Cyclosorus hudsonianus (Brack.) Ching
Cyclosorus pendens (D.D.Palmer) N.Snow
Cyclosorus sandwicensis (Brack.) Copel.
Dicot
Pteridophyte
Pteridophyte
Pteridophyte
Asteraceae
Thelypteridaceae
Thelypteridaceae
Thelypteridaceae
Polygonum capitatum Buch.-Ham. ex D. Don
Persicaria capitata (Buch.-Ham. ex D.Don) Masam.
Dicot
Polygonaceae
Polygonum chinense L.
Persicaria chinense (L.) Nakai
Dicot
Polygonaceae
Polygonum convolvulus L.
Fallopia convolvulus (L.) Á.Löve
Dicot
Polygonaceae
Polygonum glabrum Willd.
Persicaria glabra (Willd.) M.Gómez
Dicot
Polygonaceae
Polygonum punctatum Elliott
Persicaria punctata (Elliott) Small
Dicot
Polygonaceae
Potamogeton pectinatus L.
Stuckenia pectinata (L.) Börner
Monocot
Potamogetonaceae
Pouteria sandwicensis (A.Gray) Baehni & O.Deg.
Pritchardia affinis Becc.
Pritchardia aylmer-robinsonii H.St.John
Pritchardia elliptica Rock & Caum
Planchonella sandwicensis (A.Gray) Pierre
Pritchardia maideniana Becc.
Pritchardia remota Becc.
Pritchardia glabrata Becc. & Rock
Dicot
Monocot
Monocot
Monocot
Sapotaceae
Arecaceae
Arecaceae
Arecaceae
Pritchardia kahukuensis Caum
Pritchardia lanaiensis Becc. & Rock
Pritchardia limahuliensis H.St.John
Pritchardia martii (Gaudich.) H.Wendl.
Pritchardia glabrata Becc. & Rock
Pritchardia napaliensis H.St.John
Monocot
Monocot
Monocot
Arecaceae
Arecaceae
Arecaceae
D-21
Literature source
proposed syn. under Dracaena
konaensis (H.St.John) Jankal.,
along with Pleomele konaensis
H.St.John (Jankalski 2008:21)
proposed syn. under Dracaena
rockii (H.St.John) Jankal., along
with Pleomele rockii H.St.John
(Jankalski 2008:21)
proposed resurrection of Pleomele
stenophylla H.St.John (as Dracaena
halemanuensis Jankal. (Jankalski
2008:21)
Wagner & Herbst 1995:16/ Khan
& Jarvis 1989
Snow et al. 2011:15
Snow et al. 2011:15
Snow et al. 2011:15
Ronse Decraene & Akeroyd
1988:367/Herbst & Wagner
1999:29
Ronse Decraene & Akeroyd
1988:367/Herbst & Wagner
1999:29
Ronse Decraene & Akeroyd
1988:369/Herbst & Wagner
1999:29
Wilson 1990:632/Herbst & Wagner
1999:30
Herbst & Wagner 1999:30/ Hinds
& Freeman 2005:586
Haynes et al. 1998:241/Herbst &
Wagner 1999:30
Swenson et al. 2007/Wagner et al.
2012:69
Hodel 2007:S-26
Hodel 2007:S-39
Hodel 2007:S-14
Hodel 2007:S-20; Wagner et al.
2012:77
Hodel 2007:S-14
Hodel 2007:S-37
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Pseuderanthemum fasciculatum sensu Herbarium
Pacificum Staff (1998), non (Oerst.) Leonard
Pteridium aquilinum (L.) Kuhn var. decompositum
(Gaudich.) R.M.Tryon
Current disposition
Pseuderanthemum variabile (R.Br.) Radlk.
Group
Family
Literature source
Dicot
Acanthaceae
Herbst & Wagner 1999:12
Pteridophyte
Dennstaedtiaceae
Thomson 2004:800
Dicot
Fabaceae
Herbst & Wagner 1999:21
Dicot
Fabaceae
Pueraria lobata (Willd.) Ohwi var. lobata
Pueraria lobata (Willd.) Ohwi var. thomsoni (Benth.)
Maesen
Pteridium aquilinum (L.) Kuhn subsp. decompositum
(Gaudich.) Lamoureux ex J.A.Thomson
Pueraria montana (Lour.) Merr. var. lobata (Willd.)
Sanjappa & Pradeep
Pueraria montana (Lour.) Merr. var. chinensis (Ohwi)
Sanjappa & Pradeep
Pycreus polystachyos (Rottb.) P.Beauv. subsp.
holocericeus (Link) T.Koyama
Cyperus polystachyos Rottb.
Monocot
Cyperaceae
Pycreus polystachyos (Rottb.) P.Beauv. subsp.
polystachyos
Cyperus polystachyos Rottb.
Monocot
Cyperaceae
Cyperus sanguinolentus Vahl
Monocot
Cyperaceae
Staples et al. 2002:11
Tucker 1994:72/Herbst & Wagner
1999:20/Wagner & Herbst
1999:1901
Tucker 1994:72/Herbst & Wagner
1999:20/Wagner & Herbst
1999:1901
Tucker 1994:66/Herbst & Wagner
1999:20/Wagner & Herbst
1999:1901
Pyracantha crenatoserrata (Hance) Rehder
Dicot
Rosaceae
Herbarium Pacificum Staff 1999:9
Pyracantha koidzumii Rehder
Atractocarpus fitzalanii (F.Muell.) Puttock
Polyscias sandwicensis (A.Gray) Lowry &
G.M.Plunkett
Melinis repens (Willd.) Zizka
Dicot
Dicot
Rosaceae
Rubiaceae
Herbarium Pacificum Staff 1999:9
Puttock 1999:285
Dicot
Monocot
Araliaceae
Poaceae
Lowry & Plunkett 2010:74
Herbst & Clayton 1998:29
Richardia scabra L.
Dicot
Rubiaceae
Rollandia angustifolia (Hillebr.) Rock
Cyanea koolauensis Lammers, Givnish & Sytsma
Dicot
Campanulaceae
Rollandia crispa Gaudich.
Cyanea crispa (Gaudich.) Lammers, Givnish & Sytsma
Cyanea humboldtiana (Gaudich.) Lammers, Givnish &
Sytsma
Dicot
Campanulaceae
Dicot
Campanulaceae
Dicot
Campanulaceae
Dicot
Campanulaceae
Dicot
Campanulaceae
Dicot
Campanulaceae
Dicot
Campanulaceae
Imada et al. 2008:14
Lammers 1993:439/Herbst &
Wagner 1999:17
Lammers 1993:439/Herbst &
Wagner 1999:17
Lammers 1993:439/Herbst &
Wagner 1999:17
Lammers 1998:32/Herbst &
Wagner 1999:17
Lammers 1993:439/Herbst &
Wagner 1999:17
Lammers 1993:439/Herbst &
Wagner 1999:17
Lammers 1993:439/Herbst &
Wagner 1999:17
Lammers 1993:439/Herbst &
Wagner 1999:17
Pycreus sanguinolentus (Vahl) Nees
Pyracantha angustifolia sensu Wagner et al. (1990), in
part, non (Franch.) C.K.Schneid.
Pyracantha angustifolia sensu Wagner et al. (1990), in
part, non (Franch.) C.K.Schneid.
Randia fitzalanii (F.Muell.) F.Muell. ex Benth.
Reynoldsia sandwicensis A.Gray
Rhynchelytrum repens (Willd.) C.E.Hubb.
Richardia brasiliensis sensu Wagner et al. (1990), in
part, non Gomes
Rollandia humboldtiana Gaudich.
Rollandia lanceolata Gaudich. subsp. calycina (Cham.)
Lammers
Rollandia lanceolata Gaudich. subsp. lanceolata
Rollandia longiflora Wawra
Rollandia parvifolia C.N.Forbes
Rollandia purpurellifolia Rock
Cyanea calycina (Cham.) Lammers
Cyanea lanceolata (Gaudich.) Lammers, Givnish &
Sytsma
Cyanea longiflora (Wawra) Lammers, Givnish &
Sytsma
Cyanea parvifolia (C.N.Forbes) Lammers, Givnish &
Sytsma
Cyanea purpurellifolia (Rock) Lammers, Givnish &
Sytsma
D-22
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Rollandia sessilifolia O.Deg.
Cyanea sessilifolia (O.Deg.) Lammers
Dicot
Campanulaceae
Rollandia st.-johnii Hosaka
Cyanea st.-johnii (Hosaka) Lammers, Givnish & Sytsma
Dicot
Campanulaceae
Rorippa microphylla (Boenn. ex Reichenb.) Hyl. ex
A.& D.Löve
Nasturtium microphyllum Boenn. ex Rchb.
Dicot
Brassicaceae
Ruellia brevifolia (Pohl) C.Ezcurra
Rumex crispus L. subsp. crispus
Dicot
Dicot
Acanthaceae
Polygonaceae
Literature source
Lammers 1998:32/Herbst &
Wagner 1999:18
Lammers 1993:440/Herbst &
Wagner 1999:18
Al-Shehbaz & Price
1998:124/Herbst & Wagner
1999:15
Ezcurra 1993:802/Herbst &
Wagner 1999:13
Mosyakin & Wagner 1998:42
Rytidosperma penicillatum (Labill.) Connor & Edgar
Salsola tragus L.
Santalum freycinetianum Gaudich.
Santalum haleakalae Hillebr. var. lanaiense (Rock)
Harbaugh
Monocot
Dicot
Dicot
Poaceae
Chenopodiaceae
Santalaceae
Darbyshire et al. 2010:670
Wagner et al. 1997:55
Harbaugh et al. 2010:833
Dicot
Santalaceae
Harbaugh et al. 2010:834
Santalum pyrularium A.Gray
Santalum haleakalae Hillebr. var. haleakalae
Dicot
Dicot
Santalaceae
Santalaceae
Scaevola taccada (Gaertn.) Roxb.
Dicot
Goodeniaceae
Harbaugh et al. 2010:836
Harbaugh et al. 2010:833
Green 1991:121/Shannon et al.
1997:801
Botrychium subbifoliatum Brack.
Pteridophyte
Botrychiaceae
Snow et al. 2011:14
Schefflera heptaphylla (L.) Frodin
Schiedea diffusa A.Gray subsp. diffusa
Schiedea diffusa A.Gray subsp. macraei (Sherff)
W.L.Wagner & Weller
Schiedea pubescens Hillebr.
Dicot
Dicot
Araliaceae
Caryophyllaceae
Frohlich & Lau 2012:28
Wagner et al. 2005:103
Dicot
Dicot
Caryophyllaceae
Caryophyllaceae
Wagner et al. 2005:106
Wagner et al. 2005:99
Schiedea pentandra W.L.Wagner & E.M.Harris
Schiedea spergulina A.Gray
Schiedea spergulina A.Gray
Dicot
Dicot
Dicot
Caryophyllaceae
Caryophyllaceae
Caryophyllaceae
Wagner & Harris 2000:159
Wagner et al. 2005:127
Wagner et al. 2005:127
Monocot
Poaceae
Herbst & Clayton 1998:18/ Snow
& Lau 2010:47
Monocot
Monocot
Poaceae
Cyperaceae
Monocot
Cyperaceae
Ruellia graecizans Backer
Rumex crispus L.
Rytidosperma pilosum sensu Herbst & Clayton (1998),
non (R.Br.) Connor & Edgar
Salsola kali sensu Wagner et al. (1990), non L.
Santalum freycinetianum Gaudich. var. freycinetianum
Santalum freycinetianum Gaudich. var. lanaiense Rock
Santalum freycinetianum Gaudich. var. pyrularium
(A.Gray) Stemmerm.
Santalum haleakalae Hillebr.
Scaevola sericea Vahl
Sceptridium subbifoliatum (Brack.) W.H.Wagner &
F.S.Wagner
Schefflera taiwaniana sensu Daehler & Baker (2006),
non (Nakai) Kaneh.
Schiedea diffusa A.Gray
Schiedea diffusa A.Gray var. macraei Sherff
Schiedea pubescens Hillebr. var. pubescens
Schiedea pubescens Hillebr. var. purpurascens Sherff,
invalid name
Schiedea spergulina A.Gray var. leiopoda Sherff
Schiedea spergulina A.Gray var. spergulina
Current disposition
Schizachyrium condensatum sensu Lorence et al. (1995) Andropogon glomeratus (Walter) Britton, Sterns &
et Herbst et al. (2004), in part, non (Kunth) Nees
Poggenb. var. pumilus (Vasey) L.H.Dewey
Schizachyrium scoparium sensu Lorence et al. (1995),
in part, non (Michx.) Nash
Andropogon bicornis L.
Schoenoplectus juncoides (Roxb.) Palla
Schoenoplectiella juncoides (Roxb.) Lye
Schoenoplectus lacustris (L.) Palla subsp. validus (Vahl)
T.Koyama
Schoenoplectus tabernaemontani (C.C.Gmel.) Palla
D-23
Group
Family
Snow & Lau 2010:46
Lye 2003:25/Wagner et al. 2012:82
Smith 1995:98/Kennedy et al.
2010:21
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Selenicereus grandiflorus sensu Lorence et al. (1995),
non (L.) Britton & Rose
Senecio confusus Britten
Selenicereus macdonaldiae (Hook.) Britton & Rose
Pseudogynoxys chenopodioides (Kunth) Cabrera
Dicot
Dicot
Cactaceae
Asteraceae
Senecio mikanioides Otto ex Walp.
Delairea odorata Lem.
Dicot
Asteraceae
Senna surattensis (Burm.f.) H.S.Irwin & Barneby subsp.
retusa (Vogel) Randell
Setaria gracilis Kunth
Sherardia arvensis sensu Oppenheimer (2008), non L.
Sicyocarya kipahuluensis H.St.John
Senna gaudichaudii (Hook. & Arn.) H.S.Irwin &
Barneby
Setaria parviflora (Poir.) Kerguélen
Galium aparine L.
Sicyos macrophyllus A.Gray
Sicyos lanceoloideus (H.St.John) W.L.Wagner &
D.R.Herbst
Sicyos albus (H.St.John) I.Telford
Sicyos lanceoloideus (H.St.John) W.L.Wagner &
D.R.Herbst
Dicot
Monocot
Dicot
Dicot
Fabaceae
Poaceae
Rubiaceae
Cucurbitaceae
Literature source
Herbst & Wagner 1999:16/ Staples
& Herbst 2005:215
Staples & Herbst 2005:163
Jeffrey 1986:933/Herbst & Wagner
1999:14
Staples & Herbst 2005:326 call this
Senna surattensis (Burm.f.)
H.S.Irwin & Barneby subsp. retusa
(Vogel) Randell
Herbst & Clayton 1998:35
Frohlich & Lau 2012:45
Wagner & Shannon 1999:444
Dicot
Dicot
Cucurbitaceae
Cucurbitaceae
Wagner & Shannon 1999:444
Wagner et al. 2012:30
Dicot
Cucurbitaceae
Plectranthus scutellarioides (L.) R. Br.
Sorghum halepense (L.) Pers.
Dicot
Monocot
Lamiaceae
Poaceae
Wagner & Shannon 1999:444
Grayer et al. 2010/Wagner et al.
2012:48
Clayton & Snow 2010:84
Spermacoce exilis (L.O.Williams) C.D.Adams
Spermacoce exilis (L.O.Williams) C.D.Adams
Dicot
Dicot
Rubiaceae
Rubiaceae
Lorence & Flynn 2006:4
Lorence et al. 1995:51
Spermacoce prostrata Aubl.
Dicot
Rubiaceae
Sphaerocionium lanceolatum (Hook. & Arn.) Copel.
Hymenophyllum lanceolatum Hook. & Arn.
Pteridophyte
Hymenophyllaceae
Sphaerocionium obtusum (Hook. & Arn.) Copel.
Spirodela punctata (G.Mey.) C.H.Thomps.
Sporobolus diander (Retz.) P.Beauv. (spelling change)
Sporobolus indicus sensu Wagner et al. (1990) et
Warren (1993), in part, non (L.) R.Br.
Sporobolus piliferus sensu Snow (2008), non (Trin.)
Kunth
Sporobolus pyramidalis sensu Herbst & Clayton (1998),
non P.Beauv.
Sporobolus pyramidatus sensu Wagner & Herbst
(1995), in part, non (Lam.) Hitchc.
Hymenophyllum obtusum Hook. & Arn.
Landoltia punctata (G.Mey.) Les & D.J.Crawford
Sporobolus diandrus (Retz.) P.Beauv.
Pteridophyte
Monocot
Monocot
Hymenophyllaceae
Lemnaceae
Poaceae
Oppenheimer 2003:23
Ebihara et al. 2006:230/Snow et al.
2011:13
Ebihara et al. 2006:230/Snow et al.
2011:13
Staples et al. 2002:11
Clayton & Snow 2010:47
Sporobolus africanus (Poir.) Robyns & Tournay
Monocot
Poaceae
Herbst & Clayton 1998:36
Sporobolus pyramidatus (Lam.) Hitchc.
Monocot
Poaceae
Snow & Lau 2010:56
Sporobolus elongatus R.Br.
Monocot
Poaceae
Snow 2008:41
Sporobolus elongatus R.Br.
Monocot
Poaceae
Stachytarpheta cayennensis (Rich.) Vahl
Dicot
Verbenaceae
Snow 2008:41
Munir 1992:145/Herbst & Wagner
1999:32
Sicyocarya lanceoloidea H.St.John
Sicyos alba (H.St.John) I.Telford (spelling change)
Sicyos sp. A
Solenostemon scutellarioides (L.) Codd
Sorghum halpense (L.) Pers. (spelling change)
Spermacoce capitata sensu Wagner et al. (1997), non
Ruiz & Pav.
Spermacoce mauritiana Gideon
Spermacoce ovalifolia sensu Lorence et al. (1995), non
(Mart. & Gal.) Hemsl.
Stachytarpheta dichotoma (Ruiz & Pav.) Vahl
Current disposition
D-24
Group
Family
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Stachytarpheta dichotoma sensu Wagner et al. (1990),
non (Ruiz & Pav.) Vahl
Stachytarpheta urticifolia (Salisb.) Sims
Stapelia nobilis N.E.Br.
Stenogyne kaalae Wawra
Stenogyne sherffii O.Deg.
Stipa cernua Stebbins & Love
Stylosanthes fruticosa sensu Wagner et al. (1990), non
(Retz.) Alston
Styphelia tameiameiae (Cham. & Schltdl.) F.Muell.
Symphyotrichum squamatum sensu Nesom (1994), non
(Spreng.) Nesom
Symphyotrichum subulatum (Michx.) Nesom var.
parviflorum (Nees) Sundberg
Syzygium sandwicensis (A.Gray) Nied.
Talinum triangulare (Jacq.) Willd.
Current disposition
Family
Literature source
Munir 1992:156/Herbst & Wagner
1999:32
Munir 1992:145/Herbst & Wagner
1999:32
Staples & Herbst 2005:144
Wagner & Weller 1999:449
Stachytarpheta australis Moldenke
Dicot
Verbenaceae
Stachytarpheta cayennensis (Rich.) Vahl
Stapelia gigantea N.E.Br.
Stenogyne kaalae Wawra subsp. kaalae
Stenogyne kaalae Wawra subsp. sherffii (O.Deg.)
W.L.Wagner & Weller
Dicot
Dicot
Dicot
Verbenaceae
Asclepiadaceae
Lamiaceae
Dicot
Lamiaceae
Nassella cernua (Stebbins & Love) Barkworth
Monocot
Poaceae
Wagner & Weller 1999:449
Barkworth 1990:609/Snow
2008:39
Stylosanthes scabra Vogel
Leptecophylla tameiameiae (Cham. & Schltdl.)
C.M.Weiller
Symphyotrichum expansum (Poepp. ex Spreng.)
G.L.Nesom
Symphyotrichum expansum (Poepp. ex Spreng.)
G.L.Nesom
Syzygium sandwicense (A.Gray) Müll.Berol.
Talinum fruticosum (L.) Juss.
Dicot
Fabaceae
Herbst et al. 2004:7
Dicot
Epacridaceae
Dicot
Asteraceae
Weiller 1999/Staples et al. 2003:11
Sundberg 2004:907; Nesom
2005:2133
Dicot
Dicot
Dicot
Asteraceae
Myrtaceae
Portulacaceae
Dicot
Dicot
Malvaceae
Araliaceae
Nesom 2005:2133
Wagner et al. 2012:55
Wagner & Herbst 1995:25
Pfeil et al. 2002/Wagner & Herbst
2003:33
Lowry & Plunkett 2010:73
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Monocot
Dicot
Araliaceae
Araliaceae
Araliaceae
Araliaceae
Araliaceae
Araliaceae
Araliaceae
Araliaceae
Poaceae
Meliaceae
Lowry & Plunkett 2010:74
Lowry & Plunkett 2010:74
Lowry & Plunkett 2010:74
Lowry & Plunkett 2010:74
Lowry & Plunkett 2010:74
Lowry & Plunkett 2010:74
Lowry & Plunkett 2010:75
Lowry & Plunkett 2010:75
Snow & Lau 2010:56
Herbst & Wagner 1999:24
Dicot
Scrophulariaceae
Monocot
Monocot
Cyperaceae
Commelinaceae
Staples et al. 2002:15,16
Tucker 1994:180/Herbst & Wagner
1999:20/Wagner & Herbst
1999:1901
Staples & Herbst 2005:648
Talipariti tiliaceum (L.) Fryxell
Tetraplasandra bisattenuata Sherff
Hibiscus tiliaceus L.
Polyscias bisattenuata (Sherff) Lowry & G.M.Plunkett
Polyscias flynnii (Lowry & K.R.Wood) Lowry &
Tetraplasandra flynnii Lowry & K.R.Wood
G.M.Plunkett
Tetraplasandra gymnocarpa (Hillebr.) Sherff
Polyscias gymnocarpa (Hillebr.) Lowry & G.M.Plunkett
Tetraplasandra hawaiensis A.Gray
Polyscias hawaiensis (A.Gray) Lowry & G.M.Plunkett
Tetraplasandra kavaiensis (H.Mann) Sherff
Polyscias kavaiensis (H.Mann) Lowry & G.M.Plunkett
Tetraplasandra lydgatei (Hillebr.) Harms
Polyscias lydgatei (Hillebr.) Lowry & G.M.Plunkett
Tetraplasandra oahuensis (A.Gray) Harms
Polyscias oahuensis (A.Gray) Lowry & G.M.Plunkett
Tetraplasandra waialealae Rock
Polyscias waialealae (Rock) Lowry & G.M.Plunkett
Tetraplasandra waimeae Wawra
Polyscias waimeae (Wawra) Lowry & G.M.Plunkett
Thysanolaena maxima (Roxb.) Kuntze
Thysanolaena latifolia (Roxb ex Hornem.) Honda
Toona ciliata M.Roem. var. australis (F.Muell.) Bahadur Toona ciliata M.Roem.
Torenia asiatica sensu Wagner et al. (1990), in part, non
L.
Torenia glabra Osbeck
Torulinium odoratum (L.) S.Hooper subsp. auriculatum
(Nees & Meyen ex Kunth) T.Koyama
Tradescantia geniculata Jacq.
Group
Cyperus odoratus L.
Gibasis geniculata (Jacq.) Rohw.
D-25
Synonymy List Arranged By Old Name
Old name (synonym, misapplied name)
Trematolobelia lustrialis H.St.John
Trematolobelia macrostachys (Hook. & Arn.) Zahlbr.
var. kaalae (O.Deg.) E.Wimm.
Trisetum glomeratum sensu Wagner et al. (1990), in
part, non (Kunth) Trin.
Current disposition
Trematolobelia wimmeri O.Deg. & I.Deg.
Group
Dicot
Family
Campanulaceae
Literature source
Lammers 2009:140
Trematolobelia kaalae (O.Deg. ) Lammers
Dicot
Campanulaceae
Lammers 2009:134
Trisetum inaequale Whitney
Monocot
Poaceae
Monocot
Poaceae
Pteridophyte
Dicot
Monocot
Monocot
Dicot
Dicot
Monocot
Hymenophyllaceae
Asteraceae
Poaceae
Poaceae
Sterculiaceae
Asteraceae
Araceae
Dicot
Asteraceae
Herbst & Clayton 1998:37
Zuloaga et al. 2003:631/ Wagner &
Herbst 2003:58/ Clayton & Snow
2010:60
Ebihara et al. 2006:238/Snow et al.
2011:13
Wagner et al. 1997:52
Wagner et al. 2012:101
Wagner et al. 2012:101
Wagner et al. 2012:71
Wagner et al. 1997:55
Herbst et al. 2004:3
Staples et al. 2002:6; Wagner &
Herbst 2003:11
Urochloa subquadripara (Trin.) R.D.Webster
Urochloa distachya (L.) T.Q.Nguyen
Crepidomanes draytonianum (Brack.) Ebihara & K.
Vandenboschia draytoniana (Brack.) Copel.
Iwats.
Vernonia cinerea (L.) Less. var. parviflora (Reinw.) DC. Cyanthillium cinereum (L.) H.Rob.
Vulpia bromoides (L.) Gray
Festuca bromoides L.
Vulpia myuros (L.) C.C.Gmel.
Festuca myuros L.
Waltheria pyrolifolia A.Gray
Waltheria indica L.
Wedelia trilobata (L.) A.S.Hitchc.
Sphagneticola trilobata (L.) Pruski
Xanthosoma roseum Schott
Xanthosoma robustum Schott
Zinnia maritima Kunth [Note: not a syn., but Z. palmeri
sometimes considered a syn. of Z. maritima
Zinnia palmeri A.Gray
D-26
Newly Described Taxa
Appendix E. Newly described taxa since 1990 in the Hawaiian Islands
This list compiles all newly described species and infraspecific taxa following
publication of Manual of the Flowering Plants of Hawai‘i (Wagner et al. 1990) and
Hawai‘i's Ferns and Fern Allies (Palmer 2003), and treated as recognized taxa in
this publication. This list is arranged in a nested alphabetical sequence by major
group, family, and basionym. In some cases, the original name has since changed,
and these new combinations are noted. The original citation describing the taxon is
provided, along with its island distribution. Two of the novelties are non-native taxa
in Passiflora and Ceratopteris.
C. arborescens subsp. waikoluensis [Lammers 1991]). Six species were accepted in
the Manual as described: Cyrtandra hematos, Geranium hanaense, Panicum lineale,
Phyllostegia warshaueri, Sicyos erostratus, and S. waimanaloensis, and another
(Cyrtandra kamoolaensis) was accepted but not treated. Two species were accepted
under new combinations: Sicyos hillebrandii var. anunu [=Sicyos anunu] and
Cladocarpa herbstii [=Sicyos herbstii]. Finally, one species, Cyrtandra
limahuliensis, was treated in the Manual but later synonymized into C. kealiae
subsp. kealiae (Wagner & Lorence 2000). Except for Cyrtandra kamoolaensis, none
of these taxa are included in this appendix.
The reader will note that some of the new taxa were actually published prior to
the Manual publication date in 1990. These were taxa described by prominent
Hawaiian botanist Dr. Harold St. John, who, in a flurry of publication activity
between 1987 and 1989, described over 750 new taxa of Hawaiian vascular plants
(Wagner et al. 1990). Because St. John’s novelties were published so close to the
Manual publication deadline schedule, however, there was not enough time to
examine most of the type specimens and accept or synonymize them in the Manual;
thus, most of the St. John names are not included there.
Most of the new St. John names have since been dealt with in a variety of
taxonomic papers [Lammers 1991 (Clermontia); Marr & Bohm 1997 (Lysimachia);
Herbst & Wagner 1999; Wagner & Shannon 1999 (Sicyos); Wagner 1999a
(Phyllostegia); Wagner & Lorence 2000; Wagner et al. 2005 (Schiedea); Lammers
2005 (Delissea)] and in the Smithsonian Institution’s Flora of the Hawaiian Islands
website (http://botany.si.edu/pacificislandbiodiversity/hawaiianflora/index.htm). In
this appendix, we include only those St. John novelties published between 1987 and
1989 that have been accepted by subsequent workers (either as originally published
or as a new combination), as a full accounting of all names published during this
period was not within the scope of this checklist.
St. John novelties that were included in the Manual included 12 new species of
Touchardia (all synonymized under T. latifolia) and 13 new species of Clermontia
(all synonymized, although one, Clermontia waikoluensis, was soon resurrected as
Group
Family
Dicot
Dicot
Amaranthaceae
Araliaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Campanulaceae
Campanulaceae
Campanulaceae
Campanulaceae
Campanulaceae
Campanulaceae
Campanulaceae
Campanulaceae
Scientific name
Nototrichium divaricatum Lorence
Tetraplasandra flynnii Lowry & K.R.Wood [=Polyscias
flynnii (Lowry & K.R.Wood) Lowry & G.M.Plunkett
Dubautia carrii B.G.Baldwin & Friar
Dubautia hanaulaensis B.G.Baldwin
Dubautia kalalauensis B.G.Baldwin & G.D.Carr
Dubautia kenwoodii G.D.Carr
Dubautia syndetica G.D.Carr & Lorence
Clermontia grandiflora Gaudich. subsp. maxima Lammers
Cyanea dolichopoda Lammers & Lorence
Cyanea duvalliorum Lammers & H.Oppenh.
Cyanea kuhihewa Lammers
Cyanea magnicalyx Lammers
Cyanea maritae Lammers & H.Oppenh.
Cyanea minutiflora Lammers
Cyanea pseudofauriei Lammers
Status
end
end
end
end
end
end
end
end
end
end
end
end
end
end
end
E-1
Literature source
Lorence 1996:64 (SPNOV/K)
Lowry & Wood 2000:40 (SPNOV/K [as Tetraplasandra flynnii]); Lowry &
Plunkett 2010:74 (COMBNOV:Syn. T. flynnii)
Baldwin & Friar 2010:2 (SPNOV/Mo)
Baldwin & Friar 2010:5 (SPNOV/WM)
Baldwin & Carr 2005:259 (SPNOV/K)
Carr 1998:8 (SPNOV/K)
Carr & Lorence 1998:4 (SPNOV/K)
Lammers 1991:77 (SSPNOV/EM)
Lammers & Lorence 1993:432 (SPNOV/K)
Lammers 2004:89 (SPNOV/EM)
Lammers 1996:238 (SPNOV/K)
Lammers 2004:85 (SPNOV/WM)
Lammers 2004:92 (SPNOV/EM)
Lammers 2004:98 (SPNOV/K)
Lammers 2004:96 (SPNOV/K)
Newly Described Taxa
Dicot
Campanulaceae
Dicot
Campanulaceae
Dicot
Campanulaceae
Dicot
Campanulaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Campanulaceae
Campanulaceae
Campanulaceae
Caryophyllaceae
Caryophyllaceae
Caryophyllaceae
Caryophyllaceae
Caryophyllaceae
Dicot
Dicot
Caryophyllaceae
Cucurbitaceae
Dicot
Euphorbiaceae
Dicot
Dicot
Fabaceae
Fabaceae
Dicot
Dicot
Dicot
Gesneriaceae
Gesneriaceae
Gesneriaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Gesneriaceae
Gesneriaceae
Gesneriaceae
Goodeniaceae
Lamiaceae
Lamiaceae
Lamiaceae
Lamiaceae
Lamiaceae
Lamiaceae
Delissea eleeleensis H.St.John [=Cyanea eleeleensis
(H.St.John) Lammers
Delissea habenata H.St.John [=Cyanea habenata
(H.St.John) Lammers
Delissea kahiliensis H.St.John [=Cyanea kahiliensis
(H.St.John) Lammers
end
Delissea kolekoleensis H.St.John [=Cyanea kolekoleensis
(H.St.John) Lammers
Delissea takeuchii Lammers
Delissea waianaeensis Lammers
Lobelia wahiawa Lammers
Schiedea attenuata W.L.Wagner, Weller & A.K.Sakai
Schiedea jacobii W.L.Wagner, Weller & A.C.Medeiros
Schiedea kauaiensis H.St.John
Schiedea laui W.L.Wagner & Weller
Schiedea pentandra W.L.Wagner & E.M.Harris
end
Schiedea perlmanii W.L.Wagner & Weller
Sicyocarya lanceoloidea H.St.John [=Sicyos lanceoloideus
(H.St.John) W.L.Wagner & D.R.Herbst
Chamaesyce eleanoriae Lorence & W.L.Wagner
[=Euphorbia eleanoriae (Lorence & W.L.Wagner)
Govaerts
Kanaloa kahoolawensis Lorence & K.R.Wood
Mucuna sloanei Fawc. & Rendle var. persericea WilmotDear
Cyrtandra ferripilosa H.St.John
Cyrtandra kamoolaensis H.St.John
Cyrtandra kealiae Wawra subsp. urceolata W.L.Wagner &
Lorence
Cyrtandra nanawaleensis H.St.John
Cyrtandra paliku W.L.Wagner, K.R.Wood & Lorence
Cyrtandra wagneri Lorence & Perlman
Scaevola hobdyi W.L.Wagner
Phyllostegia kaalaensis H.St.John
Phyllostegia kahiliensis H.St.John
Phyllostegia micrantha H.St.John
Phyllostegia pilosa H.St.John
Phyllostegia renovans W.L.Wagner
Stenogyne kauaulaensis K.R.Wood & H.Oppenheimer
end
end
end
end
end
end
end
end
end
end
end
end
end
St. John 1987a:341 (SPNOV/K [as Delissea eleeleensis]); Lammers 1992:129
(COMBNOV:Syn. D. eleeleensis)
St. John 1987a:342 (SPNOV/K [as Delissea habenata]); Lammers 1998:33
(COMBNOV:Syn. D. habenata)
St. John 1987a:343 (SPNOV/K [as Delissea kahiliensis]); Wagner et al.
1990:464 (K [as Cyanea spathulata subsp. longipetiolata); Lammers 1998:33
(COMBNOV:Syn. D. kahiliensis, C. spathulata subsp. longipetiolata)
St. John 1987a:344 (SPNOV/K [as Delissea kolekoleensis]); Lammers 1992:130
(COMBNOV:Syn. D. kolekoleensis)
Lammers 2005:38 (SPNOV/O)
Lammers 2005:34 (SPNOV/O)
Lammers 2007:802 (SPNOV/K)
Wagner et al. 1994:187 (SPNOV/K)
Wagner, Weller et al. 1999:284 (SPNOV/EM)
St. John 1988a:177 (SPNOV/K)
Wagner et al. 2005:82 (SPNOV/Mo)
Wagner et al. 1990:519 (O [as Schiedea pubescens var. purpurascens]); Wagner
& Harris 2000:159 (SPNOV/O [S. pubescens var. purpurascens, an invalid
name])
Wagner et al. 2005:71 (SPNOV/K)
St. John 1987g:52 (SPNOV/K [as Sicyocarya lanceoloidea]); Wagner &
Shannon 1999:444 (COMBNOV:Syn. Sicyocarya lanceoloidea)
Lorence & Wagner 1996:68 (SPNOV/K) [as Chamaesyce eleanoriae]; Govaerts
et al. 2000:717 (COMBNOV:Syn. C. eleanoriae)
end
end
Lorence & Wood 1994:137 (SPNOV/Ka)
Wilmot-Dear 1990:27 (VARNOV/EM)
end
end
end
St. John 1987e:497 (SPNOV/EM)
St. John 1987f:499 (SPNOV/K);
Wagner & Lorence 2000:19 (SSPNOV/K)
end
end
end
end
end
end
end
end
end
end
St. John 1987e:500 (SPNOV/H)
Wagner et al. 2001:146 (SPNOV/K)
Lorence & Perlman 2007:357 (SPNOV/H)
Wagner 1996:225 (SPNOV/WM)
St. John 1987c:176 (SPNOV/O)
St. John 1987c:176 (SPNOV/K,)
St. John 1987c:178 (SPNOV/O,)
St. John 1987c:179 (SPNOV/EM)
Wagner 1999a:280 (SPNOV/K)
Wood & Oppenheimer 2008:544 (SPNOV/WM)
E-2
Newly Described Taxa
Dicot
Dicot
Dicot
Loganiaceae
Malvaceae
Passifloraceae
Dicot
Primulaceae
Dicot
Primulaceae
Labordia lorenciana K.R.Wood, W.L.Wagner & T.Motley
Hibiscadelphus woodii Lorence & W.L.Wagner
Passiflora tarminiana Coppens & Barney
end
end
nat
end
Lysimachia caliginis H.St.John [=L. remyi Hillebr.]
Dicot
Dicot
Primulaceae
Primulaceae
end
Lysimachia haupuensis H.St.John [=L. hillebrandii Hook.f.
ex A.Gray]
Lysimachia iniki K.L.Marr
end
end
Lysimachia kahiliensis H.St.John [=L. daphnoides
(A.Gray) Hillebr.]
Dicot
Primulaceae
Dicot
Dicot
Dicot
Primulaceae
Primulaceae
Primulaceae
Dicot
Rubiaceae
Dicot
Monocot
Monocot
Monocot
Monocot
Rubiaceae
Arecaceae
Arecaceae
Arecaceae
Arecaceae
Monocot
Monocot
Monocot
Monocot
Monocot
Pteridophyte
Pteridophyte
Arecaceae
Arecaceae
Poaceae
Poaceae
Poaceae
Athyriaceae
Pteridaceae
Pteridophyte
Thelypteridaceae
end
Lysimachia ovoidea H.St.John [=L. hillebrandii Hook.f. ex
A.Gray]
Lysimachia pendens K.L.Marr
Lysimachia scopulensis K.L.Marr
Lysimachia waianaeensis H.St.John [=L. hillebrandii
Hook.f. ex A.Gray]
Hedyotis flynnii W.L.Wagner & Lorence [=Kadua flynnii
(W.L.Wagner & Lorence) W.L.Wagner & Lorence
Kadua haupuensis Lorence & W.L.Wagner
Pritchardia bakeri Hodel
Pritchardia flynnii Lorence & Gemmill
Pritchardia gordonii Hodel
Pritchardia limahuliensis H.St.John [=Pritchardia
napaliensis H.St.John
Pritchardia perlmanii Gemmill
Pritchardia woodii Hodel
Festuca aloha Catalán, Soreng & P.M.Peterson
Festuca molokaiensis Soreng, P.M.Peterson & Catalán
Panicum longivaginatum H.St.John
Deparia cataracticola M.Kato
Ceratopteris gaudichaudii Brongn. var. vulgaris Masuyama
& Watano
Pneumatopteris pendens D.D.Palmer [=Cyclosorus
pendens (D.D.Palmer) N.Snow
end
end
end
end
end
end
end
end
end
end
end
end
end
end
end
nat
end
E-3
Wood et al. 2007:195 (SPNOV/K)
Lorence & Wagner 1995:183 (SPNOV/K)
Coppens d'Eeckenbrugge et. al. 2001:9 (SPNOV, NOM: P. mollissima
misapplied)
St. John 1987g:43 (SPNOV/M [as Lysimachia caliginis]);Marr & Bohm
1997:277, SSPNOV [as L. remyi subsp. caliginis], NOM:Syn. L. caliginis;
Herbst & Wagner 1999:31 (NOM:Syn. L. remyi subsp. caliginis)
St. John 1987g:45 (SPNOV/K [as Lysimachia haupuensis]); Marr & Bohm
1997:268 (accept L. haupuensis); Herbst & Wagner 1999:30 (NOM:Syn: L.
haupuensis)
Marr & Bohm 1997:270 (SPNOV/K)
St. John 1987g:45 (SPNOV/K [as Lysimachia kahiliensis]); Marr & Bohm 1997:
272 (accept L. kahiliensis); Herbst & Wagner 1999:30 (NOM:Syn. L.
kahiliensis)
St. John 1987g:48 (SPNOV/K [as Lysimachia ovoidea]); Marr & Bohm
1997:268 (accept L. ovoidea); Herbst & Wagner 1999:30 (NOM:Syn: L.
ovoidea)
Marr & Bohm 1997:275 (SPNOV/K)
Marr & Bohm 1997:265 (SPNOV/K)
St. John 1987g:50 (SPNOV/O [as Lysimachia waianaeensis]); Marr & Bohm
1997:268 (accept L. waianaeensis); Herbst & Wagner 1999:30 (NOM:Syn: L.
waianaeensis)
Wagner & Lorence 1998:311 (SPNOV/K) [as Hedyotis flynnii]; Terrell et al.
2005:833 (COMBNOV:Syn. H. flynnii)
Lorence et al. 2010:138 (SPNOV/K)
Hodel 2009:173 (SPNOV/O)
Lorence & Gemmill 2004:185 (SPNOV/K)
Hodel 2007:S-15 (SPNOV/H)
St. John 1988a:177 (SPNOV/K [as Pritchardia limahuliensis]); Hodel 2007:S-37
(NOM:Syn. P. limahuliensis)
Gemmill 1998:18 (SPNOV/K)
Hodel 2007:S-44 (SPNOV/EM)
Catalán et al. 2009:51 (SPNOV/K)
Catalán et al. 2009:54 (SPNOV/Mo)
St. John 1987b:370 (SPNOV/H)
Kato 2001:3 (SPNOV/K)
Masuyama & Watano 2010:79 (VARNOV/O)
Palmer 2005:81 (SPNOV/K, O, Mo, EM, WM, H [as Pneumatopteris pendens]);
Snow et al. 2011:15 (COMBNOV:Syn. P. pendens)
Resurrected Taxa
Appendix F. Taxa resurrected since 1990 in the Hawaiian Islands
This list compiles all taxa that were treated as taxonomic synonyms in the
Manual of the Flowering Plants of Hawai‘i (Wagner et al. 1990) or Hawai‘i's Ferns
and Fern Allies (Palmer 2003), and have subsequently been resurrected as
recognized entities. This list is arranged in a nested alphabetical sequence by major
group, family, and scientific name.
Group
Family
In the literature source column, the disposition of the taxon in Wagner et al.
(1990) [or in one case, Palmer (2003)] is provided first, followed by a citation for
the source of the resurrection. In two examples in the genus Pritchardia, species
were resurrected and once again synonymized.
Scientific name
Status
Literature source
Dicot
Araliaceae
Polyscias bisattenuata (Sherff) Lowry & G.M.Plunkett
end
Wagner et al. 1990:235 (as Tetraplasandra bisattenuata, syn. of T. oahuensis);
Lowry & Plunkett 2010:73 (RESURRECT/K, COMBNOV:Syn. T.
bisattenuata)
Dicot
Araliaceae
Polyscias lydgatei (Hillebr.) Lowry & G.M.Plunkett
end
Wagner et al. 1990:235 (as Tetraplasandra lydgatei, syn. of T. oahuensis);
Motley 2005:106 (RESURRECT/O); Lowry & Plunkett 2010:74
(COMBNOV:Syn. T. lydgatei)
Dicot
Asteraceae
Dubautia waianapanapaensis G.D.Carr
end
Wagner et al. 1990:297 (syn. of Dubautia dolosa); Carr 1995:1580
(RESURRECT/EM, NOM:Syn. D. dolosa)
Dicot
Asteraceae
Melanthera populifolia (Sherff) W.L.Wagner & H.Rob.
end
Wagner et al. 1990:341 (as Lipochaeta populifolia, syn. of L. subcordata);
Wagner & Robinson 2002:554 (RESURRECT/L, COMBNOV:Syn. L.
populifolia)
Dicot
Brassicaceae
Lepidium orbiculare H.St.John
end
Wagner et al. 1990:409 (syn. of Lepidium serra); Wagner & Herbst 2003:13
(RESURRECT/K)
Dicot
Campanulaceae
Clermontia arborescens (H.Mann) Hillebr. subsp.
waikoluensis (H.St.John) Lammers
end
Wagner et al. 1990:427 (as Clermontia waikoluensis, syn. of C. arborescens
subsp. waihiae); Lammers 1991:25 (RESURRECT/Mo, L, COMBNOV &
STATNOV:Syn. Delissea fauriei)
Dicot
Campanulaceae
Clermontia peleana Rock subsp. singuliflora (Rock)
Lammers
end
Wagner et al. 1990:435 (as Clermontia singuliflora, syn. of C. peleana);
Lammers 1991:32 (RESURRECT/M, H, COMBNOV & STATNOV:Syn. C.
singuliflora)
Dicot
Campanulaceae
Cyanea lobata H.Mann subsp. baldwinii (C.N.Forbes &
G.C.Munro) Lammers
end
Wagner et al. 1990:454 (as Cyanea baldwinii, syn. of C. lobata); Lammers
1999:387 (RESURRECT/L, COMBNOV & STATNOV:Syn. C. baldwinii)
Dicot
Campanulaceae
Cyanea mauiensis (Rock) Lammers
end
Wagner et al. 1990:451 (as Cyanea grimesiana var. mauiensis, syn. of C.
grimesiana subsp. grimesiana); Lammers 1998:31 (RESURRECT/M,
COMBNOV & STATNOV:Syn. C. grimesiana var. mauiensis)
Dicot
Campanulaceae
Cyanea munroi (Hosaka) Lammers
end
Wagner et al. 1990:451 (as Cyanea grimesiana var. munroi, syn. of C.
grimesiana subsp. grimesiana); Lammers 1998:31 (RESURRECT/Mo, L,
COMBNOV & STATNOV:Syn. C. grimesiana var. munroi)
Dicot
Campanulaceae
Cyanea remyi Rock
end
Wagner et al. 1990:466 (not placed, possibly referable to Cyanea truncata);
Lammers & Lorence 1993:434 (RESURRECT/K)
Dicot
Campanulaceae
Cyanea salicina H.Lév.
end
Wagner et al. 1990:461 (syn. of Cyanea recta); Wagner & Herbst 1999:1868
(RESURRECT/K, based on Lammers unpubl.)
F-1
Resurrected Taxa
Group
Family
Scientific name
Status
Literature source
Dicot
Campanulaceae
Cyanea sessilifolia (O.Deg.) Lammers
end
Wagner et al. 1990:484 (as Rollandia sessilifolia, syn. of R. longiflora);
Lammers 1998:32 (RESURRECT/O, COMBNOV:Syn. R. sessilifolia)
Dicot
Campanulaceae
Delissea argutidentata (E.Wimm.) H.St.John
end
Wagner et al. 1990:471 (syn. of Delissea undulata subsp. undulata); Lammers
2005:22 (RESURRECT/H)
Dicot
Campanulaceae
Delissea fauriei H.Lév.
end
Wagner et al. 1990:449 (species dubia); Lammers 1991:25 (NOM:syn. of
Clermontia arborescens subsp. waikoluensis); Lammers 2005:42
(RESURRECT/Mo)
Dicot
Campanulaceae
Delissea subcordata Gaudich. subsp. obtusifolia (Wawra)
Lammers
end
Wagner et al. 1990:471 (as Delissea subcordata var. obtusifolia, syn. of D.
subcordata); Lammers 2005:34 (RESURRECT/O, COMBNOV &
STATNOV:Syn. D. subcordata var. obtusifolia)
Dicot
Campanulaceae
Lobelia gloria-montis Rock subsp. longibracteata (Rock)
Lammers
end
Wagner et al. 1990:476 (as Lobelia gloria-montis var. longibracteata, syn. of L.
gloria-montis); Lammers 2007:806 (RESURRECT/EM, WM, COMBNOV &
STATNOV, Syn. L. gloria-montis var. longibracteata)
Dicot
Campanulaceae
Trematolobelia auriculata H.St.John
end
Wagner et al. 1990:486 (syn. of Trematolobelia macrostachys); Lammers
2009:138 (RESURRECT/L)
Dicot
Campanulaceae
Trematolobelia kaalae (O.Deg.) Lammers
end
Wagner et al. 1990:486 (as Trematolobelia macrostachys var. kaalae, syn. of T.
macrostachys); Lammers 2009:134 (RESURRECT/O, STATNOV, Syn. T.
macrostachys var. kaalae)
Dicot
Campanulaceae
Trematolobelia rockii H.St.John
end
Wagner et al. 1990:486 (syn. of Trematolobelia macrostachys); Lammers
2009:138 (RESURRECT/Mo)
Dicot
Campanulaceae
Trematolobelia wimmeri O.Deg. & I.Deg.
end
Wagner et al. 1990:486 (syn. of Trematolobelia grandifolia); Lammers
2009:140 (RESURRECT/H)
Dicot
Caryophyllaceae
Schiedea diffusa A.Gray subsp. macraei (Sherff)
W.L.Wagner & Weller
end
Wagner et al. 1990:511 (as Schiedea diffusa var. macraei, syn. of S. diffusa);
Wagner et al. 2005:106 (RESURRECT/H, COMBNOV & STATNOV:Syn. S.
diffusa var. macraei)
Dicot
Caryophyllaceae
Schiedea hawaiiensis Hillebr.
end
Wagner et al. 1990:511 (syn. of Schiedea diffusa); Herbst & Wagner 1999:18
(RESURRECT/H)
Dicot
Caryophyllaceae
Schiedea sarmentosa O.Deg. & Sherff
end
Wagner et al. 1990:517 (syn. of Schiedea menziesii); Wagner et al. 2005:116
(RESURRECT/Mo)
Dicot
Fabaceae
Acacia koaia Hillebr.
end
Wagner et al. 1990:641 (syn. of Acacia koa); Wagner & Herbst 1999:1875
(RESURRECT/K, O?, Mo, L, M, H)
Dicot
Gesneriaceae
Cyrtandra wainihaensis H.Lév.
end
Wagner et al. 1990:762 (syn. of Cyrtandra kauaiensis); Wagner & Herbst
2003:29 (RESURRECT/K)
Dicot
Lamiaceae
Phyllostegia haliakalae Wawra
end
Wagner et al. 1990:821 [as syn. of P. mollis]; Wagner 1999b:269
(RESURRECT, Mo, L, EM, NOM:Syn. P. imminuta)
Dicot
Lamiaceae
Phyllostegia parviflora (Gaudich.) Benth. var. lydgatei
(Sherff) W.L.Wagner
end
Wagner et al. 1990:822 (as Phyllostegia mollis var. lydgatei, syn. of P.
parviflora var. parviflora); Wagner 1999a:282 (RESURRECT/O,
COMBNOV:Syn. P. mollis var. lydgatei)
Dicot
Lamiaceae
Stenogyne kaalae Wawra subsp. sherffii (O.Deg.)
W.L.Wagner & Weller
end
Wagner et al. 1990:838 (as Stenogyne sherffii, syn. of S. kaalae); Wagner &
Weller 1999:449 (RESURRECT/O, COMBNOV & STATNOV:Syn. S. sherffii)
F-2
Resurrected Taxa
Group
Family
Scientific name
Status
Literature source
Dicot
Lamiaceae
Stenogyne kealiae Wawra
end
Wagner et al. 1990:841 (syn. of Stenogyne purpurea); Wagner & Weller
1991:50 (RESURRECT/K)
Dicot
Loganiaceae
Labordia triflora Hillebr.
end
Wagner et al. 1990:861 (syn. of Labordia tinifolia var. lanaiensis); Motley
1995:221 (RESURRECT/Mo)
Dicot
Malvaceae
Hibiscus brackenridgei A.Gray subsp. molokaianus (Rock
ex Caum) F.D.Wilson
end
Wagner et al. 1990:883 (as H. brackenridgei var. molokaiana, syn. of Hibiscus
brackenridgei subsp. brackenridgei); Wilson 1993:278 (RESURRECT/Mo,
COMBNOV & STATNOV:Syn. H. brackenridgei var. molokaiana)
Dicot
Myoporaceae
Myoporum degeneri (G.L.Webster) O.Deg. & I.Deg.
end
Wagner et al. 1990:928 (syn. of Myoporum sandwicense); Chinnock 2007:145
(RESURRECT/EM) [Note: not yet accepted; status under study]
Dicot
Myoporaceae
Myoporum sandwicense A.Gray subsp. lanaiense
(G.L.Webster) Chinnock
end
Wagner et al. 1990:928 (as Myoporum sandwicense var. lanaiense, syn. of M.
sandwicense); Chinnock 2007:144 (RESURRECT/L, COMBNOV &
STATNOV) [Note: not yet accepted; status under study]
Dicot
Myoporaceae
Myoporum stellatum (G.L.Webster) O.Deg. & I.Deg.
end
Wagner et al. 1990:928 (syn. of Myoporum sandwicense); Chinnock
2007:146/Wagner et al. 2005– (RESURRECT/O)
Dicot
Pittosporaceae
Pittosporum halophilum Rock
end
Wagner et al. 1990:1040 (syn. of Pittosporum confertiflorum); Wagner &
Herbst 1999:1887 (RESURRECT/Mo)
Dicot
Rubiaceae
Kadua cordata Cham. & Schltdl. subsp. waimeae (Wawra)
W.L.Wagner & Lorence
end
Wagner et al. 1990:1150 (as Hedyotis glaucifolia var. waimeae, syn. of H.
schlechtendahliana var. schlechtendahliana); Wagner & Lorence 1998:316
(RESURRECT/K, COMBNOV & STATNOV [as H. schlechtendahliana subsp.
waimeae, NOM:Syn. H. glaucifolia var. waimeae]; Terrell et al. 2005:833
(COMBNOV:Syn. H. schlechtendahliana subsp. waimeae)
Dicot
Rutaceae
Melicope hiiakae (B.C.Stone) T.G.Hartley & B.C.Stone
end
Wagner et al. 1990:1206 (as Pelea hiiakae, syn. of P. wawraeana); Hartley &
Stone 1989:122 (COMBNOV:Syn. P. hiiakae); Wagner & Herbst 1999:1894
(RESURRECT/O)
Dicot
Santalaceae
Santalum involutum H.St.John
end
Wagner et al. 1990:1221 (syn. of Santalum freycinetianum var. pyrularium);
Harbaugh et al. 2010:835 (RESURRECT/K)
Dicot
Sterculiaceae
Waltheria pyrolifolia A.Gray
end
Wagner et al. 1990:1280 (as Waltheria pyrolaefolia, syn. of W. indica);
Saunders 2011:79 (RESURRECT/WM); Wagner et al. 2012:71 (NOM:Syn. W.
pyrifolia)
Monocot
Arecaceae
Pritchardia lanaiensis Becc. & Rock [=P. glabrata Becc. &
Rock
end
Wagner et al. 1990:1365 (Pritchardia lanaiensis unplaced, ID doubtful)); Herbst
& Wagner 1999:13 (RESURRECT/L); Hodel 2007:S-14 (NOM:Syn. P.
lanaiensis)
Monocot
Arecaceae
Pritchardia kahukuensis Caum [=P. martii (Gaudich.)
H.Wendl.
end
Wagner et al. 1990:1372 (O (syn. of Pritchardia martii]); Hodel 2007:S-20
(RESURRECT/O); Wagner et al. 2012:77 (P. kahukuensis not recognized)
Pteridophyte
Aspleniaceae
Asplenium diellaciniatum Viane
end
Palmer 2003:117 (as Diellia laciniata, syn. of D. erecta; Wagner et al. 2012:103
(RESURRECT/K, NOM:Syn. D. laciniata [D. Lorence, in prep.)
Pteridophyte
Pteridaceae
Doryopteris tryonii O.Deg. & I.Deg.
end
Palmer 2003:133 (syn. of Doryopteris decora); Yesilyurt 2005: 557
(RESURRECT/K) [Note: not yet accepted; status under study]
F-3
New Naturalized Taxa
Appendix G. New naturalized taxa since 1990 in the Hawaiian Islands
This list compiles all newly naturalized vascular plant taxa in the Hawaiian
Islands since publication of the Manual of the Flowering Plants of Hawai‘i (Wagner
et al. 1990) and Hawai‘i's Ferns and Fern Allies (Palmer 2003). It is arranged in a
nested alphabetical sequence by major group, family, and scientific name.
Group
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Family
Acanthaceae
Acanthaceae
Acanthaceae
Acanthaceae
Acanthaceae
Acanthaceae
Acanthaceae
Acanthaceae
Acanthaceae
Acanthaceae
Acanthaceae
Aizoaceae
Amaranthaceae
Amaranthaceae
Amaranthaceae
Amaranthaceae
Anacardiaceae
Annonaceae
Annonaceae
Annonaceae
Annonaceae
Apiaceae
Apocynaceae
Apocynaceae
Apocynaceae
Apocynaceae
Apocynaceae
Apocynaceae
Apocynaceae
Aquifoliaceae
Araliaceae
The literature citation for the first recorded documentation of naturalization is
also provided, along with the island(s) on which it was first collected. Some taxa
have undergone name changes since being published, and the old name is so noted
following the citation. Records preceded by an asterisk (*) have been subsequently
judged to be not truly naturalized.
Scientific name
Aphelandra aurantiaca (Scheidw.) Lindl.
Barleria repens Nees
Hypoestes phyllostachya Baker
Justicia carnea Lindl.
Justicia spicigera Schltdl.
Megaskepasma erythrochlamys Lindau
Odontonema cuspidatum (Nees) Kuntze
Pseuderanthemum variabile (R.Br.) Radlk.
Ruellia devosiana E.Morren
Sanchezia parvibracteata Sprague & Hutch.
Sanchezia speciosa Leonard
Sesuvium verrucosum Raf.
Amaranthus graecizans L.
Amaranthus retroflexus L.
Blutaparon vermiculare (L.) Mears
Celosia argentea L.
Rhus taitensis Guill.
Annona cherimola Mill.
Annona muricata L.
Artabotrys hexapetalus (L.f.) Bhandari
*Polyalthia suberosa (Roxb.) Thwaites
Torilis arvensis (Huds.) Link
Alstonia macrophylla Wall. ex G.Don
Carissa macrocarpa (Eckl.) A.DC.
Nerium oleander L.
Rauvolfia vomitoria Afzel.
Stemmadenia litoralis (Kunth) L.Allorge
*Tabernaemontana pandacaqui Poir.
Vinca major L.
Ilex cassine L.
Schefflera arboricola (Hayata) Merr.
Literature source
Daehler & Baker 2006:4 (NEWNAT/O)
Staples et al. 2002:3 (NEWNAT/O)
Herbst & Wagner 1999:12 (NEWNAT/H)
Oppenheimer 2004:8 (NEWNAT/EM)
Staples et al. 2002:4 (NEWNAT/O, Mo)
Frohlich & Lau 2010:3 (NEWNAT/O)
Lorence et al. 1995:20 (NEWNAT/K)
Herbarium Pacificum Staff 1998:8 (NEWNAT/O [as Pseuderanthemum fasciculatum])
Lorence et al. 1995:20 (NEWNAT/K)
Parker & Parsons 2012a:55 (NEWNAT/H)
Lorence et al. 1995:20 (NEWNAT/K)
Frohlich & Lau 2012:27 (NEWNAT/O, Mo, EM)
Herbst et al. 2004:3 (NEWNAT/O)
Wagner et al. 1997:51 (NEWNAT/H)
Wagner & Herbst 1995:14 (NEWNAT/O)
Oppenheimer 2003:5 (NEWNAT/H)
Daehler & Baker 2006:4 (NEWNAT/O)
Staples et al. 2002:4 (NEWNAT/H)
Wysong et al. 2007:2 (NEWNAT/Mo)
Nagata 1995:11 (NEWNAT/O, H)
Frohlich & Lau 2010:4 (NEWNAT/O); Wagner et al. 2012:6 (O record adventive)
Staples et al. 2002:5 (NEWNAT/K, KEY)
Wagner et al. 1997:51 (NEWNAT/H)
Lau & Frohlich 2012:7 (NEWNAT/O)
Parker & Parsons 2012a:55 (NEWNAT/H)
Kennedy et al. 2010:19 (NEWNAT/H)
Lau & Frohlich 2012:7 (NEWNAT/O); Parker & Parsons 2012a:56 (NEWNAT/H)
Frohlich & Lau 2010:4 (NEWNAT/O); Wagner et al. 2012:6 (O record adventive)
Starr et al. 2002:17 (NEWNAT/EM)
Imada et al. 2000:9 (NEWNAT/O, H)
Starr et al. 2003:24 (NEWNAT/EM)
G-1
New Naturalized Taxa
Group
Family
Dicot
Araliaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Araliaceae
Araliaceae
Araliaceae
Asclepiadaceae
Asclepiadaceae
Asclepiadaceae
Asclepiadaceae
Asclepiadaceae
Asclepiadaceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Asteraceae
Basellaceae
Begoniaceae
Begoniaceae
Betulaceae
Bignoniaceae
Scientific name
*Schefflera elegantissima (Veitch ex Masters) Lowry &
Frodin
Schefflera heptaphylla (L.) Frodin
Schefflera insularum (Seem.) Harms
Tetrapanax papyrifer (Hook.) K.Koch
Calotropis gigantea (L.) W.T.Aiton
Calotropis procera (Aiton) W.T.Aiton
Cryptostegia grandiflora Roxb. ex R.Br.
Cryptostegia madagascariensis Bojer ex Decne.
Cynanchum gerrardii (Harv.) Liede
Hoya australis R.Br. ex Traill
Arctotheca calendula (L.) Levyns
Centaurea cyanus L.
Centratherum punctatum subsp. punctatum
Chromolaena odorata (L.) R.M.King & H.Rob.
Crepis vesicaria L. subsp. taraxacifolia (Thuill.) Thuill.
Dyssodia tenuiloba (DC.) B.L.Rob.
Euchiton gymnocephalus (DC.) Holub
Euchiton japonicus (Thunb.) Anderb.
Eupatorium capillifolium (Lam.) Small
Florestina tripteris DC.
Gamochaeta americana (Mill.) Weddell
Gamochaeta argyrinea G.L.Nesom
Gamochaeta pensylvanica (Willd.) Cabrera
Gamochaeta simplicicaulis (Willd. ex Spreng.) Cabrera
Kalimeris indica (L.) Sch.Bip. subsp. indica
Leucanthemum xsuperbum (J.W.Ingram) D.H.Kent
Perityle emoryi Torr.
Pseudognaphalium attenuatum (DC.) Anderb.
Pseudogynoxys chenopodioides (Kunth) Cabrera
Senecio madagascariensis Poir.
Soliva sessilis Ruiz & Pav.
Tagetes erecta L.
Vernonia elliptica DC.
Basella alba L.
Begonia cucullata Willd.
Begonia fusca Liebm.
Alnus nepalensis D.Don
Catalpa longissima (Jacq.) Dum.-Cours.
Literature source
Frohlich & Lau 2010:4 (NEWNAT/O); Wagner et al. 2012:7 (O record adventive)
Frohlich & Lau 2012:28 (NEWNAT/O)
Frohlich & Lau 2012:29 (NEWNAT/O)
Lau & Frohlich 2012:8 (NEWNAT/O); Parker & Parsons 2012a:56 (NEWNAT/H)
Starr et al. 2003:25 (NEWNAT/EM, WM)
Wood & LeGrande 2006:19 (NEWNAT/Le)
Lau & Frohlich 2012:8 (NEWNAT/O)
Staples et al. 2006:6 (NEWNAT/Mo)
Frohlich & Lau 2012:29 (NEWNAT/O)
Lorence & Flynn 1999:3 (NEWNAT/EM)
Starr & Starr 2011:23 (NEWNAT/EM)
Wagner & Herbst 1995:15 (NEWNAT/L)
Lorence et al. 1995:23 (NEWENAT/K)
Frohlich & Lau 2012:30 (NEWNAT/O)
Staples et al. 2002:5 (NEWNAT/H)
Starr et al. 2002:18 (NEWNAT/O, EM, WM)
Nesom 2002:516, 518 (NEWNAT/H, KEY)
Wagner et al. 1997:54 (NEWNAT/H, KEY)
Staples et al. 2003:9 (NEWNAT/H)
Oppenheimer & Bartlett 2000:2 (NEWNAT/WM)
Alford 2012:2 (NEWNAT/EM)
Alford 2012:3 (NEWNAT/O, EM, WM, H)
Alford 2012:3 (NEWNAT/Le, K, O, Mo, L, EM, WM, Ka, H)
Alford 2012:4 (NEWNAT/Mo)
Nagata 1995:11 (NEWNAT/K [as Boltonia indica])
Pratt & Bio 2012:75 (NEWNAT/H)
Starr et al. 2010:62 (NEWNAT/Ka)
Nesom 2001a:1186 (NEWNAT/H)
Parker & Parsons 2012a:57 (NEWNAT/H)
Lorence et al. 1995:24 (NEWNAT/K, H)
Wagner & Herbst 1995:16 (NEWNAT/H)
Kennedy et al. 2010:20 (NEWNAT/O)
Lau & Frohlich 2012:9 (NEWNAT/O)
Nagata 1995:11 (NEWNAT/O)
Lorence et al. 1995:25 (NEWNAT/K, H)
Miller 2003 (NEWNAT/K, O)
Wysong et al. 2007:3 (NEWNAT/Mo)
Lau & Frohlich 2012:9 (NEWNAT/O)
G-2
New Naturalized Taxa
Group
Family
Dicot
Dicot
Dicot
Dicot
Dicot
Bignoniaceae
Bignoniaceae
Bignoniaceae
Bignoniaceae
Bignoniaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Bignoniaceae
Bignoniaceae
Bignoniaceae
Bignoniaceae
Boraginaceae
Boraginaceae
Boraginaceae
Boraginaceae
Boraginaceae
Brassicaceae
Brassicaceae
Brassicaceae
Brassicaceae
Buddlejaceae
Cactaceae
Cactaceae
Cactaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Cactaceae
Cactaceae
Campanulaceae
Campanulaceae
Capparaceae
Caryophyllaceae
Caryophyllaceae
Caryophyllaceae
Casuarinaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Celastraceae
Chenopodiaceae
Chenopodiaceae
Chenopodiaceae
Chrysobalanaceae
Scientific name
Mansoa hymenaea (DC.) A.H.Gentry
Podranea ricasoliana (Tanfani) Sprague
Pyrostegia venusta (Ker Gawl.) Miers
*Radermachera sinica (Hance) Hemsl.
Tabebuia aurea (Silva Manso) Benth. & Hook.f. ex
S.Moore
Tabebuia heterophylla (DC.) Britton
Tabebuia pallida (Lindl.) Miers
Tabebuia rosea (Bertol.) DC.
Tecoma castanifolia (D.Don) Melch.
Carmona retusa (Vahl) Masam.
Cordia alliodora (Ruiz & Pav.) Oken
Cordia dichotoma G.Forst.
Cordia lutea Lam.
Myosotis azorica H.C.Watson
Hirschfeldia incana (L.) Lagr.-Foss.
Raphanus raphanistrum L.
Rapistrum rugosum (L.) All.
Sinapis arvensis L.
Buddleja davidii Franch.
Acanthocereus tetragonus (L.) Hummelinck
Harrisia bonplandii (Parm. ex Pfeiff.) Britton & Rose
Hylocereus costaricensis (F.A.C.Weber) Britton & Rose
Pereskia aculeata Mill.
Selenicereus macdonaldiae (Hook.) Britton & Rose
Platycodon grandiflorus (Jacq.) A.DC.
Wahlenbergia marginata (Thunb.) A.DC.
Capparis mollicella Standl.
Cerastium glomeratum Thuill.
Sagina procumbens L.
Silene armeria L.
Casuarina cunninghamiana Miq. x Casuarina glauca
Sieber ex Spreng.
Catha edulis (Vahl) Forssk. ex Endl.
Atriplex canescens (Pursh) Nutt.
Atriplex maximowicziana Makino
Enchylaena tomentosa R.Br. var. tomentosa
Chrysobalanus icaco L.
Literature source
Lau & Frohlich 2012:10 (NEWNAT/O)
Starr et al. 2004:21 (NEWNAT/EM)
Frohlich & Lau 2012:31 (NEWNAT/K, O)
Frohlich & Lau 2008:4 (NEWNAT/O); Wagner et al. 2012:18 (O record adventive)
Frohlich & Lau 2010:5 (NEWNAT/O)
Oppenheimer 2003:8 (NEWNAT/WM)
Frohlich & Lau 2012:31 (NEWNAT/K, O)
Frohlich & Lau 2010:6 (NEWNAT/O)
Herbarium Pacificum Staff 1998:9 (NEWNAT/O)
Lorence et al. 1995:25 (NEWNAT/K, WM)
Lau & Frohlich 2012:10 (NEWNAT/O)
Herbarium Pacificum Staff 1996:3 (NEWNAT/O)
Parker & Parsons 2012a:58 (NEWNAT/H)
Starr et al. 2008:44 (NEWNAT/EM)
Lorence et al. 1995:26 (NEWNAT/EM, H)
Staples et al. 2003:9 (NEWNAT/K, O, Mo, EM, H)
Lorence et al. 1995:27 (NEWNAT/K)
Herbarium Pacificum Staff 1998:9 (NEWNAT/Mo)
Shannon & Wagner 1996:13 (NEWNAT/K)
Lorence et al. 1995:27 (NEWNAT/K)
Lorence et al. 1995:28 (NEWNAT/K)
Lorence et al. 1995:28 (NEWNAT/K) [Note: BISH voucher (Flynn & Hume 3571) has since
been annotated as Hylocereus cf. trigonus (Haw.) Safford by B. Leuenberger, 16 Oct 2000]
Parker & Parsons 2012b:66 (NEWNAT/Mo?, H)
Lorence et al. 1995:28 (NEWNAT/K [as Selenicereus grandiflorus])
Lau & Frohlich 2012:12 (NEWNAT/O)
Herbst et al. 2004:4, 5 (NEWNAT/H)
Lau & Frohlich 2012:12 (NEWNAT/O)
Starr et al. 2004:21 (NEWNAT/EM)
Oppenheimer 2003:9 (NEWNAT/L, EM, WM)
Starr & Starr 2012:88 (NEWNAT/EM)
Lorence et al. 1995:31 (NEWNAT/K)
Frohlich & Lau 2012:32 (NEWNAT/O)
Staples et al. 2003:10 (NEWNAT/H)
Wagner et al. 1997:55 (NEWNAT/H)
Imada et al. 2000:11 (NEWNAT/M)
Lau & Frohlich 2012:13 (NEWNAT/O)
G-3
New Naturalized Taxa
Group
Family
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Clethraceae
Combretaceae
Convolvulaceae
Convolvulaceae
Convolvulaceae
Convolvulaceae
Convolvulaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Convolvulaceae
Convolvulaceae
Convolvulaceae
Cornaceae
Crassulaceae
Crassulaceae
Crassulaceae
Crassulaceae
Crassulaceae
Crassulaceae
Crassulaceae
Crassulaceae
Dilleniaceae
Dipsacaceae
Elaeocarpaceae
Ericaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Scientific name
Literature source
Clethra lanata M.Martens & Galeotti
Terminalia melanocarpa F.Muell.
Argyreia nervosa (Burm.f.) Bojer
Convolvulus erubescens Sims
Dichondra micrantha Urb.
Evolvulus alsinoides (L.) L.
Ipomoea carnea Jacq. subsp. fistulosa (Mart. ex Choisy)
D.F.Austin
Merremia peltata (L.) Merr.
Merremia umbellata (L.) Hallier f.
Poranopsis paniculata (Roxb.) Roberty
Cornus kousa Buerg. ex Hance
Crassula multicava Lem.
Crassula ovata (Mill.) Druce
Kalanchoe beharensis Drake
Kalanchoe crenata (Andrews) Haw.
Kalanchoe daigremontiana Raym.-Hamet & H.Perrier
Kalanchoe fedtschenkoi Raym.-Hamet & H.Perrier
Kalanchoe lateritia Engl.
Kalanchoe laxiflora Baker
Dillenia suffruticosa (Griff.) Martelli
Scabiosa palaestrina L.
Elaeocarpus angustifolius Blume
Erica lusitanica Rudolph
Aleurites montana (Lour.) E.H.Wilson
Breynia disticha J.R.Forst. & G.Forst. cv. ‘Roseo-picta’
Croton guatemalensis Lotsy
Euphorbia lactea Haw.
Euphorbia leucocephala Lotsy
Euphorbia serpens Kunth
Euphorbia tirucalli L.
Flueggea virosa (Roxb. ex Willd.) Voigt
Homalanthus populifolius Graham
Homalanthus populneus (Geiseler) Pax
Jatropha multifida L.
Jatropha podagrica Hook.
Manihot esculenta Crantz
Pedilanthus tithymaloides (L.) Poit. subsp. padifolius (L.)
Dressler
Frohlich & Lau 2012:33 (NEWNAT/K)
Parker & Parsons 2012a:58 (NEWNAT/H)
Lorence et al. 1995:32 (NEWNAT/K)
Herbst 1998:2 (NEWNAT/O)
Oppenheimer & Bartlett 2002:6 (NEWNAT/O, EM, WM)
Oppenheimer 2004:10 (NEWNAT/WM)
Lau & Frohlich 2012:13 (NEWNAT/O)
Wood 2012:93 (NEWNAT/K)
Imada et al. 2000:11 (NEWNAT/O)
Starr et al. 2004:21 (NEWNAT/EM)
Pratt & Bio 2012:75 (NEWNAT/H)
Lorence et al. 1995:33 (NEWNAT/K)
Lorence et al. 1995:32 (NEWNAT/K)
Lau & Frohlich 2012:14 (NEWNAT/O)
Herbst et al. 2004:6 (NEWNAT/O)
Lorence et al. 1995:33 (NEWNAT/K)
Lorence et al. 1995:34 (NEWNAT/K)
Kraus 2003:76 (NEWNAT/O)
Frohlich & Lau 2012:34 (NEWNAT/O [as Bryophyllum laxiflorum])
Sato 1998 (NEWNAT/O))
Herbst & Wagner 1996:10 (NEWNAT/H)
Daehler & Baker 2006:7 (NEWNAT/O)
Starr et al. 2008:45 (NEWNAT/EM)
Herbst 1998:2 (NEWNAT/H)
Lorence et al. 1995:35 (NEWNAT/EM)
Starr et al. 2004:22 (NEWNAT/EM)
Frohlich & Lau 2012:35 (NEWNAT/K, O)
Frohlich & Lau 2012:35 (NEWNAT/K)
Wagner et al. 1997:56 (NEWNAT/K [as Chamaesyce serpens])
Lorence et al. 1995:35 (NEWNAT/K)
Starr et al. 2004:22 (NEWNAT/EM)
Oppenheimer 2004:11 (NEWNAT/EM, H [as Omalanthus populifolius])
Frohlich & Lau 2012:35 (NEWNAT/O)
Parker & Parsons 2012a:59 (NEWNAT/H)
Parker & Parsons 2012a:59 (NEWNAT/H)
Frohlich & Lau 2012:36 (NEWNAT/O)
Frohlich & Lau 2012:37 (NEWNAT/O)
G-4
New Naturalized Taxa
Group
Family
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Dicot
Dicot
Dicot
Fabaceae
Fabaceae
Fabaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Scientific name
Phyllanthus acidus (L.) Skeels
Phyllanthus reticulatus Poir.
*Sauropus androgynus (L.) Merr.
Acacia aneura F.Muell. ex Benth.
*Acacia auriculiformis A.Cunn. ex Benth.
Acacia catechu (L.f.) Willd.
Acacia mangium Willd.
Acacia melanoxylon R.Br. ex Aiton
*Acacia retinodes Schltdl.
Adenanthera pavonina L.
Aeschynomene paniculata Willd. ex Vogel
Albizia niopoides (Benth.) Burkart
Albizia saponaria (Lour.) Blume ex Miq.
Bauhinia glauca (Benth.) Benth. subsp. tenuiflora
(C.B.Clarke) K.Larsen & S.S.Larsen
Bauhinia monandra Kurz
Caesalpinia crista L.
Calliandra houstoniana (Mill.) Standl. var. calothyrsa
(Meisn.) Barneby
Calliandra surinamensis Benth.
Calopogonium galactioides (Kunth) Benth. ex Hemsl.
Cassia javanica L. var. indochinensis Gagnep.
Centrosema molle Mart. ex Benth.
Clitoria heterophylla Lam.
Crotalaria juncea L.
Crotalaria sagittalis L.
Crotalaria trichotoma Bojer
Crotalaria verrucosa L.
Dalea emarginata (Torr. & A.Gray) Shinners
Derris elliptica (Wall.) Benth.
Desmodium barbatum (L.) Benth. & Oerst.
Desmodium heterophyllum (Willd.) DC.
Enterolobium cyclocarpum (Jacq.) Griseb.
Erythrina crista-galli L.
Erythrina poeppigiana (Walp.) O.F.Cook
Flemingia macrophylla (Willd.) Merr.
Flemingia strobilifera (L.) Aiton & W.T.Aiton
Gliricidia sepium (Jacq.) Walp.
Glycine microphylla Tindale
Literature source
Parker & Parsons 2012a:59 (NEWNAT/H)
Parker & Parsons 2012b:67 (NEWNAT/H)
Frohlich & Lau 2008:5 (NEWNAT/O); Wagner et al. 2012:35 (O record adventive)
Wagner & Herbst 1995:19 (NEWNAT/L)
Frohlich & Lau 2007:9 (NEWNAT/O); Wagner et al. 2012:36 (O record adventive)
Frohlich & Lau 2010:10 (NEWNAT/O)
Frohlich & Lau 2008:6 (NEWNAT/O)
Lorence et al. 1995:36 (NEWNAT/K)
Starr & Starr 2011:25 (NEWNAT/EM); Wagner et al. 2012:36 (EM record adventive)
Wagner & Herbst 1995:19 (NEWNAT/K)
Hughes 1995:5 (NEWNAT/Mo)
Lau & Frohlich 2012:14 (NEWNAT/O)
Lorence et al. 1995:36 (NEWNAT/K)
Frohlich & Lau 2012:37 (NEWNAT/O)
Lorence & Flynn 1999:4 (NEWNAT/H)
Frohlich & Lau 2008:6 (NEWNAT/O)
Imada et al. 2008:13 (NEWNAT/L)
Starr & Starr 2011:26 (NEWNAT/WM)
Starr et al. 2003:27 (NEWNAT/EM)
Starr & Starr 2011:26 (NEWNAT/EM)
Flynn & Lorence 1998:5 (NEWNAT/K [as Centrosema pubescens])
Staples et al. 2002:9, 10 (NEWNAT/H)
Wagner et al. 1997:56 (NEWNAT/O?, H)
Pratt & Bio 2012:76 (NEWNAT/H)
Starr et al. 2006:35 (NEWNAT/EM)
Wagner & Herbst 1995:19 (NEWNAT/O)
Herbarium Pacificum Staff 1996:4 (NEWNAT/O)
Parker & Parsons 2012a:60 (NEWNAT/H)
Oppenheimer 2010:35 (NEWNAT/Mo)
Wagner & Herbst 1995:20 (NEWNAT/Mo?, EM)
Parker & Parsons 2012b:68 (NEWNAT/H)
Lau & Frohlich 2012:15 (NEWNAT/O); Parker & Parsons 2012b:68 (NEWNAT/H)
Daehler & Baker 2006:7 (NEWNAT/O)
Parker & Parsons 2012b:69 (NEWNAT/H)
Lorence et al. 1995:37 (NEWNAT/EM)
Parker & Parsons 2012b:69 (NEWNAT/H)
Lau & Frohlich 2012:15 (NEWNAT/O)
G-5
New Naturalized Taxa
Group
Family
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Dicot
Fabaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fabaceae
Fagaceae
Flacourtiaceae
Flacourtiaceae
Fumariaceae
Geraniaceae
Geraniaceae
Scientific name
Inga feuillei DC.
Inga sertulifera DC. subsp. leptopus (Benth.) T.D.Penn.
Kummerowia striata (Thunb.) Schindl.
Leucaena diversifolia (Schltdl.) Benth.
Leucaena xspontanea C.E.Hughes & S.A.Harris
Lupinus angustifolius L.
Lupinus arboreus Sims
Macrotyloma axillare (E.Mey.) Verdc. var. glabrum
(E.Mey.) Verdc.
Paraserianthes lophantha (Willd.) I.Nielsen subsp.
montana (Jungh.) I.Nielsen
Peltophorum pterocarpum (DC.) K.Heyne
Piscidia piscipula (L.) Sargent
*Platymiscium stipulare Benth.
Pueraria phaseoloides (Roxb.) Benth.
Senna multijuga (Rich.) H.S.Irwin & Barneby var.
multijuga
Senna siamea (Lam.) H.S.Irwin & Barneby
Senna spectabilis (DC.) H.S.Irwin & Barneby
Sophora tomentosa L.
Stylosanthes guianensis (Aubl.) Sw. var. guianensis
Stylosanthes guianensis (Aubl.) Sw. var. intermedia
(Vogel) Hassler
Stylosanthes viscosa Sw.
Tamarindus indica L.
Tephrosia candida Roxb. ex DC.
Trifolium campestre Schreb.
Vigna hosei (Craib) Backer
Vigna luteola (Jacq.) Benth.
Vigna speciosa (Kunth) Verdc.
Vigna vexillata (L.) A.Rich.
Zornia cf. reticulata Sm.
Zornia gemella (Willd.) Vogel
Quercus suber L.
Dovyalis hebecarpa (Gardner) Warb.
Flacourtia indica (N.L.Burm.) Merr.
Fumaria officinalis L.
Erodium sp.
Pelargonium capitatum (L.) L'Hér. ex Aiton
Literature source
Parker & Parsons 2012b:69 (NEWNAT/H)
Lorence & Flynn 2006:2 (NEWNAT/K)
Oppenheimer 2003:13 (NEWNAT/WM)
Starr et al. 2008:45 (NEWNAT/EM)
Oppenheimer 2004:12 (NEWNAT/EM)
Pratt & Bio 2012:76 (NEWNAT/H)
Staples et al. 2002:10 (NEWNAT/H)
Staples et al. 2003:11 (NEWNAT/O)
Oppenheimer & Bartlett 2002:8 (NEWNAT/EM)
Frohlich & Lau 2012:38 (NEWNAT/K)
Lau & Frohlich 2012:16 (NEWNAT/O)
Frohlich & Lau 2008:6 (NEWNAT/O); Wagner et al. 2012:41 (O record adventive)
Frohlich & Lau 2008:7 (NEWNAT/O)
Herbarium Pacificum Staff 1998:10 (NEWNAT/K)
Lau & Frohlich 2012:16 (NEWNAT/O)
Lau & Frohlich 2012:17 (NEWNAT/O)
Pratt & Bio 2012:77 (NEWNAT/H)
Herbst et al. 2004:7, 8 (NEWNAT/K, Mo?)
Herbst et al. 2004:7, 8 (NEWNAT/Mo?)
Herbst et al. 2004:7 (NEWNAT/O, WM)
Wysong et al. 2007:4 (NEWNAT/Mo)
Oppenheimer & Bartlett 2000:5 (NEWNAT/WM)
Starr & Starr 2011:27 (NEWNAT/EM)
Staples et al. 2006:7 (NEWNAT/O)
Staples et al. 2003:12 (NEWNAT/O)
Herbst 1998:3 (NEWNAT/O)
Wagner et al. 2012:43 (NEWNAT/K)
Starr et al. 2006:35 (NEWNAT/EM)
Staples et al. 2006:7 (NEWNAT/O)
Herbst & Wagner 1996:11 (NEWNAT/H)
Herbst 1998:3 (NEWNAT/H)
Lau & Frohlich 2012:17 (NEWNAT/O)
Starr & Starr 2011:27 (NEWNAT/EM)
Hughes 1995:6 (NEWNAT/Mo)
Starr et al. 2008:46 (NEWNAT/EM)
G-6
New Naturalized Taxa
Group
Family
Dicot
Dicot
Grossulariaceae
Grossulariaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Hydrangeaceae
Lamiaceae
Lamiaceae
Lamiaceae
Lamiaceae
Lamiaceae
Lamiaceae
Lythraceae
Magnoliaceae
Malpighiaceae
Malvaceae
Malvaceae
Malvaceae
Malvaceae
Melastomataceae
Melastomataceae
Melastomataceae
Melastomataceae
Melastomataceae
Meliaceae
Meliaceae
Meliaceae
Meliaceae
Meliaceae
Moraceae
Moraceae
Moraceae
Moraceae
Moraceae
Moraceae
Moraceae
Myricaceae
Myrtaceae
Myrtaceae
Myrtaceae
Myrtaceae
Scientific name
Brexia madagascariensis (Lam.) Thouars ex Ker Gawl.
Escallonia rubra (Ruiz & Pav.) Pers. var. macrantha
(Hook. & Arn.) Reiche
Philadelphus karwinskyanus Koehne
Hyptis capitata Jacq.
Hyptis verticillata Jacq.
Plectranthus cylindraceus Hochst. ex Benth.
Plectranthus neochilus Schltr.
Plectranthus prostratus Gürke
Plectranthus verticillatus (L.f.) Druce
Cuphea subuligera Koehne
Liriodendron tulipifera L.
Hiptage benghalensis (L.) Kurz
Bastardia viscosa (L.) Kunth
*Hibiscus makinoi Jotani & Ohba
Malva neglecta Wallr.
Sida ciliaris L.
Medinilla cumingii Naudin
Medinilla heterophylla A.Gray
Medinilla magnifica Lindl.
Miconia calvescens DC.
Tibouchina granulosa (Desr.) Cogn.
Azadirachta indica A.Juss.
Cedrela odorata L.
Khaya nyasica Stapf ex Baker f.
Sandoricum koetjape (Burm.f.) Merr.
Swietenia macrophylla King
Artocarpus heterophyllus Lam.
Fatoua villosa (Thunb.) Nakai
Ficus cf. platypoda (A.Cunn. ex Miq.) A.Cunn. ex Miq.
Ficus macrophylla Desf. ex Pers.
Ficus nota (Blanco) Merr.
*Ficus religiosa L.
Ficus rubiginosa Desf. ex Vent.
Morella cerifera (L.) Small
Angophora costata (Gaertn.) Britten
Eucalyptus punctata DC.
Eugenia brasiliensis Lam.
Leptospermum morrisonii Joy Thomps.
Literature source
Starr et al. 2003:28 ( (NEWNAT/EM)
Parker & Parsons 2012a:60 (NEWNAT/H)
Lorence et al. 1995:38 (NEWNAT/K, EM)
Wagner & Herbst 1995:21 (NEWNAT/O)
Starr & Starr 2011:28 (NEWNAT/EM)
Parker & Parsons 2010:42 (NEWNAT/H)
Lau & Frohlich 2012:18 (NEWNAT/O)
Staples et al. 2002:11 (NEWNAT/O)
Starr et al. 2004:23 (NEWNAT/O, EM)
Parker & Parsons 2012a:60 (NEWNAT/H)
Parker & Parsons 2012b:70 (NEWNAT/H)
Daehler & Baker 2006:8 (NEWNAT/O)
Staples et al. 2002:12 (NEWNAT/H)
Frohlich & Lau 2008:7 (NEWNAT/O); Wagner et al. 2012:49 (O record adventive)
Herbarium Pacificum Staff 1998:10 (NEWNAT/EM)
Wagner et al. 1997:59 (NEWNAT/O)
Oppenheimer 2004:13 (NEWNAT/EM)
Frohlich & Lau 2010:12 (NEWNAT/O)
Frohlich & Lau 2007:10 (NEWNAT/O)
Conant 1996:1 (NEWNAT/K, O)
Lau & Frohlich 2012:18 (NEWNAT/O)
Lau & Frohlich 2012:19 (NEWNAT/O)
Starr et al. 2006:37 (NEWNAT/EM)
Daehler & Baker 2006:9 (NEWNAT/O)
Starr et al. 2004:24 (NEWNAT/EM, WM)
Meidell et al. 1998:6 (NEWNAT/O, WM)
Lau & Frohlich 2012:19 (NEWNAT/O); Parker & Parsons 2012b:70 (NEWNAT/H)
Wagner & Herbst 1995:22 (NEWNAT/O)
Oppenheimer & Bartlett 2000:7 (NEWNAT/O, EM, WM)
Oppenheimer & Bartlett 2000:6 (NEWNAT/WM)
Herbarium Pacificum Staff 1998:10 (NEWNAT/O, H)
Frohlich & Lau 2008:7 (NEWNAT/O); Wagner et al. 2012:52 (O record adventive)
Oppenheimer 2003:15 (NEWNAT/WM)
Meidell et al. 1997:17 (NEWNAT/WM [as Myrica cerifera])
Starr et al. 2006:37 (NEWNAT/EM)
Oppenheimer 2004:14 (NEWNAT/WM)
Starr & Starr 2011:29 (NEWNAT/EM)
Herbarium Pacificum Staff 1999:5 (NEWNAT/EM)
G-7
New Naturalized Taxa
Group
Family
Scientific name
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Myrtaceae
Myrtaceae
Myrtaceae
Myrtaceae
Myrtaceae
Myrtaceae
Myrtaceae
Myrtaceae
Nymphaeaceae
Nymphaeaceae
Leptospermum petersonii F.M.Bailey
Melaleuca armillaris (Sol. ex Gaertn.) Sm.
Metrosideros kermadecensis W.R.B.Oliv.
Pimenta dioica (L.) Merr.
Pimenta racemosa (Mill.) J.W.Moore
Syncarpia glomulifera (Sm.) Nied.
Syzygium grande (Wight) Walp.
Syzygium syzygioides (Miq.) Merr. & L.M.Perry
Nymphaea caerulea Savigny
Nymphaea capensis Thunb.
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Nymphaeaceae
Nymphaeaceae
Ochnaceae
Ochnaceae
Oleaceae
Oleaceae
Oleaceae
Oleaceae
Oleaceae
Oleaceae
Oleaceae
Onagraceae
Onagraceae
Onagraceae
Nymphaea xdaubenyana W.T.Baxter ex Daubeny
Nymphaea lotus L.
Ochna serrulata (Hochst.) Walp.
Ochna thomasiana Engl. & Gilg
Fraxinus griffithii C.B.Clarke
Jasminum fluminense Vell.
Jasminum multiflorum (Burm.f.) Andrews
Jasminum polyanthum Franch.
Ligustrum lucidum W.T.Aiton
Ligustrum sinense Lour.
Noronhia emarginata (Lam.) Poir.
Oenothera biennis L.
Oenothera kunthiana (Spach) Munz
*Oenothera speciosa Nutt.
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Passifloraceae
Phytolaccaceae
Piperaceae
Piperaceae
Piperaceae
Pittosporaceae
Plumbaginaceae
Polemoniaceae
Polygalaceae
Polygonaceae
Polygonaceae
Polygonaceae
Polygonaceae
Passiflora maliformis L.
Phytolacca dioica L.
Piper aduncum L.
Piper auritum Kunth
Piper betle L.
Pittosporum pentandrum (Blanco) Merr.
Plumbago auriculata Lam.
Collomia linearis Nutt.
Polygala virgata Thunb.
Coccoloba uvifera (L.) L.
Emex australis Steinh.
Rumex conglomeratus Murray
Rumex crispus L. subsp. fauriei (Rech.) Mosyakin &
Literature source
Herbarium Pacificum Staff 1999:5 (NEWNAT/H)
Starr & Starr 2012:89 (NEWNAT/EM)
Starr et al. 2004:25 (NEWNAT/EM)
Lorence et al. 1995:42 (NEWNAT/K)
Frohlich & Lau 2012:41 (NEWNAT/O)
Oppenheimer 2003:16 (NEWNAT/WM)
Daehler & Baker 2006:9 (NEWNAT/O)
Daehler & Baker 2006:9 (NEWNAT/O)
Herbarium Pacificum Staff 1998:11 (NEWNAT/H)
Herbarium Pacificum Staff 1998:11 (NEWNAT/K [as Nymphaea capensis var.
zanzibariensis])
Oppenheimer 2003:17 (NEWNAT/WM)
Staples et al. 2003:16 (NEWNAT/O)
Herbarium Pacificum Staff 1998:12 (NEWNAT/H)
Imada et al. 2000:13 (NEWNAT/O)
Oppenheimer 2003:17 (NEWNAT/EM)
Herbarium Pacificum Staff 1996:5 (NEWNAT/O, H)
Parker & Parsons 2012b:71 (NEWNAT/H)
Imada 2007:38 (NEWNAT/H)
Staples et al. 2003:16 (NEWNAT/H)
Lorence & Flynn 1999:5 (NEWNAT/K)
Flynn & Lorence 2002:15 (NEWNAT/K)
Starr et al. 2008:47 (NEWNAT/EM)
Starr et al. 2004:25 (NEWNAT/EM)
Starr & Starr 2011:29 (NEWNAT/EM); Wagner et al. 2012:56 (not naturalized; probably
persisting & spreading only by rhizomes)
Lorence et al. 1995:43 (NEWNAT/K)
Frohlich & Lau 2012:43 (NEWNAT/O)
Starr et al. 2004:25 (NEWNAT/EM)
Staples et al. 2006:8 (NEWNAT/ K, O)
Frohlich & Lau 2008:8 (NEWNAT/O)
Herbarium Pacificum Staff 1999:6 (NEWNAT/O, H)
Herbarium Pacificum Staff 1999:7 (NEWNAT/EM)
Wagner & Herbst 1995:23 (NEWNAT/EM)
Starr & Starr 2011:31 (NEWNAT/EM)
Herbst 1998:3 (NEWNAT/O)
Parker & Parsons 2012b:71 (NEWNAT/H)
Wagner et al. 1997:60 (NEWNAT/K)
Mosyakin & Wagner 1998:40 (NEWNAT/O?, L)
G-8
New Naturalized Taxa
Group
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Family
Polygonaceae
Proteaceae
Proteaceae
Proteaceae
Rhamnaceae
Rhamnaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rosaceae
Rubiaceae
Rubiaceae
Rubiaceae
Rubiaceae
Rubiaceae
Rubiaceae
Rubiaceae
Rubiaceae
Rubiaceae
Rubiaceae
Rutaceae
Rutaceae
Rutaceae
Rutaceae
Santalaceae
Sapindaceae
Sapindaceae
Sapindaceae
Sapindaceae
Scientific name
W.L.Wagner
Triplaris weigeltiana (Rchb.) Kuntze
Banksia integrifolia L.f.
Macadamia integrifolia Maiden & Betche
Macadamia tetraphylla L.A.S.Johnson
Alphitonia excelsa (Fenzl) Reissek ex Benth.
*Ziziphus mauritiana Lam.
Cotoneaster pannosus Franch.
Eriobotrya japonica (Thunb.) Lindl.
Heteromeles arbutifolia (Lindl.) M.Roem.
Photinia davidiana (Decne.) Cardot
Prunus campanulata Maxim.
Prunus grisea (C.Müll.) Kalkman
Prunus persica (L.) Batsch
Pyracantha crenatoserrata (Hance) Rehder
Pyracantha koidzumii Rehder
Rosa xdamascena Mill.
Rosa laevigata Michx.
Rubus discolor Weihe & Nees
Rubus glaucus Benth.
Atractocarpus fitzalanii (F.Muell.) Puttock
Coffea liberica Hiern
Diodia apiculata (Roem. & Schult.) K.Schum.
Galium aparine L.
Pentas lanceolata (Forssk.) Deflers
Richardia scabra L.
Serissa japonica (Thunb.) Thunb.
Spermacoce exilis (L.O.Williams) C.D.Adams
Spermacoce latifolia Aubl.
Spermacoce prostrata Aubl.
Melicope elleryana (F.Muell.) T.G.Hartley
Murraya paniculata (L.) Jack
Ruta graveolens L.
Triphasia trifolia (Burm.f.) P.Wilson
Santalum album L.
*Allophyllus cobbe (L.) Raeusch.
Cupaniopsis anacardioides (A.Rich.) Radlk.
Filicium decipiens (Wight & Arn.) Thwaites
Koelreuteria elegans (Seem.) A.C.Sm. subsp. formosana
Literature source
Lau & Frohlich 2012:20 (NEWNAT/O)
Starr et al. 2006:40 (NEWNAT/EM)
Oppenheimer 2010:38 (NEWNAT/Mo)
Oppenheimer 2004:16 (NEWNAT/EM)
Daehler & Baker 2006:10 (NEWNAT/O)
Frohlich & Lau 2010:15 (NEWNAT/O); Wagner et al. 2012:61 (O record adventive)
Lorence et al. 1995:49 (NEWNAT/K)
Lorence et al. 1995:49 (NEWNAT/K)
Herbst & Wagner 1996:12 (NEWNAT/H)
Herbarium Pacificum Staff 1996:6 (NEWNAT/EM?, H)
Lorence & Flynn 2006:4 (NEWNAT/K)
Daehler & Baker 2006:11 (NEWNAT/O)
Oppenheimer 2003:22 (NEWNAT/EM)
Herbarium Pacificum Staff 1999:9 (NEWNAT/K, H)
Herbarium Pacificum Staff 1999:9 (NEWNAT/K, H)
Staples et al. 2002:14 (NEWNAT/Mo, EM, H)
Nagata 1995:12 (NEWNAT/L)
Conant 1996:1 (NEWNAT/O)
Starr et al. 2003:31 (NEWNAT/EM)
Daehler & Baker 2006:11 (NEWNAT/O [as Randia fitzalanii])
Staples et al. 2002:14 (NEWNAT/O)
Imada et al. 2000:15 (NEWNAT/O)
Starr & Starr 2011:31 (NEWNAT/EM)
Nagata 1995:12 (NEWNAT/H)
Lorence et al. 1995:50 (NEWNAT/K)
Wagner et al. 1997:61 (NEWNAT/H)
Wagner et al. 1997:61 (NEWNAT/H [as Spermacoce capitata])
Lorence et al. 1995:51 (NEWNAT/K)
Lorence et al. 1995:52 (NEWNAT/H [as Spermacoce ovalifolia])
Frohlich & Lau 2012:46 (NEWNAT/O)
Kraus 2003:77 (NEWNAT/O)
Starr et al. 2004:27 (NEWNAT/EM)
Lau & Frohlich 2012:22 (NEWNAT/O)
Frohlich & Lau 2008:9 (NEWNAT/O)
Frohlich & Lau 2008:9 (NEWNAT/O); Wagner et al. 2012:69 (O record adventive)
Frohlich & Lau 2010:15 (NEWNAT/O)
Staples et al. 2002:15 (NEWNAT/O)
Oppenheimer 2003:25 (NEWNAT/WM)
G-9
New Naturalized Taxa
Group
Family
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Sapindaceae
Sapotaceae
Sapotaceae
Scrophulariaceae
Scrophulariaceae
Scrophulariaceae
Scrophulariaceae
Scrophulariaceae
Scrophulariaceae
Simaroubaceae
Solanaceae
Solanaceae
Solanaceae
Solanaceae
Solanaceae
Sterculiaceae
Sterculiaceae
Sterculiaceae
Theaceae
Tiliaceae
Urticaceae
Urticaceae
Verbenaceae
Verbenaceae
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Dicot
Verbenaceae
Verbenaceae
Verbenaceae
Verbenaceae
Verbenaceae
Verbenaceae
Violaceae
Vitaceae
Vitaceae
Vitaceae
Vitaceae
Vitaceae
Vitaceae
Scientific name
(Hayata) F.G.Mey.
Majidea zanquebarica J.Kirk ex Oliv.
Chrysophyllum oliviforme L.
Sideroxylon persimile (Hemsl.) T.D.Penn.
Hebe speciosa (A.Cunn.) Cockayne & Allan
Lindernia procumbens (Krock.) Philcox
Mazus pumilus (Burm.f.) Steenis
Otacanthus azureus (Linden) Ronse
Paulownia tomentosa (Thunb.) Steud.
Torenia glabra Osbeck
Ailanthus altissima (Mill.) Swingle
Physalis ixocarpa Brot. ex Hornem.
Physalis lanceifolia Nees
Solanum melongena L.
Solanum villosum Mill.
Streptosolen jamesonii (Benth.) Miers
Heritiera littoralis Dryand.
Sterculia apetala (Jacq.) H.Karst.
Sterculia foetida L.
Camellia sinensis (L.) Kuntze
Grewia micrantha Bojer
Boehmeria nivea (L.) Gaudich.
*Procris pedunculata (J.R.Forst. & G.Forst.) Wedd.
Citharexylum spinosum L.
Clerodendrum buchananii (Roxb.) Walp. var. fallax
(Lindl.) Bakh.
Clerodendrum glabrum E.Mey.
Clerodendrum macrostegium Schauer
Duranta erecta L.
Phyla nodiflora (L.) Greene
Vitex parviflora Juss.
Vitex trifolia L.
Viola odorata L.
Cissus quadrangularis L.
Cissus repens Lam.
Cissus rhombifolia Vahl
Cissus rotundifolia (Forssk.) Vahl
Cissus verticillata (L.) Nicolson & C.E.Jarvis
Tetrastigma pubinerve Merr. & Chun
Literature source
Parker & Parsons 2012a:61 (NEWNAT/H)
Lorence et al. 1995:54 (NEWNAT/K)
Starr et al. 2002:24 (NEWNAT/O, L?, EM)
Shannon & Wagner 1996:14 (NEWNAT/H)
Lorence et al. 1995:54 (NEWNAT/K)
Oppenheimer 2004:17 (NEWNAT/EM)
Parker & Parsons 2012a:61 (NEWNAT/H)
Parker & Parsons 2012a:61 (NEWNAT/H)
Staples et al. 2002:15, 16 (NEWNAT/O, H)
Frohlich & Lau 2012:47 (NEWNAT/K)
Nagata 1995: 13 (NEWNAT/Mo)
Wagner & Herbst 1995:26 (NEWNAT/EM)
Frohlich & Lau 2012:47 (NEWNAT/O)
Imada et al. 2008:14 (NEWNAT/H)
Lorence & Flynn 1997:12 (NEWNAT/K)
Lorence & Flynn 2006:3 (NEWNAT/K)
Oppenheimer 2003:26 (NEWNAT/WM)
Frohlich & Lau 2010:16 (NEWNAT/O)
Lorence & Flynn 2006:5 (NEWNAT/K)
Lau & Frohlich 2012:22 (NEWNAT/O)
Parker & Parsons 2012a:62 (NEWNAT/H)
Frohlich & Lau 2008:9 (NEWNAT/O); Wagner et al. 2012:72 (O record adventive)
Herbarium Pacificum Staff 1998:13 (NEWNAT/O)
Starr et al. 2006:41 (NEWNAT/EM)
Frohlich & Lau 2012:48 (NEWNAT/K)
Lorence & Flynn 1997:12 (NEWNAT/K)
Lorence et al. 1995:55 (NEWNAT/K)
Starr et al. 2002:26 (NEWNAT/WM)
Frohlich & Lau 2008:10 (NEWNAT/O)
Wysong et al. 2007:7 (NEWNAT/Mo)
Shannon & Wagner 1996:15 (NEWNAT/H)
Frohlich & Lau 2008:10 (NEWNAT/O)
Lau & Frohlich 2012:23 (NEWNAT/O)
Starr et al. 2004:28 (NEWNAT/WM)
Lorence & Flynn 1997:12 (NEWNAT/K)
Frohlich & Lau 2012:48 (NEWNAT/K, O)
Oppenheimer & Bartlett 2000:9 (NEWNAT/WM)
G-10
New Naturalized Taxa
Group
Family
Dicot
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Zygophyllaceae
Araucariaceae
Cupressaceae
Cupressaceae
Cupressaceae
Cupressaceae
Cupressaceae
Pinaceae
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Gymnosperm
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Pinaceae
Pinaceae
Pinaceae
Pinaceae
Podocarpaceae
Podocarpaceae
Podocarpaceae
Podocarpaceae
Taxodiaceae
Agavaceae
Aloeaceae
Araceae
Araceae
Araceae
Araceae
Araceae
Araceae
Araceae
Araceae
Araceae
Araceae
Arecaceae
Arecaceae
Arecaceae
Arecaceae
Arecaceae
Arecaceae
Arecaceae
Arecaceae
Arecaceae
Scientific name
Literature source
Guaiacum officinale L.
Araucaria columnaris (G.Forst.) Hook.
Callitris columellaris F.Muell.
Callitris endlicheri (Parl.) F.M.Bailey
Cupressus lusitanica Mill.
Cupressus macrocarpa Hartw.
Juniperus bermudiana L.
Pinus caribaea Morelet var. hondurensis (Sénécl.)
W.H.Barrett & Golfari
Pinus elliottii Engelm.
Pinus patula Schiede ex Schltdl. & Cham.
Pinus radiata D.Don
Pinus taeda L.
Dacrycarpus imbricatus (Blume) de Laub.
Nageia falcata (Thunb.) Carrière
Podocarpus elatus R.Br. ex Endl.
Podocarpus macrophyllus (Thunb.) Sweet
Cryptomeria japonica (Thunb. ex L.f.) D.Don
Sansevieria trifasciata Prain
Aloe vera (L.) Burm.f.
Aglaonema commutatum Schott
Alocasia cucullata (Lour.) G.Don
Caladium bicolor (Aiton) Vent.
Dieffenbachia maculata (Loddiges) G.Don
Philodendron erubescens K.Koch & Augustin
Philodendron lacerum (Jacq.) Schott
Philodendron pinnatifidum (Jacq.) Schott
Philodendron scandens K.Koch & Sello
Syngonium podophyllum Schott
Xanthosoma robustum Schott
Arenga pinnata (Wurmb) Merr.
Caryota mitis Lour.
Coccothrinax barbadensis (Lodd. ex Mart.) Becc.
Licuala spinosa Wurmb
Phoenix hybrid
Pinanga coronata (Blume ex Mart.) Blume
Ptychosperma macarthurii (Veitch) H.Wendl. ex Hook.f.
Roystonea regia (Kunth) O.F.Cook
Syagrus romanzoffiana (Cham.) Glassman
Lau & Frohlich 2012:24 (NEWNAT/O)
Oppenheimer 2002:20 (NEWNAT/WM)
Oppenheimer 2002:20 (NEWNAT/WM)
Lau & Frohlich 2012:14 (NEWNAT/O)
Frohlich & Lau 2012:34 (NEWNAT/O)
Starr et al. 2008:44 (NEWNAT/EM)
Oppenheimer 2002:20, 21 (NEWNAT/WM)
Oppenheimer 2003:19 (NEWNAT/EM, WM)
Wysong et al. 2007:6 (NEWNAT/Mo)
Oppenheimer 2002:20, 21 (NEWNAT/EM)
Oppenheimer 2002:20, 21 (NEWNAT/EM)
Oppenheimer 2008:31 (NEWNAT/L)
Daehler & Baker 2006:10 (NEWNAT/O)
Oppenheimer 2002:20, 22 (NEWNAT/EM)
Lau & Frohlich 2012:20 (NEWNAT/O)
Lau & Frohlich 2012:20 (NEWNAT/O)
Oppenheimer 2002:20, 22 (NEWNAT/WM)
Imada et al. 2000:9 (NEWNAT/O)
Lorence et al. 1995:21 (NEWNAT/K)
Imada 2007:34 (NEWNAT/O)
Staples et al. 2003:8 (NEWNAT/EM, H)
Resslar 2010 (NEWNAT/H)
Lorence & Flynn 2006:2 (NEWNAT/K)
Flynn & Lorence 2002:14 (NEWNAT/K)
Oppenheimer 2004:9 (NEWNAT/EM)
Flynn & Lorence 2002:15 (NEWNAT/K)
Oppenheimer 2008:23 (NEWNAT/K, EM)
Oppenheimer 2006:10 (NEWNAT/WM)
Staples & Woolliams 1997:13 (NEWNAT/K [as Xanthosoma roseum])
Daehler & Baker 2006:5 (NEWNAT/O)
Daehler & Baker 2006:5 (NEWNAT/O)
Frohlich & Lau 2012:29 (NEWNAT/K, O)
Lau & Frohlich 2012:8 (NEWNAT/O)
Oppenheimer 2003:6 (NEWNAT/EM, Mo?)
Daehler & Baker 2006:5 (NEWNAT/O)
Daehler & Baker 2006:6 (NEWNAT/O)
Frohlich & Lau 2007:9 (NEWNAT/O)
Daehler & Baker 2006:6 (NEWNAT/O)
G-11
New Naturalized Taxa
Group
Family
Monocot
Arecaceae
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Arecaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Bromeliaceae
Commelinaceae
Commelinaceae
Commelinaceae
Commelinaceae
Commelinaceae
Costaceae
Costaceae
Costaceae
Costaceae
Cyperaceae
Cyperaceae
Cyperaceae
Cyperaceae
Cyperaceae
Cyperaceae
Cyperaceae
Cyperaceae
Cyperaceae
Cyperaceae
Cyperaceae
Cyperaceae
Scientific name
Literature source
Washingtonia filifera (Linden ex André) H.Wendl. ex de
Bary
Washingtonia robusta H.Wendl.
*Aechmea bracteata (Sw.) Griseb.
Guzmania lindenii (André) Mez
Guzmania monostachia (L.) Rusby ex Mez
Tillandsia balbisiana Schult.f.
Tillandsia bulbosa Hook.
Tillandsia caput-medusae E.Morren
Tillandsia fasciculata Sw.
Tillandsia gardneri Lindl.
*Tillandsia juncea (Ruiz & Pav.) Poir.
Tillandsia polystachia (L.) L.
Tillandsia recurvata (L.) L.
Tillandsia stricta Sol.
*Tillandsia usneoides (L.) L.
Werauhia sanguinolenta (Cogn. & Marchal) J.R.Grant
Callisia repens L.
Palisota bracteosa C.B.Clarke
Tradescantia fluminensis Vell.
Tradescantia spathacea Sw.
Tradescantia zebrina Hort. ex Bosse
Costus dubius (Afzel.) K.Schum.
Costus pulverulentus C.Presl
Costus scaber Ruiz & Pav.
Costus woodsonii Maas
Carex inversa R.Br.
Carex longii Mack.
Cyperus confertus Sw.
Cyperus cyperoides (L.) Kuntze
Cyperus eragrostis Lam.
Cyperus prolifer Lam.
Cyperus sphacelatus Rottb.
Eleocharis olivacea Torr.
Eleocharis schaffneri Boeck.
Fimbristylis ferruginea (L.) Vahl
Fimbristylis littoralis Gaudich.
Rhynchospora radicans (Schltdl. & Cham.) H.Pfeiff.
subsp. microcephala (Bertero ex Spreng.) W.W.Thomas
Oppenheimer & Bartlett 2002:3 (NEWNAT/WM)
Oppenheimer & Bartlett 2002:3 (NEWNAT/WM)
Frohlich & Lau 2008:4 (NEWNAT/O); Wagner et al. 2012:77 (O record adventive)
Frohlich & Lau 2010:6 (NEWNAT/O)
Frohlich & Lau 2010:7 (NEWNAT/O)
Frohlich & Lau 2010:7 (NEWNAT/O)
Frohlich & Lau 2010:7 (NEWNAT/O)
Oppenheimer 2006:11 (NEWNAT/EM)
Frohlich & Lau 2010:8 (NEWNAT/O)
Frohlich & Lau 2010:8 (NEWNAT/O)
Frohlich & Lau 2010:8 (NEWNAT/O); Wagner et al. 2012:78 (O record adventive)
Frohlich & Lau 2010:9 (NEWNAT/O)
Frohlich & Lau 2010:9 (NEWNAT/O)
Frohlich & Lau 2010:10 (NEWNAT/O)
Frohlich & Lau 2007:9 (NEWNAT/O); Wagner et al. 2012:78 (O record adventive)
Lau & Frohlich 2012:11 (NEWNAT/O)
Frohlich & Lau 2010:10 (NEWNAT/O)
Herbst et al. 2004:5 (NEWNAT/O)
Staples et al. 2006:6 (NEWNAT/EM, H)
Frohlich & Lau 2010:10 (NEWNAT/O)
Lorence & Flynn 1997:10 (NEWNAT/K)
Daehler & Baker 2006:6 (NEWNAT/O)
Daehler & Baker 2006:6 (NEWNAT/O)
Frohlich & Lau 2012:33 (NEWNAT/K)
Oppenheimer 2004:11 (NEWNAT/EM)
Shaw & Douglas 1995:249 (NEWNAT/H)
Strong & Wagner 1997:37 (NEWNAT/EM, H [as Carex longii])
Herbarium Pacificum Staff 1996:3 (NEWNAT/H)
Herbarium Pacificum Staff 1996:4 (NEWNAT/H)
Staples et al. 2003:10 (NEWNAT/H)
Strong & Wagner 1997:41 (NEWNAT/K)
Oppenheimer 2008:27 (NEWNAT/Mo)
Strong & Wagner 1997:43 (NEWNAT/H)
Strong & Wagner 1997:43 (NEWNAT/K)
Strong & Wagner 1997:44 (NEWNAT/O)
Strong & Wagner 1997:45 (NEWNAT/K [as Fimbristylis miliacea])
Strong & Wagner 1997:46 (NEWNAT/H)
G-12
New Naturalized Taxa
Group
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Family
Cyperaceae
Cyperaceae
Cyperaceae
Hydrocharitaceae
Hydrocharitaceae
Hydrocharitaceae
Iridaceae
Iridaceae
Iridaceae
Iridaceae
Lemnaceae
Lemnaceae
Lemnaceae
Liliaceae
Liliaceae
Liliaceae
Liliaceae
Liliaceae
Liliaceae
Liliaceae
Liliaceae
Marantaceae
Marantaceae
Musaceae
Najadaceae
Najadaceae
Orchidaceae
Orchidaceae
Orchidaceae
Orchidaceae
Orchidaceae
Orchidaceae
Orchidaceae
Orchidaceae
Orchidaceae
Orchidaceae
Orchidaceae
Poaceae
Poaceae
Scientific name
Literature source
Rhynchospora tenuis subsp. tenuis
Schoenoplectus mucronatus (L.) Palla subsp. mucronatus
Schoenus apogon Roem. & Schult.
Halophila decipiens Ostenf.
Vallisneria americana Michx.
Vallisneria spiralis L.
Aristea gerrardii Weim.
Dietes iridioides (L.) Sweet
Gladiolus dalenii Van Geel
Watsonia meriana (L.) Mill.
Landoltia punctata (G.Mey.) Les & D.J.Crawford
Lemna obscura (Austin) Daubs
Wolffia globosa (Roxb.) Hartog & Plas
Asparagus asparagoides (Forssk.) Deflers
Asparagus densiflorus (Kunth) Jessop
*Asparagus falcatus L.
Asparagus plumosus Baker
Dianella caerulea Sims var. assera R.J.F.Hend.
Molineria capitulata (Lour.) Herb.
Zephyranthes citrina Baker
Zephyranthes grandiflora Lindl.
Pleiostachya pruinosa (W.Bull ex Regel) K.Schum.
Stromanthe tonckat (Aubl.) Eichler
Musa velutina H.Wendl. & Drude
Najas guadalupensis (Spreng.) Magnus
Najas marina L.
Cymbidium dayanum Rchb.f.
Dendrobium antennatum Lindl.
Dendrobium crumenatum Sw.
*Dendrobium hybrid
Dendrobium mirbelianum Gaudich.
Dendrobium rhombeum Lindl.
Epidendrum nocturnum Jacq.
Habenaria rodeiensis Barb.Rodr.
Polystachya concreta (Jacq.) Garay & Sweet
Vanda tricolor Lindl.
Zeuxine strateumatica (L.) Schltr.
Agrostis capillaris L.
Agrostis hyemalis (Walter) Britton, Sterns & Poggenb.
Imada 2007:36 (NEWNAT/K)
Strong & Wagner 1997:47 (NEWNAT/H)
Herbarium Pacificum Staff 1996:4 (NEWNAT/H)
McDermid et al. 2002 (NEWNAT/O, H)
Staples et al. 2003:13 (NEWNAT/K)
Staples et al. 2003:13 (NEWNAT/O)
Herbarium Pacificum Staff 1999:4 (NEWNAT/H [as Aristea cf. compressa])
Frohlich & Lau 2012:39 (NEWNAT/O)
Starr et al. 2004:23 (NEWNAT/EM, H)
Oppenheimer 2004:12 (NEWNAT/EM)
Wagner et al. 1997:59 (NEWNAT/O, H [as Spirodela punctata])
Wagner et al. 1997:58 (NEWNAT/O, H)
Wagner et al. 1997:59 (NEWNAT/O, Mo)
Starr et al. 2002:20 (NEWNAT/EM)
Lorence & Flynn 1999:4 (NEWNAT/K)
Frohlich & Lau 2010:11 (NEWNAT/O); Wagner et al. 2012:84 (O record adventive)
Lorence et al. 1995:40 (NEWNAT/K [as Asparagus setaceus])
Lau & Frohlich 2012:18 (NEWNAT/O)
Flynn & Lorence 2002:15 (NEWNAT/K)
Lorence et al. 1995:40 (NEWNAT/K)
Oppenheimer 2011:8 (NEWNAT/L)
Daehler & Baker 2006:8 (NEWNAT/O)
Daehler & Baker 2006:9 (NEWNAT/O)
Daehler & Baker 2006:9 (NEWNAT/O)
Herbst & Wagner 1996:11 (NEWNAT/H)
Shannon & Wagner 1996:14 (NEWNAT/O)
Ackerman et al. 2011:1 (NEWNAT/H)
Frohlich & Lau 2010:13 (NEWNAT/O)
Frohlich & Lau 2010:13 (NEWNAT/O)
Ackerman et al. 2011:2 (NEWNAT/O); Wagner et al. 2012:85 (O record adventive)
Frohlich & Lau 2012:41 (NEWNAT/O)
Frohlich & Lau 2012:42 (NEWNAT/O)
Frohlich & Lau 2012:42 (NEWNAT/O)
Herbst & Wagner 1999:24 (NEWNAT/EM)
Staples et al. 2003:17 (NEWNAT/O)
Oppenheimer 2006:12 (NEWNAT/WM)
Oppenheimer 2003:18 (NEWNAT/EM)
Herbst & Clayton 1998:18 (NEWNAT/EM)
Snow & Lau 2010:46 (NEWNAT/O)
G-13
New Naturalized Taxa
Group
Family
Monocot
Monocot
Poaceae
Poaceae
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Scientific name
Literature source
Andropogon bicornis L.
Andropogon glomeratus (Walter) Britton, Sterns &
Poggenb. var. pumilus (Vasey) L.H.Dewey
Anthephora hermaphrodita (L.) Kuntze
Arrhenatherum elatius (L.) P.Beauv. ex J.Presl & C.Presl
Avena barbata Pott ex Link
Axonopus compressus (Sw.) P.Beauv.
Bothriochloa bladhii (Retz.) S.T.Blake
Bothriochloa laguroides (DC.) Herter
Bromus madritensis L.
Bromus rubens L.
Bromus sterilis L.
Cenchrus americanus (L.) Morrone
Cenchrus complanatus (Nees) Morrone
Cenchrus longisetus M.C.Johnst.
Cenchrus setigerus Vahl
Cenchrus tribuloides L.
Chloris truncata R.Br.
Cortaderia jubata (Lemoine ex Carrière) Stapf
Cortaderia selloana (Schult. & Schult.f.) Asch. & Graebn.
Cynodon aethiopicus Clayton & J.R.Harlan
Cynodon nlemfuensis Vanderyst
Deschampsia caespitosa (L.) P.Beauv. subsp. beringensis
(Hultén) W.E.Lawr.
Dichanthium tenue (R.Br.) A.Camus
Dichelachne crinita (L.f.) Hook.f.
Dichelachne micrantha (Cav.) Domin
Digitaria abyssinica (Hochst. ex A.Rich.) Stapf
Digitaria bicornis (Lam.) Roem. & Schult.
Digitaria divaricatissima (R.Br.) Hughes
Digitaria eriantha Steud.
Digitaria horizontalis Willd.
Digitaria radicosa (J.Presl) Miq.
Disakisperma dubia (Kunth) P.M.Peterson & N.Snow
Distichlis spicata (L.) Greene
Ehrharta calycina Sm.
Ehrharta erecta Lam.
Entolasia marginata (R.Br.) Hughes
Eragrostis ciliaris (L.) R.Br.
Snow & Lau 2010:46 (NEWNAT/K)
Herbst & Clayton 1998:18 (NEWNAT/K [as Andropogon glomeratus])
Herbst & Clayton 1998:19 (NEWNAT/O)
Starr et al. 2003:29 (NEWNAT/EM)
Staples et al. 2002:13 (NEWNAT/H)
Oppenheimer 2003:19 (NEWNAT/Mo, EM, WM)
Herbarium Pacificum Staff 1997:18 (NEWNAT/EM)
Herbst & Clayton 1998:19 (NEWNAT/EM)
Herbst & Wagner 1999:25 (NEWNAT/Mo. EM, H)
Herbst & Wagner 1999:25 (NEWNAT/Mo, H)
Oppenheimer 2008:32 (NEWNAT/Mo, WM)
Oppenheimer 2007:29 (NEWNAT/WM [as Pennisetum glaucum])
Herbst & Clayton 1998:32 (NEWNAT/O, L, H [as Pennisetum complanatum])
Herbst & Clayton 1998:32 (NEWNAT/H [as Pennisetum villosum])
Starr et al. 2004:26 (NEWNAT/K)
Lorence et al. 1995:44 (NEWNAT/Ka)
Herbst & Wagner 1999:25 (NEWNAT/O, Ka)
Loope & Medeiros 1992: 7 (NEWNAT/EM)
Frohlich & Lau 2010:15 (NEWNAT/O)
Herbst & Clayton 1998:22 (NEWNAT/O, H)
Herbst & Wagner 1999:25 (NEWNAT/Mo, H)
Snow & Davidse 2011:17 (NEWNAT/EM)
Herbst & Clayton 1998:22 (NEWNAT/O, EM)
Starr et al. 2003:29 (NEWNAT/EM)
Snow & Davidse 2011:18 (NEWNAT/K)
Herbst & Clayton 1998:23 (NEWNAT/K, M)
Snow & Lau 2010:50 (NEWNAT/WM)
Herbst et al. 2004:9 (NEWNAT/L, H)
Herbst & Clayton 1998:23 (NEWNAT/O?, M?, H)
Herbst & Wagner 1999:26 (NEWNAT/O, L?)
Staples et al. 2003:18 (NEWNAT/H)
Snow & Davidse 2011:18 (NEWNAT/Mo [as Leptochloa dubia])
Herbarium Pacificum Staff 1999:7 (NEWNAT/M)
Herbst & Wagner 1999:26 (NEWNAT/EM?, H)
Herbst & Clayton 1998:25 (NEWNAT/EM)
Flynn & Lorence 1998:5 (NEWNAT/H)
Herbst & Wagner 1999:26 (NEWNAT/O)
G-14
New Naturalized Taxa
Group
Family
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Monocot
Monocot
Poaceae
Poaceae
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Scientific name
Literature source
Eragrostis curvula (Schrad.) Nees
Eragrostis dielsii Pilg. ex Diels & Pritz.
Eragrostis elongata (Willd.) J.Jacq.
Eragrostis parviflora (R.Br.) Trin.
Eragrostis pilosa (L.) P.Beauv.
Eragrostis superba Peyr.
Eragrostis tenuifolia (A.Rich.) Hochst. ex Steud.
Eragrostis trichophora Coss. & Durieu
Eragrostis unioloides (Retz.) Nees ex Steud.
Eriochloa procera (Retz.) C.E.Hubb.
Glyceria notata Chevall.
Hackelochloa granularis (L.) Kuntze
Hemarthria altissima (Poir.) Stapf & C.E.Hubb.
Hordeum brachyantherum Nevski
Hordeum vulgare L.
Hyparrhenia dregeana (Nees) Stent
Hyparrhenia hirta (L.) Stapf
Ischaemum ciliare Retz.
Ischaemum timorense Kunth
Leptochloa panicea (Retz.) Ohwi subsp. brachiata (Steud.)
N.Snow
Leymus triticoides (Buckley) Pilg.
Lolium temulentum L.
Miscanthus floridulus (Labill.) Warb. ex K.Schum. &
Lauterb.
Muhlenbergia mexicana (L.) Trin.
Oplismenus hirtellus (L.) P.Beauv. subsp. undulatifolius
U.Scholz
Panicum sp. A
Paspalidium distans (Trin.) Hughes
Paspalum arundinaceum Poir.
Paspalum distichum L.
Paspalum longifolium Roxb.
Paspalum macrophyllum Kunth
Paspalum malacophyllum Trin.
Paspalum mandiocanum Trin. var. mandiocanum
Paspalum notatum Flüggé
Paspalum paniculatum L.
Paspalum pilosum Lam.
Herbst & Clayton 1998:26 (NEWNAT/O, Ka)
Snow & Lau 2010:50 (NEWNAT/Mo)
Lorence et al. 1995:45 (NEWNAT/K)
Flynn & Lorence 1998:5 NEWNAT/K)
Lorence et al. 1995:45 (NEWNAT/K)
Herbst & Clayton 1998:25, 28 (NEWNAT/H)
Herbst & Clayton 1998:26, 28 (NEWNAT/O, H)
Snow & Lau 2010:51 (NEWNAT/M)
Herbst & Wagner 1999:26 (NEWNAT/H)
Oppenheimer 2008:32 (NEWNAT/Mo)
Herbst & Wagner 1999:27 (NEWNAT/EM, H)
Herbst & Wagner 1999:27 (NEWNAT/H)
Starr et al. 2004:26 (NEWNAT/EM)
Herbst & Clayton 1998:28 (NEWNAT/H)
Herbst & Wagner 1999:27 (NEWNAT/K, O, EM, H)
Herbst & Wagner 1996:11 (NEWNAT/L)
Wagner & Herbst 1995:24 (NEWNAT/Mo, L)
Herbst & Clayton 1998:29 (NEWNAT/EM [as Ischaemum indicum])
Herbst & Wagner 1996:11 (NEWNAT/EM)
Frohlich & Lau 2012:44 (NEWNAT/O)
Herbst & Clayton 1998:29 (NEWNAT/H [as Elymus triticoides])
Herbst & Wagner 1999:28 (NEWNAT/O)
Frohlich & Lau 2012:45 (NEWNAT/O)
Herbst & Clayton 1998:30 (NEWNAT/EM)
Snow & Lau 2010:52 (NEWNAT/K, O, Mo, EM, H)
Herbst & Clayton 1998:30 (NEWNAT/Mo)
Herbst & Clayton 1998:31 (NEWNAT/Ni)
Snow & Lau 2010:53, 56 (NEWNAT/EM [as Paspalum aff. laxum])
Herbst & Wagner 1999:28 (NEWNAT/K, O)
Staples et al. 2003:18 (NEWNAT/K)
Herbst & Clayton 1998:32 (NEWNAT/H)
Oppenheimer 2004:16 (NEWNAT/WM)
Snow & Lau 2010:53, 55 (NEWNAT/O, EM [as Paspalum aff. thunbergii])
Lorence & Flynn 1999:6 (NEWNAT/K)
Herbst & Wagner 1999:28 (NEWNAT/O, H)
Snow & Lau 2010:53, 56 (NEWNAT/Mo)
G-15
New Naturalized Taxa
Group
Family
Scientific name
Literature source
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Monocot
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Poaceae
Paspalum setaceum Michx.
Paspalum unispicatum (Scribn. & Merr.) Nash
Paspalum virgatum L.
Phalaris aquatica L.
Phalaris minor Retz.
Phleum pratense L.
Phyllostachys aurea Carrière ex Rivière & C.Rivière
Piptatherum miliaceum (L.) Coss.
Polypogon fugax Nees ex Steud.
Rytidosperma biannulare (Zotov) Connor & Edgar
Schedonorus arundinaceus (Schreb.) Dumort.
Schedonorus pratensis (Huds.) P.Beauv.
Schizachyrium scoparium (Michx.) Nash
Setaria sphacelata (Schumach.) M.B.Moss ex Stapf &
C.E.Hubb.
Sorghum arundinaceum (Desv.) Stapf
Sorghum xdrummondii (Nees ex Steud.) Millsp. & Chase
Sporobolus elongatus R.Br.
Sporobolus piliferus (Trin.) Kunth
Monocot
Monocot
Monocot
Monocot
Poaceae
Poaceae
Poaceae
Poaceae
Sporobolus pyramidatus (Lam.) Hitchc.
Trisetum flavescens (L.) P.Beauv.
Triticum aestivum L.
Urochloa brizantha (Hochst. ex A.Rich.) R.D.Webster
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Monocot
Pteridophyte
Pteridophyte
Pteridophyte
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Poaceae
Potamogetonaceae
Zingiberaceae
Blechnaceae
Davalliaceae
Dicksoniaceae
Urochloa decumbens (Stapf) R.D.Webster
Urochloa distachya (L.) T.Q.Nguyen
Urochloa mollis (Sw.) Morrone & Zuloaga
Urochloa plantaginea (Link) R.D.Webster
Urochloa reptans (L.) Stapf
Zizania latifolia (Griseb.) Turcz. ex Stapf
Zoysia matrella (L.) Merr. var. pacifica Goudsw.
Potamogeton tricarinatus A.Benn.
Alpinia zerumbet (Pers.) B.L.Burtt & R.M.Sm.
Blechnum orientale L.
Davallia fejeensis Hook.
Dicksonia fibrosa Colenso
Oppenheimer 2007:29 (NEWNAT/EM)
Snow & Davidse 2011:19 (NEWNAT/K)
Snow & Lau 2010:54, 55 (NEWNAT/O)
Herbst & Clayton 1998:33 (NEWNAT/EM, H)
Herbst & Wagner 1999:28 (NEWNAT/O, Mo, H)
Herbst & Wagner 1999:29 (NEWNAT/K, O, EM?)
Staples et al. 2002:14 (NEWNAT/O)
Dougill & Banko 2002 (NEWNAT/H [as Oryzopsis miliacea])
Herbst & Clayton 1998:34 (NEWNAT/O)
Snow 2008:40 (NEWNAT/Mo, M?);
Wagner et al. 1997:60 (NEWNAT/K [as Festuca arundinacea])
Snow 2008:40 (NEWNAT/EM)
Lorence et al. 1995:47 (NEWNAT/K)
Starr et al. 2002:23 (NEWNAT/EM)
Snow & Lau 2010:56 (NEWNAT/H)
Herbst & Clayton 1998:35 (NEWNAT/O, L, H?)
Herbst & Clayton 1998:36 (NEWNAT/L)
Herbst & Clayton 1998:35, 36 (NEWNAT/H); [Note: this first record was later reidentified
as Sporobolus pyramidatus [Snow & Lau 2010:56], but authentic piliferus has also been
documented from O‘ahu [Snow 2008:41]
Wagner & Herbst 1995:24 (NEWNAT/O)
Herbst & Clayton 1998:36 (NEWNAT/EM)
Wagner & Herbst 2003:57 (NEWNAT/K, O, Mo, L, M, H)
Starr et al. 2006:39 (NEWNAT/Ka [as Brachiaria brizantha]) [Note: this first record was
later reidentified as B. decumbens [Snow & Lau 2010:49], but authentic brizantha has also
been documented from East Maui [Oppenheimer 2008:31]
Snow & Lau 2010:49 (NEWNAT/K, Mo?, Ka, H [as Brachiaria decumbens]
Lorence et al. 1995:44 (NEWNAT/K [as Brachiaria subquadripara)
Staples et al. 2002:13 (NEWNAT/O [as Brachiaria mollis])
Wagner & Herbst 1995:23 (NEWNAT/O, Mo [as Brachiaria plantaginea])
Herbst & Clayton 1998:20 (NEWNAT/O [as Brachiaria reptans])
Lorence & Flynn 1999:6 (NEWNAT/K)
Flynn & Lorence 1998:5 (NEWNAT/K)
Wagner et al. 2012:102 (NEWNAT/EM)
Flynn & Lorence 2002:16 (NEWNAT/K)
Lau & Frohlich 2012:10 (NEWNAT/O)
Frohlich & Lau 2008:5 (NEWNAT/O)
Lorence & Flynn 2006:1 (NEWNAT/H)
G-16
New Naturalized Taxa
Group
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Pteridophyte
Family
Marsileaceae
Polypodiaceae
Polypodiaceae
Pteridaceae
Salviniaceae
Selaginellaceae
Scientific name
Marsilea crenata C.Presl
Phymatosorus scolopendria (Burm.f.) Pic.Serm.
Pyrrosia piloselloides (L.) M.G.Price
Pteris tremula R.Br.
Salvinia molesta D.S.Mitch.
Selaginella umbrosa Lem. ex Hieron.
Literature source
Wilson 2003:5 (NEWNAT/O)
Oppenheimer 2006:13 (NEWNAT/WM)
Lau & Frohlich 2012:21 (NEWNAT/O)
Oppenheimer 2007:30 (NEWNAT/EM)
Wilson 2003:6 (NEWNAT/O, H)
Wilson 2003:6 (NEWNAT/H)
G-17
View publication stats
G-18
New Naturalized Taxa
Clyde Imada