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Pinus caribaea

Pinus caribaea

Pinus caribaea

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Taxonomy and nomenclature<br />

Family: Pinaceae<br />

Varieties: <strong>Pinus</strong> <strong>caribaea</strong> var. bahamensis (Griseb.)<br />

W.H.G. Barrett & Golfari, P. <strong>caribaea</strong> var. <strong>caribaea</strong>,<br />

P. <strong>caribaea</strong> var. hondurensis (Sénéclauze) W.H.G.<br />

Barrett & Golfari.<br />

Synonyms: <strong>Pinus</strong> taeda var. heterophylla Elliott, P.<br />

recurvata Rowlee.<br />

Vernacular/common names: Caribbean pine, pitch<br />

pine (Eng.); pino de la costa, ocote blanco, pino<br />

caribe, pino caribeño de Honduras (Sp.); pin jaune,<br />

pin mate (Fr.); karibische kiefer (Germ.); Honduran<br />

yellow pine (trade name).<br />

Distribution and habitat<br />

Native to Central America and the Caribbean, widely<br />

planted throughout the American, Asian and African<br />

tropics and subtropics. P. <strong>caribaea</strong> var. <strong>caribaea</strong> is<br />

confined to Cuba and the Isla de la Juventud, P.<br />

<strong>caribaea</strong> var. bahamensis is indigenous to certain islands<br />

of the Bahamas and the Caicos groups and P.<br />

<strong>caribaea</strong> var. hondurensis can be found in the eastern<br />

half of Central America south-east from the Yucatán<br />

peninsula.<br />

Grows best in frost-free areas up to 700 m altitude<br />

on more fertile sites with good drainage and annual<br />

rainfall of 1000 - 3000 mm.<br />

1, mature male strobili; 2, female strobili at receptive stage;<br />

3, female strobili two months old; 4, mature cones; 5, cone<br />

scales; 6, seeds. From: Robbins, 1994.<br />

SEED LEAFLET<br />

No. 40 September 2000<br />

<strong>Pinus</strong> <strong>caribaea</strong> Morelet<br />

Danida Forest Seed Centre<br />

Uses<br />

Used for fuel, pulpwood, timber, resin and land reclamation.<br />

Botanical description<br />

Tree up to 45 m tall and 1 m in diameter, with conical<br />

and irregular crown. Bark is grey on young trees, later<br />

dark and with fissures. Needles in fascicles of 3-5 at<br />

the ends of twigs, mostly 15-25 cm long, up to 1.5<br />

mm broad, slightly toothed, with whitish lines. The<br />

needles remain attached for 2 years.<br />

Strobili appear before the new leaves; male strobili<br />

in clusters, mostly in the lower part of crown; female<br />

strobili 2-5 together, mostly in the upper part of the<br />

crown.<br />

Fruit and seed description<br />

Seed about twice as long as broad, triangular and<br />

pointed at the ends. On average less that 6 mm long, 3<br />

mm broad and black to grey or brown. The membranous<br />

wing is up to 20 mm long, sometimes fused with<br />

the seed coat but becoming detached when moistened.<br />

There are 35-40 seeds per cone. and 59,000-<br />

72,000 dewinged seeds/kg, depending on the variety.<br />

Flowering and fruiting habit<br />

The cones mature at the onset of the rainy season but<br />

there is often variation between trees and stands. In<br />

general, cones tend to mature during the same period,<br />

despite variation in flowering times. Seed production<br />

in exotic plantations is often poor due to either cool<br />

temperatures preventing the formation of flowers or<br />

humid conditions during flowering preventing pollination.<br />

When the tree is 3-4 years old, it begins to produce<br />

female cones but seed setting is low unless there are<br />

mature pollinating trees close by.<br />

Harvest<br />

The cones are mature when: 1. more than half of the<br />

cone has turned brown, 2. when cut in two, the cone<br />

axis is dark brown, 3. the apex of the cone feels firm<br />

when pressed with the thumb 4. the seed coat is darkening<br />

in colour and 5. the inside of the seed is white<br />

and firm and filling the cavity.<br />

Collection is done directly from the tree. Care must<br />

be taken not to break the fragile branchlets as this can<br />

seriously reduce crop size for several years. Once the


cones have become brown and are still moist, the<br />

scales may start to open and pregermination occur. A<br />

cutting test will reveal this.<br />

Processing and handling<br />

After harvest the cones should not be left in sacks for<br />

more than 1-2 days and always in a cool, dry and<br />

properly ventilated place. Before drying, the cones<br />

are precured on trays with wire mesh, under shade,<br />

until the cone has turned completely brown (5-10<br />

days). After precuring, the cones open relatively easily<br />

and sun drying is normally adequate, provided the<br />

climate is not too humid.<br />

When the seeds have been extracted they are<br />

dewinged and cleaned and then dried in the sun on a<br />

sheet and raked frequently. The seeds will lose about<br />

1-1.5 % moisture per hour. Cleaning the seed before<br />

sowing to eliminate empty and dead seed can<br />

improve germination considerably.<br />

Storage and viability<br />

The seed is orthodox and can be stored at temperatures<br />

below 0°C. At 6-8% mc and 3-4° C in airtight<br />

containers, the seed can be stored for at least 5 years<br />

with little loss in viability. At room temperature, few<br />

seeds will survive one year of storage. As long as the<br />

seed is kept dry and cool, fumigation and application<br />

of fungicides are not necessary and may be harmful.<br />

Dormancy and pretreatment<br />

The seeds are not dormant, but in some places it is<br />

practice to soak the seeds in clean water for 12 hours<br />

followed by 2-3 days cold stratification at 4-5°C to<br />

obtain a more uniform germination.<br />

Sowing and germination<br />

Germination normally begins seven days after sowing<br />

and after 12-15 days reaches some 80%. The seeds can<br />

be sown directly in plastic bags with 1-2 seeds per bag,<br />

or in germination trays from where they are transplanted<br />

into bags when they are 3-4 cm long.<br />

Mycorrhiza is necessary for the sapling growth, and<br />

soil taken near established trees should be added. Suitable<br />

sowing medium is sand:forest soil in a 3:1 ratio. If<br />

the seedbeds are covered with mulch, it reduces the<br />

risk of fungi.<br />

Phytosanitary problems<br />

Cones that are inadequately stored are easily susceptible<br />

to infestation by fungal moulds, which may<br />

cause problems during germination. Infestation by<br />

pathogenic fungi may also happen especially in seed<br />

from high rainfall areas.<br />

Selected readings<br />

Birks, J.S., Barnes, R.D., 1990. Provenance variation in<br />

<strong>Pinus</strong> <strong>caribaea</strong>, P. oocarpa and P. patula ssp. tecunumanii.<br />

Tropical Forestry Papers, No. 21. Oxford Forestry Institute,<br />

University of Oxford.<br />

CATIE 1991. Pino Caribe, <strong>Pinus</strong> <strong>caribaea</strong>, Especies de<br />

árbol de uso múltiple en América Central. Turrialba<br />

(C:R:). Coleccíon guías silviculturales no. 8, 59 pp.<br />

Dvorak, W.S., Ross, K.D., Liu, Y., 1993. Performance of<br />

<strong>Pinus</strong> <strong>caribaea</strong> var. hondurensis in Brazil, Colombia and<br />

Venezuela. CAMCORE Bulletin on Tropical Forestry, No.<br />

11, 47 pp.<br />

Lamb, A.F.A., 1973. Fast growing timber trees of the lowland<br />

tropics. No. 6. <strong>Pinus</strong> <strong>caribaea</strong>, vol. 1. Commonwealth<br />

Forestry Institute, 254 pp.<br />

Napier, I.A. & Willan, R.L. 1983. Nursery Techniques for<br />

Tropical and Subtropical Pines. DFSC Tecnical Note No. 4.<br />

Soerianegara, I., Lemmens, R.H.M.J., eds., 1993. Plant<br />

Resources of South-East Asia No. 5(1). Timber trees: major<br />

commercial timbers. Wageningen, Netherlands: Pudoc Scientific<br />

Publishers. 610 pp.<br />

Robbins, A.M.J. 1994. <strong>Pinus</strong> <strong>caribaea</strong>. Seed Leaflet No.<br />

2. DFSC<br />

P. <strong>caribaea</strong> var. hondurensis. Alamikamba, Nicaragua.<br />

Photo: E.A. Gutierrez, CAMCORE.<br />

THIS NOTE WAS PREPARED IN COLLABORATION<br />

WITH CENTRO AGRONÓMICO TROPICAL DE<br />

INVESTIGACIÓN Y ENSEÑANZA<br />

Authors: Rodolfo Salazar, CATIE<br />

Dorthe Jøker, DFSC<br />

Danida Forest Seed Centre Phone: +45-49190500<br />

Krogerupvej 21 Fax: +45-49160258<br />

DK-3050 Humlebaek Email: dfsc@sns.dk<br />

Denmark Website: www.dfsc.dk

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